treatments-xml/data/A2/44/87/A24487FE3E7D6A706D3FFD614A69FD62.xml

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<mods:title id="3478F397C1808438BA92116EDBF92DD5">A revision of Oocyclus Sharp of Mexico and Central America (Coleoptera: Hydrophilidae)</mods:title>
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<taxonomicName id="EDED4D6B3E7D6A726D3FFD614B91FD12" ID-CoL="49RW4" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[264,664,644,670]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="1899EAFA3E7D6A726D3FFD614B91FD12" bold="true" box="[264,664,644,670]" pageId="12" pageNumber="13">
<emphasis id="1899EAFA3E7D6A726D3FFD6148F2FD12" bold="true" box="[264,507,644,670]" italics="true" pageId="12" pageNumber="13">Oocyclus maculatus</emphasis>
<bibRefCitation id="4E7C4B193E7D6A726E34FD614B91FD12" author="Sharp" box="[515,664,644,670]" pageId="12" pageNumber="24" refString="Sharp, D. (1882). Insecta, Coleoptera, Vol. 1, Part 2. In: Godman, F. D, Schvin, O.: Biologia Centrali-Americana (16): XV + 824 pp. London. (pp 1 - 144 issued in 1882)." type="book chapter" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="2A5236E83E7D6A726D3FFD4248E9FD4D" blockId="12.[264,664,644,705]" box="[264,480,679,705]" pageId="12" pageNumber="13">
(
<figureCitation id="B2D62A6D3E7D6A726D26FD424888FD4D" box="[273,385,679,705]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="31.[264,368,1871,1895]" captionStartId-1="32.[264,368,1591,1615]" captionTargetBox-0="[271,1292,292,1841]" captionTargetBox-1="[264,1322,310,1566]" captionTargetId-0="figure@31.[264,1323,281,1861]" captionTargetId-1="figure@32.[264,1323,308,1567]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="FIGURE 8. Oocyclus maculatus Sharp holotype. Dorsal and lateral habitus." captionText-1="FIGURE 9. Oocyclus maculatus Sharp (specimen from Panama). Dorsal and lateral habitus." httpUri-0="https://zenodo.org/record/169617/files/figure.png" httpUri-1="https://zenodo.org/record/169618/files/figure.png" pageId="12" pageNumber="13">Figs. 8–9</figureCitation>
, 
<figureCitation id="B2D62A6D3E7D6A726DB9FD4248D1FD4D" box="[398,472,679,705]" captionStart="FIGURES 20 – 23" captionStartId="40.[264,382,1812,1836]" captionTargetBox="[329,1218,291,1800]" captionTargetId="figure@40.[264,1323,289,1801]" captionTargetPageId="40" captionText="FIGURES 20 – 23. Oocyclus aedeagi, dorsal views. Scale bars = 0.2 mm. – 20. O. maculatus Sharp (holotype). – 21. O. maculatus Sharp (specimen from Panama). – 22. O. muscus n. sp. (holotype). – 23. O. ornatus n. sp. (holotype)." httpUri="https://zenodo.org/record/158182/files/figure.png" pageId="12" pageNumber="13">20–21</figureCitation>
)
</paragraph>
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<subSubSection id="62F765633E7D6A726D3FFD124B9FFC81" box="[264,662,758,782]" pageId="12" pageNumber="13" type="reference_group">
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<treatmentCitationGroup id="0AFD11C63E7D6A726D3FFD124B9FFC81" box="[264,662,758,782]" pageId="12" pageNumber="13">
<taxonomicName id="EDED4D6B3E7D6A726D3FFD124B9AFC81" ID-CoL="49RW4" authority="Sharp, 1882: 63" authorityName="Sharp" authorityPageNumber="63" authorityYear="1882" box="[264,659,758,782]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="1899EAFA3E7D6A726D3FFD1248E9FC82" box="[264,480,759,782]" italics="true" pageId="12" pageNumber="13">Oocyclus maculatus</emphasis>
<treatmentCitation id="AB4C10F93E7D6A726DD1FD134B9AFC81" author="Sharp" box="[486,659,758,781]" page="63" pageId="12" pageNumber="13" year="1882">
<bibRefCitation id="4E7C4B193E7D6A726DD1FD134B63FC81" author="Sharp" box="[486,618,758,781]" pageId="12" pageNumber="24" refString="Sharp, D. (1882). Insecta, Coleoptera, Vol. 1, Part 2. In: Godman, F. D, Schvin, O.: Biologia Centrali-Americana (16): XV + 824 pp. London. (pp 1 - 144 issued in 1882)." type="book chapter" year="1882">Sharp, 1882</bibRefCitation>
: 63
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.
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<emphasis id="1899EAFA3E7D6A726D3FFCA448CBFCD7" bold="true" box="[264,450,833,859]" pageId="12" pageNumber="13">
<typeStatus id="F556884A3E7D6A726D3FFCA4484CFCD7" box="[264,325,833,859]" pageId="12" pageNumber="13">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="F556884A3E7D6A726DE2FCA44B4BFCD7" box="[469,578,833,859]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
male. Fixed by monotypy. “ 
<taxonomicName id="EDED4D6B3E7D6A726FA8FCA44D99FCD7" box="[927,1168,833,859]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="maculatus">Oocyclus maculatus</taxonomicName>
/ 
<typeStatus id="F556884A3E7D6A726895FCA44DD5FCD7" box="[1186,1244,833,859]" pageId="12" pageNumber="13">Type</typeStatus>
D. S./ Cerro Zunil Guatem­/ ala, 
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./ Champion.[written on same card as specimen]”, “ 
<typeStatus id="F556884A3E7D6A726D25FC744846FC27" box="[274,335,913,939]" pageId="12" pageNumber="13">Type</typeStatus>
[red disc]”, “Cerro Zunil/ 
<collectingCountry id="52FA76783E7D6A726EA5FC744A19FC27" box="[658,784,913,939]" name="Guatemala" pageId="12" pageNumber="13">Guatemala</collectingCountry>
/ Champion”, “B. C. A. Col. 
<date id="5E5310283E7D6A726845FC744DA9FC27" box="[1138,1184,913,939]" pageId="12" pageNumber="13">I. 2.</date>
/ 
<taxonomicName id="EDED4D6B3E7D6A726883FC744C2BFC27" box="[1204,1314,913,939]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Oocyclus</taxonomicName>
/ 
<taxonomicName id="EDED4D6B3E7D6A726D3FFC5C488BFC5F" box="[264,386,953,979]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="maculatus">maculatus</taxonomicName>
, / Sharp.” (
<collectionCode id="4CFCAE2D3E7D6A726E32FC5C4B6AFC5F" box="[517,611,953,979]" pageId="12" pageNumber="13">BMNH</collectionCode>
).
</paragraph>
<paragraph id="2A5236E83E7D6A736D0FFC044D5BFE72" blockId="12.[264,1323,833,1899]" lastBlockId="13.[264,1323,284,1870]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="1899EAFA3E7D6A726D0FFC044D6CFC77" bold="true" box="[312,1125,993,1019]" pageId="12" pageNumber="13">
Other Material Examined (320). 
<collectingCountry id="52FA76783E7D6A726EEAFC044A90FC77" box="[733,921,993,1019]" name="Costa Rica" pageId="12" pageNumber="13">COSTA RICA</collectingCountry>
: Alajuela Prov.,
</emphasis>
On Rt. 126, 
<quantity id="ED159B0D3E7D6A726931FC044824FBAF" metricMagnitude="3" metricUnit="m" metricValue="9.4" pageId="12" pageNumber="13" unit="km" value="9.4">9.4 km</quantity>
S. Vara Blanca, 
<date id="5E5310283E7D6A726DC6FBEC4B78FBAF" box="[497,625,1033,1059]" pageId="12" pageNumber="13" value="2003-06-25">25.vi.2003</date>
, rock face seeps, A. E. Z. Short leg. (15 
<collectionCode id="4CFCAE2D3E7D6A72687DFBEC4D9FFBAF" box="[1098,1174,1033,1059]" pageId="12" pageNumber="13">AEZS</collectionCode>
, 25 
<collectionCode id="4CFCAE2D3E7D6A7268F1FBEC4C15FBAF" box="[1222,1308,1033,1059]" pageId="12" pageNumber="13">INBIO</collectionCode>
); On Rt. 126, 
<quantity id="ED159B0D3E7D6A726DAAFBD44B09FBC7" box="[413,512,1073,1099]" metricMagnitude="4" metricUnit="m" metricValue="1.54" pageId="12" pageNumber="13" unit="km" value="15.4">15.4 km</quantity>
SE of Poasito, 
<quantity id="ED159B0D3E7D6A726E8CFBD44BF7FBC7" box="[699,766,1073,1099]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="12" pageNumber="13" unit="m" value="960.0">960m</quantity>
elev., 
<date id="5E5310283E7D6A726F79FBD44AB6FBC7" box="[846,959,1073,1099]" pageId="12" pageNumber="13" value="2004-01-16">16.i.2004</date>
, rock seeps/waterfall, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­056 (22 
<collectionCode id="4CFCAE2D3E7D6A726F34FBBC4A50FBFF" box="[771,857,1113,1139]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<quantity id="ED159B0D3E7D6A726F58FBBC4AA5FBFF" box="[879,940,1113,1139]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="km" value="1.0">1 km</quantity>
S. Cariblanco, 
<quantity id="ED159B0D3E7D6A726851FBBC4DA0FBFF" box="[1126,1193,1113,1139]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="12" pageNumber="13" unit="m" value="870.0">870m</quantity>
elev., wall seep, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­055 (4 
<collectionCode id="4CFCAE2D3E7D6A726FAFFB644AE6FB17" box="[920,1007,1153,1179]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<quantity id="ED159B0D3E7D6A726833FB644D5CFB17" box="[1028,1109,1153,1179]" metricMagnitude="3" metricUnit="m" metricValue="6.1" pageId="12" pageNumber="13" unit="km" value="6.1">6.1 km</quantity>
N. Los Lagos, on Los Lagos­Colonia Rd., 
<quantity id="ED159B0D3E7D6A726E04FB4C4B7CFB4F" box="[563,629,1193,1219]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="12" pageNumber="13" unit="m" value="870.0">870m</quantity>
elev., seeps on Rio San Lorenzo, 
<date id="5E5310283E7D6A726830FB4C4D71FB4F" box="[1031,1144,1193,1219]" pageId="12" pageNumber="13" value="2003-01-12">12.i.2003</date>
, A. Short &amp; R. Roughley leg., CR­03­04 (18 
<collectionCode id="4CFCAE2D3E7D6A726E46FB344BB4FB67" box="[625,701,1233,1259]" pageId="12" pageNumber="13">AEZS</collectionCode>
, 63 
<collectionCode id="4CFCAE2D3E7D6A726EC4FB344A40FB67" box="[755,841,1233,1259]" pageId="12" pageNumber="13">INBIO</collectionCode>
); Same locality but 
<date id="5E5310283E7D6A726870FB344DB1FB67" box="[1095,1208,1233,1259]" pageId="12" pageNumber="13" value="2004-01-10">10.i.2004</date>
, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­024 (3 
<collectionCode id="4CFCAE2D3E7D6A726ED1FB1C4A34FA9F" box="[742,829,1273,1299]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<quantity id="ED159B0D3E7D6A726F67FB1C4A85FA9F" box="[848,908,1273,1299]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="km" value="6.0">6 km</quantity>
N. Los Lagos, small stream nr. Rio San Lorenzo, 
<date id="5E5310283E7D6A726D99FAC44B26FAB7" box="[430,559,1313,1339]" pageId="12" pageNumber="13" value="2003-06-12">12.vi.2003</date>
, A. E. Z. Short leg. (1 
<collectionCode id="4CFCAE2D3E7D6A726F74FAC44A90FAB7" box="[835,921,1313,1339]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<quantity id="ED159B0D3E7D6A726F98FAC44AE4FAB7" box="[943,1005,1313,1339]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="13" unit="km" value="7.0">7 km</quantity>
N. Bajos del Toro, at 
<taxonomicName id="EDED4D6B3E7D6A7268CFFAC44848FAEF" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="44" phylum="Arthropoda" rank="species" species="catarata" status="sp. nov.">Catarata</taxonomicName>
del Toro, 
<quantity id="ED159B0D3E7D6A726D8AFAAC4B06FAEF" box="[445,527,1353,1379]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="12" pageNumber="13" unit="m" value="1210.0">1210m</quantity>
elev., 
<date id="5E5310283E7D6A726E69FAAC4BC6FAEF" box="[606,719,1353,1379]" pageId="12" pageNumber="13" value="2004-01-15">15.i.2004</date>
, seeps, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­ 0 52 (1 
<collectionCode id="4CFCAE2D3E7D6A726D69FA9448BCFA07" box="[350,437,1393,1419]" pageId="12" pageNumber="13">INBIO</collectionCode>
); San Ramon Biological Reserve, 
<quantity id="ED159B0D3E7D6A726F67FA944A9AFA07" box="[848,915,1393,1419]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="13" unit="m" value="800.0">800m</quantity>
elev., 
<date id="5E5310283E7D6A726FD4FA944D49FA07" box="[995,1088,1393,1419]" pageId="12" pageNumber="13" value="1995-03">iii.1995</date>
, G. Carballo leg. (1 
<collectionCode id="4CFCAE2D3E7D6A726D3FFA7C4857FA3F" box="[264,350,1433,1459]" pageId="12" pageNumber="13">INBIO</collectionCode>
); Alberto Brenes Biological Reserve, Rio San Lorencito, 
<quantity id="ED159B0D3E7D6A72683DFA7C4D44FA3F" box="[1034,1101,1433,1459]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="13" unit="m" value="800.0">800m</quantity>
elev., 
<date id="5E5310283E7D6A7268ABFA7C4C2DFA3F" box="[1180,1316,1433,1459]" pageId="12" pageNumber="13" value="1999-06-30">30.VI.1999</date>
, C. Rojas leg. (1 
<collectionCode id="4CFCAE2D3E7D6A726DFFFA244B17FA57" box="[456,542,1473,1499]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<emphasis id="1899EAFA3E7D6A726E04FA244BE3FA57" bold="true" box="[563,746,1473,1499]" pageId="12" pageNumber="13">Cartago Prov.,</emphasis>
<taxonomicName id="EDED4D6B3E7D6A726ECEFA244A59FA57" box="[761,848,1473,1499]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="tapanti" status="sp. nov.">Tapanti</taxonomicName>
N. P., 
<date id="5E5310283E7D6A726F97FA244D29FA57" box="[928,1056,1473,1499]" pageId="12" pageNumber="13" value="2003-06-22">22.vi.2003</date>
, rock seeps/waterfalls, A. E. Z. Short leg. (8 
<collectionCode id="4CFCAE2D3E7D6A726E23FA0C4B63F98F" box="[532,618,1513,1539]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<taxonomicName id="EDED4D6B3E7D6A726EB6FA0C4BD1F98F" box="[641,728,1513,1539]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="tapanti" status="sp. nov.">Tapanti</taxonomicName>
N. P., 
<quantity id="ED159B0D3E7D6A726F19FA0C4A88F98F" box="[814,897,1513,1539]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="13" unit="km" value="3.5">3.5 km</quantity>
past Ranger Station, 
<date id="5E5310283E7D6A7268B1FA0C4DE0F98F" box="[1158,1257,1513,1539]" pageId="12" pageNumber="13" value="2004-01-06">6.i.2004</date>
, rock seep, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­002 (25 
<collectionCode id="4CFCAE2D3E7D6A726F94F9F44AE6F9A7" box="[931,1007,1553,1579]" pageId="12" pageNumber="13">AEZS</collectionCode>
, 54 
<collectionCode id="4CFCAE2D3E7D6A726817F9F44D7EF9A7" box="[1056,1143,1553,1579]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<taxonomicName id="EDED4D6B3E7D6A7268BBF9F44DEAF9A7" box="[1164,1251,1553,1579]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="tapanti" status="sp. nov.">Tapanti</taxonomicName>
N. P., 
<quantity id="ED159B0D3E7D6A726D3FF9DC4852F9DF" box="[264,347,1593,1619]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="12" pageNumber="13" unit="km" value="3.1">3.1 km</quantity>
after gate, 
<date id="5E5310283E7D6A726DD5F9DC4B4CF9DF" box="[482,581,1593,1619]" pageId="12" pageNumber="13" value="2004-01-08">8.i.2004</date>
, wall seep, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­013 (29 
<collectionCode id="4CFCAE2D3E7D6A726D3FF9844857F9F7" box="[264,350,1633,1659]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<taxonomicName id="EDED4D6B3E7D6A726D43F98448C5F9F7" box="[372,460,1633,1659]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="tapanti" status="sp. nov.">Tapanti</taxonomicName>
N. P., 
<quantity id="ED159B0D3E7D6A726E2BF9844B66F9F7" box="[540,623,1633,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="km" value="1.5">1.5 km</quantity>
after gate, 
<quantity id="ED159B0D3E7D6A726EC2F9844A4EF9F7" box="[757,839,1633,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="12" pageNumber="13" unit="m" value="1620.0">1620m</quantity>
elev., 
<date id="5E5310283E7D6A726FA3F9844AFFF9F7" box="[916,1014,1633,1659]" pageId="12" pageNumber="13" value="2004-01-08">8.i.2004</date>
, rock seep, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­011 (6 
<collectionCode id="4CFCAE2D3E7D6A726E8AF96C4A1DF92F" box="[701,788,1673,1699]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<taxonomicName id="EDED4D6B3E7D6A726F1BF96C4A8AF92F" box="[812,899,1673,1699]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="tapanti" status="sp. nov.">Tapanti</taxonomicName>
N. P., 
<quantity id="ED159B0D3E7D6A726FEAF96C4D3AF92F" box="[989,1075,1673,1699]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="12" pageNumber="13" unit="km" value="7.6">7.6 km</quantity>
after gate, 
<date id="5E5310283E7D6A7268F4F96C4C2FF92F" box="[1219,1318,1673,1699]" pageId="12" pageNumber="13" value="2004-01-08">8.i.2004</date>
, 1660m elev., rock seep, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­019 (13 
<collectionCode id="4CFCAE2D3E7D6A7268FFF9544C17F947" box="[1224,1310,1713,1739]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<taxonomicName id="EDED4D6B3E7D6A726D3FF93C4856F97F" box="[264,351,1753,1779]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="tapanti" status="sp. nov.">Tapanti</taxonomicName>
N. P., 
<quantity id="ED159B0D3E7D6A726D83F93C4B01F97F" box="[436,520,1753,1779]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="12" pageNumber="13" unit="km" value="4.6">4.6 km</quantity>
after gate, 
<date id="5E5310283E7D6A726EA5F93C4BFCF97F" box="[658,757,1753,1779]" pageId="12" pageNumber="13" value="2004-01-08">8.i.2004</date>
, 
<quantity id="ED159B0D3E7D6A726F35F93C4A5DF97F" box="[770,852,1753,1779]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="m" value="1500.0">1500m</quantity>
elev., waterfall, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­015 (1 
<collectionCode id="4CFCAE2D3E7D6A726E78F8E44BAFF897" box="[591,678,1793,1819]" pageId="12" pageNumber="13">INBIO</collectionCode>
); Guayabo Nat. Monument, 
<quantity id="ED159B0D3E7D6A726FC9F8E44D47F897" box="[1022,1102,1793,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="13" unit="m" value="1100.0">1100m</quantity>
elev., 
<date id="5E5310283E7D6A726897F8E44C0DF897" box="[1184,1284,1793,1819]" pageId="12" pageNumber="13" value="1994-07">vii.1994</date>
, G. Fonseca leg. (1 
<collectionCode id="4CFCAE2D3E7D6A726DE5F8CC4B21F8CF" box="[466,552,1833,1859]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<emphasis id="1899EAFA3E7D6A726E74F8CC4A21F8CF" bold="true" box="[579,808,1833,1859]" pageId="12" pageNumber="13">Puntarenas Prov.,</emphasis>
Las Cruces Biological Station, Rio Jaba, 3800’ elev., 
<date id="5E5310283E7D6A726DACF8B44B12F8E7" box="[411,539,1873,1899]" pageId="12" pageNumber="13" value="2003-06-19">19.vi.2003</date>
, A. E. Z. Short leg. (2 
<collectionCode id="4CFCAE2D3E7D6A726F07F8B44A8EF8E7" box="[816,903,1873,1899]" pageId="12" pageNumber="13">INBIO</collectionCode>
); 
<emphasis id="1899EAFA3E7D6A726FADF8B44D53F8E7" bold="true" box="[922,1114,1873,1899]" pageId="12" pageNumber="13">San Jose Prov.,</emphasis>
Rt. 243, 
<quantity id="ED159B0D3E7D6A7268FEF8B44C23F8E7" box="[1225,1322,1873,1899]" metricMagnitude="4" metricUnit="m" metricValue="1.91" pageId="12" pageNumber="13" unit="km" value="19.1">19.1 km</quantity>
NE of Dominical, 
<date id="5E5310283E7C6A736DD8FEF94B67FEBA" box="[495,622,284,310]" pageId="13" pageNumber="14" value="2003-06-18">18.vi.2003</date>
, “waterfall and seeps”, A. E. Z. Short leg. (9 
<collectionCode id="4CFCAE2D3E7C6A736886FEF94DF4FEBA" box="[1201,1277,284,310]" pageId="13" pageNumber="14">AEZS</collectionCode>
, 14 
<collectionCode id="4CFCAE2D3E7C6A736D3FFEA14857FED2" box="[264,350,324,350]" pageId="13" pageNumber="14">INBIO</collectionCode>
); Rt. 2 nr. Division, 
<date id="5E5310283E7C6A736E7EFEA14BC0FED2" box="[585,713,324,350]" pageId="13" pageNumber="14" value="2003-06-21">21.vi.2003</date>
, waterfall/seeps, A. E. Z. Short leg. (1 
<collectionCode id="4CFCAE2D3E7C6A7368BCFEA14DEBFED2" box="[1163,1250,324,350]" pageId="13" pageNumber="14">INBIO</collectionCode>
); Station 
<collectingCountry id="52FA76783E7C6A736D76FE8948DDFE0A" box="[321,468,364,390]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="13" pageNumber="14">Santa Elena</collectingCountry>
, “Las Nubes”, 
<quantity id="ED159B0D3E7C6A736EA5FE894BEDFE0A" box="[658,740,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="13" pageNumber="14" unit="m" value="1210.0">1210m</quantity>
elev., 
<date id="5E5310283E7C6A736F0EFE894AD7FE0A" box="[825,990,364,390]" pageId="13" pageNumber="14" value="1996-07-05" valueMax="1996-07-19" valueMin="1996-07-05">5–19.vii.1996</date>
, E. Alfaro leg. (1 
<collectionCode id="4CFCAE2D3E7C6A7368F0FE894C14FE0A" box="[1223,1309,364,390]" pageId="13" pageNumber="14">INBIO</collectionCode>
); same locality but 
<date id="5E5310283E7C6A736DD5FE714B42FE22" box="[482,587,404,430]" pageId="13" pageNumber="14" value="1995-10-01">1.x.1995</date>
, A. M. Maroto, “de luz” leg. (2 
<collectionCode id="4CFCAE2D3E7C6A736FE9FE714D3CFE22" box="[990,1077,404,430]" pageId="13" pageNumber="14">INBIO</collectionCode>
). 
<emphasis id="1899EAFA3E7C6A73687BFE764DC5FE21" bold="true" box="[1100,1228,403,429]" pageId="13" pageNumber="14">
<collectingCountry id="52FA76783E7C6A73687BFE764DC5FE21" box="[1100,1228,403,429]" name="Panama" pageId="13" pageNumber="14">PANAMA</collectingCountry>
</emphasis>
: “V. de Chirique”, 
<quantity id="ED159B0D3E7C6A736DBDFE594B01FE5A" box="[394,520,444,470]" metricMagnitude="2" metricUnit="m" metricValue="6.1341" metricValueMax="12.192" metricValueMin="0.0762" pageId="13" pageNumber="14" unit="ft" value="2012.5" valueMax="4000.0" valueMin="25.0">25­4000 ft</quantity>
, Champion, leg. (1 
<collectionCode id="4CFCAE2D3E7C6A736ED9FE594A42FE5A" box="[750,843,444,470]" pageId="13" pageNumber="14">BMNH</collectionCode>
). Representative specimens deposited in 
<collectionCode id="4CFCAE2D3E7C6A736D3FFE01486AFE72" box="[264,355,484,510]" pageId="13" pageNumber="14">BMNH</collectionCode>
, 
<collectionCode id="4CFCAE2D3E7C6A736D59FE0148CEFE72" box="[366,455,484,510]" pageId="13" pageNumber="14">CMNC</collectionCode>
, 
<collectionCode id="4CFCAE2D3E7C6A736DE4FE014B27FE72" box="[467,558,484,510]" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="13" pageNumber="14">CMNH</collectionCode>
, 
<collectionCode id="4CFCAE2D3E7C6A736E0DFE014B8AFE72" box="[570,643,484,510]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="13" pageNumber="14">CUIC</collectionCode>
, 
<collectionCode id="4CFCAE2D3E7C6A736EB9FE014BE8FE72" box="[654,737,484,510]" pageId="13" pageNumber="14">ESMC</collectionCode>
, 
<collectionCode id="4CFCAE2D3E7C6A736EDAFE014A4CFE72" LSID="urn:lsid:biocol.org:col:34795" box="[749,837,484,510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14">FMNH</collectionCode>
, 
<collectionCode id="4CFCAE2D3E7C6A736F67FE014A9BFE72" LSID="urn:lsid:biocol.org:col:33791" box="[848,914,484,510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="13" pageNumber="14">MCZ</collectionCode>
, 
<collectionCode id="4CFCAE2D3E7C6A736FA9FE014AE0FE72" box="[926,1001,484,510]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="14">NMW</collectionCode>
, 
<collectionCode id="4CFCAE2D3E7C6A736FC2FE014D47FE72" LSID="urn:lsid:biocol.org:col:34871" box="[1013,1102,484,510]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="14">USNM</collectionCode>
.
</paragraph>
<paragraph id="2A5236E83E7C6A736D0FFDEE4DF5FD12" blockId="13.[264,1323,284,1870]" pageId="13" pageNumber="14">
<emphasis id="1899EAFA3E7C6A736D0FFDEE4B58FDA9" bold="true" box="[312,593,523,549]" pageId="13" pageNumber="14">Differential Diagnosis.</emphasis>
Characterized by the pale anterolateral corners of the pronotum, the five spines on the mesosternal process, the mentum and stipes lighter than the venter of the head, and the green and/or blue maculae on the dorsum. Dorsal coloration sometimes similar to 
<taxonomicName id="EDED4D6B3E7C6A736E33FD614B88FD11" box="[516,641,644,669]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="1899EAFA3E7C6A736E33FD614B88FD11" box="[516,641,644,669]" italics="true" pageId="13" pageNumber="14">O. ornatus</emphasis>
</taxonomicName>
, but that species has hairs on the mesosternal process.
</paragraph>
</subSubSection>
<subSubSection id="62F765633E7C6A706D0FFD494A21FD4A" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="2A5236E83E7C6A736D0FFD494C08FC3A" blockId="13.[264,1323,284,1870]" pageId="13" pageNumber="14">
<emphasis id="1899EAFA3E7C6A736D0FFD494B90FD4A" bold="true" box="[312,665,684,710]" pageId="13" pageNumber="14">Description. Size and Form</emphasis>
. Length= 
<quantity id="ED159B0D3E7C6A736F21FD494A94FD4A" box="[790,925,684,710]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.3" metricValueMin="2.6" pageId="13" pageNumber="14" unit="mm" value="2.95" valueMax="3.3" valueMin="2.6">2.6–3.3mm</quantity>
. EL/EW=1.28. Oval, moderately convex. Elytra slightly longer than wide. 
<emphasis id="1899EAFA3E7C6A736F3EFD314A46FD62" bold="true" box="[777,847,724,750]" pageId="13" pageNumber="14">Color</emphasis>
. Dorsum black, with green iridescence on head, pronotum and broad lateral margins of elytra. Each elytron with 10–12 green to bluish green iridescent patches. Maxillary and labial palpi yellow. Mentum and stipes yellow to light brown, lighter than venter of head. Anterolateral corners of pronotum with pale spot. Sternum and ventrites brown. Legs yellow, with femora slightly darkened basally and along anterior edge. Epipleura and pseudoepipleura yellow to light brown.
</paragraph>
<paragraph id="2A5236E83E7C6A736D0FFC214C20F8C2" blockId="13.[264,1323,284,1870]" pageId="13" pageNumber="14">
<emphasis id="1899EAFA3E7C6A736D0FFC214872FC52" bold="true" box="[312,379,964,990]" pageId="13" pageNumber="14">Head</emphasis>
. Punctation on frons, clypeus and labrum moderately fine; distance between punctures 2– 
<date id="5E5310283E7C6A736D95FC0948A6FB8A" box="[418,431,1004,1030]" pageId="13" pageNumber="14">3</date>
x width of one puncture. Scape subequal in length as segments 2–5, segment 2 subequal or longer in length than segments 3–5, segments 3 and 4 subequal in length, apical segment 1.5x length of penultimate segment. Maxillary palpi ¾ width of labrum; segments 2 and 3 subequal with apical segment slightly longer. Labial palpi subequal to length of mentum; apical two segments subequal in length. Labrum with 2–3 setiferous systematic punctures centrally. Clypeus without detectable systematic punctures. Frons with irregular row of systematic punctures mesad of each eye. Mentum quadrate, anterior margin moderately convex; anterior half with several setiferous punctures. 
<emphasis id="1899EAFA3E7C6A736D3FFAC9486CFACA" bold="true" box="[264,357,1324,1350]" pageId="13" pageNumber="14">Thorax</emphasis>
. Dorsal punctation of pronotum moderately fine, slightly more impressed on elytra; uniform in size; distance between punctures 2– 
<date id="5E5310283E7C6A736FBEFAB14A9FFAE2" box="[905,918,1364,1390]" pageId="13" pageNumber="14">3</date>
x width of one puncture. Posterolateral corners of pronotum rounded. Lateral margin of pronotum with row of setiferous punctures; anterior and posterior series of setiferous systematic punctures moderately distinct, slightly larger then general punctation; setae recumbent. Rows of punctures on elytra indistinct and not generally detectable from surrounding punctation; several punctures bearing short recumbent seta. Lateral margin of elytra with row of setiferous punctures, denser in posterior half. Prosternum carinate medially, with two spines anteromedially which are as long as or slightly longer than narrowest width of prosternum. Mesosternal process about as long as wide, with 5 distinct spines positioned toward apex. Metasternum with oval glabrous area posteromedially, about as long as wide; less than half the total length of metasternum. Procoxae densely pubescent, some specimens with a few scattered spines or thickened setae. Protibia with 5–7 spines on dorsal face. Protarsal segments 1–4 small, subequal in length; segment 5 as long as segments 1–4. 
<collectingCountry id="52FA76783E7C6A73682EF8D14D5DF8C2" box="[1049,1108,1844,1870]" name="India" pageId="13" pageNumber="14">Hind</collectingCountry>
femora glabrous.
</paragraph>
<paragraph id="2A5236E83E7F6A706D3FFEFE4ADEFE5A" blockId="14.[264,1323,283,750]" pageId="14" pageNumber="15">
<emphasis id="1899EAFA3E7F6A706D3FFEFE488BFEB9" bold="true" box="[264,386,283,309]" pageId="14" pageNumber="15">Abdomen</emphasis>
. Ventrites sparsely pubescent; setae as long or longer as longest setae around margin of metasternal glabrous area. Fifth exposed ventrite entire. Aedeagus (
<figureCitation id="B2D62A6D3E7F6A70688BFEA1482FFE0A" captionStart="FIGURES 20 – 23" captionStartId="40.[264,382,1812,1836]" captionTargetBox="[329,1218,291,1800]" captionTargetId="figure@40.[264,1323,289,1801]" captionTargetPageId="40" captionText="FIGURES 20 – 23. Oocyclus aedeagi, dorsal views. Scale bars = 0.2 mm. – 20. O. maculatus Sharp (holotype). – 21. O. maculatus Sharp (specimen from Panama). – 22. O. muscus n. sp. (holotype). – 23. O. ornatus n. sp. (holotype)." httpUri="https://zenodo.org/record/158182/files/figure.png" pageId="14" pageNumber="15">Figs. 20– 21</figureCitation>
) with outer edges of parameres broadly curved with tips slightly curved inward; ventral sides much broader than dorsal sides. Tip of median lobe bifid, nearly reaching the apex of the parameres. Basal piece narrowed and rounded basally.
</paragraph>
<paragraph id="2A5236E83E7F6A706D0FFE064A21FD4A" blockId="14.[264,1323,283,750]" pageId="14" pageNumber="15">
<emphasis id="1899EAFA3E7F6A706D0FFE0648BDFE71" bold="true" box="[312,436,483,509]" pageId="14" pageNumber="15">Variation:</emphasis>
<taxonomicName id="EDED4D6B3E7F6A706D89FE014B53FE71" box="[446,602,484,509]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="1899EAFA3E7F6A706D89FE014B53FE71" box="[446,602,484,509]" italics="true" pageId="14" pageNumber="15">O. maculatus</emphasis>
</taxonomicName>
expresses the most variation of any of the species here examined. The dorsal coloration and iridescence varies from a few, very subdued green maculae to a number of bright green or blue maculae similar to 
<taxonomicName id="EDED4D6B3E7F6A706F8FFDD14D3FFDC1" box="[952,1078,564,589]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="1899EAFA3E7F6A706F8FFDD14D3FFDC1" box="[952,1078,564,589]" italics="true" pageId="14" pageNumber="15">O. ornatus</emphasis>
</taxonomicName>
. The marginal band of green iridescence on each elytron may also very in intensity. The shape of the parameres also exhibits some variation (
<figureCitation id="B2D62A6D3E7F6A706EDBFD614A77FD12" box="[748,894,644,670]" captionStart="FIGURES 20 – 23" captionStartId="40.[264,382,1812,1836]" captionTargetBox="[329,1218,291,1800]" captionTargetId="figure@40.[264,1323,289,1801]" captionTargetPageId="40" captionText="FIGURES 20 – 23. Oocyclus aedeagi, dorsal views. Scale bars = 0.2 mm. – 20. O. maculatus Sharp (holotype). – 21. O. maculatus Sharp (specimen from Panama). – 22. O. muscus n. sp. (holotype). – 23. O. ornatus n. sp. (holotype)." httpUri="https://zenodo.org/record/158182/files/figure.png" pageId="14" pageNumber="15">Figs. 20–21</figureCitation>
). It is possible that this taxon may represent a complex of closely related species.
</paragraph>
</subSubSection>
<subSubSection id="62F765633E7F6A706D0FFD314A69FD62" box="[312,864,724,750]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="2A5236E83E7F6A706D0FFD314A69FD62" blockId="14.[264,1323,283,750]" box="[312,864,724,750]" pageId="14" pageNumber="15">
<emphasis id="1899EAFA3E7F6A706D0FFD3148D8FD62" bold="true" box="[312,465,724,750]" pageId="14" pageNumber="15">Distribution</emphasis>
. 
<collectingCountry id="52FA76783E7F6A706DE8FD314B68FD62" box="[479,609,724,750]" name="Guatemala" pageId="14" pageNumber="15">Guatemala</collectingCountry>
, 
<collectingCountry id="52FA76783E7F6A706E5AFD314BF9FD62" box="[621,752,724,750]" name="Costa Rica" pageId="14" pageNumber="15">Costa Rica</collectingCountry>
, 
<collectingCountry id="52FA76783E7F6A706ECCFD314A55FD62" box="[763,860,724,750]" name="Panama" pageId="14" pageNumber="15">Panamá</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>