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<document id="B77037603F950850E3D99141DE135371" ID-DOI="10.5281/zenodo.158165" ID-GBIF-Dataset="fb07d896-f75a-4f51-aeb0-2700da02c70e" ID-ISSN="1175­5326" ID-Zenodo-Dep="158165" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038221360" checkinUser="plazi" docAuthor="Short, A. E. Z. &amp; Perkins, P. D." docDate="2004" docId="A24487FE3E7B6A726D3FFCA94D1BFDAA" docLanguage="en" docName="zt00783.pdf" docOrigin="Zootaxa 783" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Oocyclus grandis Short &amp; Perkins, 2004, n. sp." docType="treatment" docVersion="10" lastPageNumber="13" masterDocId="5E7DFF863E716A7E6C37FFE54909FF8C" masterDocTitle="A revision of Oocyclus Sharp of Mexico and Central America (Coleoptera: Hydrophilidae)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="11" updateTime="1698574933192" updateUser="plazi">
<mods:mods id="EDC926AB9BC1AD02493D0F09955D728B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F6BE70383467E3EE3047D8BFBDB017F7">
<mods:title id="AE5C5BE5C40F496697E84F09F2E0FA13">A revision of Oocyclus Sharp of Mexico and Central America (Coleoptera: Hydrophilidae)</mods:title>
</mods:titleInfo>
<mods:name id="1A821E0022BA011E624561875EF6F857" type="personal">
<mods:role id="23C04A08B1C6A0960EA5800D8FBB3CD7">
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</mods:role>
<mods:namePart id="C9947200F031832A0FEC4442CBE8B721">Short, A. E. Z.</mods:namePart>
</mods:name>
<mods:name id="00D2E4B567603AF652D8ACC328BBB1EC" type="personal">
<mods:role id="58F98A49B1EA2681AE1DD83AD25EC44C">
<mods:roleTerm id="A936543AD60452885E3065BBC93B9B1A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD37257D04230C0B96E2F9682528835C">Perkins, P. D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2445733AB0A14A5E2CAD1E2082D2C77E">text</mods:typeOfResource>
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<mods:title id="0999B1363F24FA3FCC183A0A5824FD64">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="068A5D19F98FDDA3A49936F6C43EA949">
<mods:date id="B412CF04B800651557C5E291593A6EAF">2004</mods:date>
<mods:detail id="17E15D668D941086ADB3D0784D2F188C" type="volume">
<mods:number id="CAFEADA927A29BAEABB4354DCCE60C0C">783</mods:number>
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</mods:extent>
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</mods:relatedItem>
<mods:classification id="D8875695BD2C5969D2403BB2119546F0">journal article</mods:classification>
<mods:identifier id="D13AFE6940EC430C9661FE5E0349BF15" type="DOI">10.5281/zenodo.158165</mods:identifier>
<mods:identifier id="55F9A1A8AF4F5A5FEA037882690A3A08" type="GBIF-Dataset">fb07d896-f75a-4f51-aeb0-2700da02c70e</mods:identifier>
<mods:identifier id="C65B5D48E2E08B31D9EBA14027DCEAB2" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="B01849A12F703C184D3085D5C7111001" type="Zenodo-Dep">158165</mods:identifier>
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</mods:mods>
<treatment id="A24487FE3E7B6A726D3FFCA94D1BFDAA" ID-DOI="http://doi.org/10.5281/zenodo.6269927" ID-GBIF-Taxon="119609252" ID-Zenodo-Dep="6269927" LSID="urn:lsid:plazi:treatment:A24487FE3E7B6A726D3FFCA94D1BFDAA" httpUri="http://treatment.plazi.org/id/A24487FE3E7B6A726D3FFCA94D1BFDAA" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="62F765633E7B6A746D3FFCA948BDFC05" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="2A5236E83E7B6A746D3FFCA94B2AFCEA" blockId="10.[264,547,844,905]" box="[264,547,844,870]" pageId="10" pageNumber="11">
<heading id="711A81843E7B6A746D3FFCA94B2AFCEA" bold="true" box="[264,547,844,870]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="1899EAFA3E7B6A746D3FFCA94B2AFCEA" bold="true" box="[264,547,844,870]" pageId="10" pageNumber="11">
<taxonomicName id="EDED4D6B3E7B6A746D3FFCA948D0FCEA" ID-CoL="49RVT" box="[264,473,844,870]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="1899EAFA3E7B6A746D3FFCA948D0FCEA" bold="true" box="[264,473,844,870]" italics="true" pageId="10" pageNumber="11">Oocyclus grandis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="03AA57813E7B6A746DD6FCA94B2AFCEA" box="[481,547,844,870]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2A5236E83E7B6A746D3FFC8A48BDFC05" blockId="10.[264,547,844,905]" box="[264,436,879,905]" pageId="10" pageNumber="11">
(
<figureCitation id="B2D62A6D3E7B6A746D26FC8A488BFC05" box="[273,386,879,905]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStartId-0="29.[264,368,1771,1795]" captionStartId-1="30.[264,368,1707,1731]" captionTargetBox-0="[327,1284,334,1710]" captionTargetBox-1="[264,1322,328,1683]" captionTargetId-0="figure@29.[264,1323,308,1747]" captionTargetId-1="figure@30.[264,1323,308,1683]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURE 6. Oocyclus grandis n. sp. holotype. Dorsal and lateral habitus." captionText-1="FIGURE 7. Oocyclus grandis n. sp. holotype. Ventral habitus." httpUri-0="https://zenodo.org/record/169615/files/figure.png" httpUri-1="https://zenodo.org/record/169616/files/figure.png" pageId="10" pageNumber="11">Figs. 6–7</figureCitation>
,
<figureCitation id="B2D62A6D3E7B6A746DBAFC8A48A2FC05" box="[397,427,879,905]" captionStart="FIGURES 16 – 19" captionStartId="39.[264,382,1803,1827]" captionTargetBox="[383,1201,291,1793]" captionTargetId="figure@39.[264,1323,289,1798]" captionTargetPageId="39" captionText="FIGURES 16 – 19. Oocyclus aedeagi, dorsal views. Scale bars = 0.2 mm. – 16. O. brevicornis Sharp (specimen from Guatemala). – 17. O. catarata n. sp. (holotype). – 18. O. funestus n. sp. (holotype). – 19. O. grandis n. sp. (holotype)." httpUri="https://zenodo.org/record/158181/files/figure.png" pageId="10" pageNumber="11">19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62F765633E7B6A756D3FFC5B4A1AFED2" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="2A5236E83E7B6A746D3FFC5B4B41FB8D" blockId="10.[264,1324,958,1905]" pageId="10" pageNumber="11">
<emphasis id="1899EAFA3E7B6A746D3FFC5B48B2FC54" bold="true" box="[264,443,958,984]" pageId="10" pageNumber="11">
<typeStatus id="F556884A3E7B6A746D3FFC5B484CFC54" box="[264,325,958,984]" pageId="10" pageNumber="11">Type</typeStatus>
Locality.
</emphasis>
Waterfall on the Río Carmona,
<quantity id="ED159B0D3E7B6A746F7BFC5A4A82FC55" box="[844,907,959,985]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="km" value="2.0">2 km</quantity>
N. of Carmona, Guanacaste Province,
<collectingCountry id="52FA76783E7B6A746D70FC0248C3FB8D" box="[327,458,999,1025]" name="Costa Rica" pageId="10" pageNumber="11">Costa Rica</collectingCountry>
,
<quantity id="ED159B0D3E7B6A746DE1FC024B03FB8D" box="[470,522,999,1025]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="m" value="30.0">30m</quantity>
elev.
</paragraph>
<paragraph id="2A5236E83E7B6A756D0FFBEB4A1AFED2" blockId="10.[264,1324,958,1905]" lastBlockId="11.[264,1324,284,1870]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="1899EAFA3E7B6A746D0FFBEB48F9FBA4" bold="true" box="[312,496,1038,1064]" pageId="10" pageNumber="11">
<typeStatus id="F556884A3E7B6A746D0FFBEB487CFBA4" box="[312,373,1038,1064]" pageId="10" pageNumber="11">Type</typeStatus>
Material.
</emphasis>
<typeStatus id="F556884A3E7B6A746DC9FBEA4B66FBA5" box="[510,623,1039,1065]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
. Male: “
<collectingCountry id="52FA76783E7B6A746EF8FBEA4A8BFBA5" box="[719,898,1039,1065]" name="Costa Rica" pageId="10" pageNumber="11">COSTA RICA</collectingCountry>
: Guanacaste Prov./Nandayure;
<quantity id="ED159B0D3E7B6A7468D9FBEA4C23FBA5" box="[1262,1322,1039,1065]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="km" value="2.0">2 km</quantity>
N of Carmona/ Waterfall margin; w/insecticide/ alt.
<quantity id="ED159B0D3E7B6A746FBEFBD24AC8FBDD" box="[905,961,1079,1105]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="m" value="30.0">30m</quantity>
;
<date id="5E5310283E7B6A746FE6FBD24D6AFBDD" box="[977,1123,1079,1105]" pageId="10" pageNumber="11" value="2003-01-15">15­Jan­2003</date>
(CR­03­06)/ A. Short, R. Roughley, W. Porras”, “
<typeStatus id="F556884A3E7B6A746E98FBBA4A45FBF5" box="[687,844,1119,1145]" pageId="10" pageNumber="11" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="EDED4D6B3E7B6A746F54FBBA4AD8FBF5" box="[867,977,1119,1145]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Oocyclus</taxonomicName>
/
<taxonomicName id="EDED4D6B3E7B6A746FDFFBBA4D34FBF5" box="[1000,1085,1119,1145]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="grandis">grandis</taxonomicName>
/ Short &amp; Perkins” (
<collectionCode id="4CFCAE2D3E7B6A746D27FB624861FB2D" box="[272,360,1159,1185]" pageId="10" pageNumber="11">INBIO</collectionCode>
).
<emphasis id="1899EAFA3E7B6A746DBFFB634AF8FB2C" bold="true" box="[392,1009,1158,1184]" pageId="10" pageNumber="11">
<typeStatus id="F556884A3E7B6A746DBFFB634B0CFB2C" box="[392,517,1158,1184]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
(383).
<collectingCountry id="52FA76783E7B6A746E5EFB634A2CFB2C" box="[617,805,1158,1184]" name="Costa Rica" pageId="10" pageNumber="11">COSTA RICA</collectingCountry>
: Alajuela Prov.,
</emphasis>
<quantity id="ED159B0D3E7B6A746FCCFB624D47FB2D" box="[1019,1102,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="6.1" pageId="10" pageNumber="11" unit="km" value="6.1">6.1 km</quantity>
N. Los Lagos, on Los Lagos­Colonia Rd.,
<quantity id="ED159B0D3E7B6A746E04FB4A4B7CFB45" box="[563,629,1199,1225]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="10" pageNumber="11" unit="m" value="870.0">870m</quantity>
elev., seeps on Rio San Lorenzo,
<date id="5E5310283E7B6A746830FB4A4D71FB45" box="[1031,1144,1199,1225]" pageId="10" pageNumber="11" value="2003-01-12">12.i.2003</date>
, A. Short &amp; R. Roughley leg., CR­03­04 (1
<collectionCode id="4CFCAE2D3E7B6A746E68FB324BBCFB7D" box="[607,693,1239,1265]" pageId="10" pageNumber="11">INBIO</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746EFCFB334AB9FB7C" bold="true" box="[715,944,1238,1264]" pageId="10" pageNumber="11">Guanacaste Prov.,</emphasis>
Cerro Azul, road between Carmona &amp; Bella Vista,
<quantity id="ED159B0D3E7B6A746E3FFB1A4B42FA95" box="[520,587,1279,1305]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="m" value="800.0">800m</quantity>
elev.,
<date id="5E5310283E7B6A746EAAFB1A4A07FA95" box="[669,782,1279,1305]" pageId="10" pageNumber="11" value="2003-01-16">16.i.2003</date>
, waterfall/seep, A. Short, R. Roughley &amp; W. Porras, leg. (15
<collectionCode id="4CFCAE2D3E7B6A746DF0FAC24B1AFACD" box="[455,531,1319,1345]" pageId="10" pageNumber="11">AEZS</collectionCode>
, 28
<collectionCode id="4CFCAE2D3E7B6A746E70FAC24B94FACD" box="[583,669,1319,1345]" pageId="10" pageNumber="11">INBIO</collectionCode>
); same data as
<typeStatus id="F556884A3E7B6A746F64FAC24A8CFACD" box="[851,901,1319,1345]" pageId="10" pageNumber="11">type</typeStatus>
(8
<collectionCode id="4CFCAE2D3E7B6A746F87FAC24AF5FACD" box="[944,1020,1319,1345]" pageId="10" pageNumber="11">AEZS</collectionCode>
, 15
<collectionCode id="4CFCAE2D3E7B6A746807FAC24D8FFACD" box="[1072,1158,1319,1345]" pageId="10" pageNumber="11">INBIO</collectionCode>
);
<emphasis id="1899EAFA3E7B6A7468ABFAC34840FAE4" bold="true" pageId="10" pageNumber="11">Puntarenas Prov.</emphasis>
,
<quantity id="ED159B0D3E7B6A746D6AFAAA48BAFAE5" box="[349,435,1359,1385]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="km" value="1.2">1.2 km</quantity>
SE of Coronado,
<date id="5E5310283E7B6A746EAEFAAA4A10FAE5" box="[665,793,1359,1385]" pageId="10" pageNumber="11" value="2003-06-18">18.vi.2003</date>
, waterfall, A. E. Z. Short leg. (1
<collectionCode id="4CFCAE2D3E7B6A7468F7FAAA4C05FAE5" box="[1216,1292,1359,1385]" pageId="10" pageNumber="11">AEZS</collectionCode>
, 6
<collectionCode id="4CFCAE2D3E7B6A746D3FFA924857FA1D" box="[264,350,1399,1425]" pageId="10" pageNumber="11">INBIO</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746D41FA934B20FA1C" bold="true" box="[374,553,1398,1424]" pageId="10" pageNumber="11">San Jose Prov</emphasis>
.,
<quantity id="ED159B0D3E7B6A746E75FA924BAFFA1D" box="[578,678,1399,1425]" metricMagnitude="4" metricUnit="m" metricValue="1.91" pageId="10" pageNumber="11" unit="km" value="19.1">19.1 km</quantity>
NE of Dominical on rt. 243,
<date id="5E5310283E7B6A746824FA924D9BFA1D" box="[1043,1170,1399,1425]" pageId="10" pageNumber="11" value="2003-06-18">18.vi.2003</date>
, waterfall &amp; seeps, A. E. Z. Short leg. (1
<collectionCode id="4CFCAE2D3E7B6A746E6AFA7A4BBDFA35" box="[605,692,1439,1465]" pageId="10" pageNumber="11">INBIO</collectionCode>
).
<emphasis id="1899EAFA3E7B6A746EFFFA7B4A81FA34" bold="true" box="[712,904,1438,1464]" pageId="10" pageNumber="11">
<collectingCountry id="52FA76783E7B6A746EFFFA7B4A81FA34" box="[712,904,1438,1464]" name="Guatemala" pageId="10" pageNumber="11">GUATEMALA</collectingCountry>
</emphasis>
:
<emphasis id="1899EAFA3E7B6A746FAEFA7B4D4AFA34" bold="true" box="[921,1091,1438,1464]" pageId="10" pageNumber="11">Baja Verapaz</emphasis>
,
<quantity id="ED159B0D3E7B6A746864FA7A4D92FA35" box="[1107,1179,1439,1465]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="10" pageNumber="11" unit="mi" value="20.0">20 mi</quantity>
. S. Rabinal,
<date id="5E5310283E7B6A746D3FFA224881FA6D" box="[264,392,1479,1505]" pageId="10" pageNumber="11" value="1974-06-12">12.vi.1974</date>
, M. E. &amp; P. D. Perkins leg. (3
<collectionCode id="4CFCAE2D3E7B6A746EC1FA224A59FA6D" LSID="urn:lsid:biocol.org:col:34871" box="[758,848,1479,1505]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746F52FA234AF2FA6C" bold="true" box="[869,1019,1478,1504]" pageId="10" pageNumber="11">Chiquimula</emphasis>
,
<quantity id="ED159B0D3E7B6A74683CFA224D4DFA6D" box="[1035,1092,1479,1505]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="10" pageNumber="11" unit="mi" value="4.0">4 mi</quantity>
. N. Quezaltepeque,
<date id="5E5310283E7B6A746D3FFA0A488EF985" box="[264,391,1519,1545]" pageId="10" pageNumber="11" value="1974-06-17">17.vi.1974</date>
, M. E. &amp; P. D. Perkins leg. (2
<collectionCode id="4CFCAE2D3E7B6A746EC1FA0A4A59F985" LSID="urn:lsid:biocol.org:col:34871" box="[758,848,1519,1545]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746F52FA0B4AB6F984" bold="true" box="[869,959,1518,1544]" pageId="10" pageNumber="11">Jalapa,</emphasis>
<quantity id="ED159B0D3E7B6A746F88FA0A4D0EF985" box="[959,1031,1519,1545]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="10" pageNumber="11" unit="mi" value="12.0">12 mi</quantity>
. NW Jalapa,
<date id="5E5310283E7B6A746891FA0A4C2FF985" box="[1190,1318,1519,1545]" pageId="10" pageNumber="11" value="1974-06-15">15.vi.1974</date>
, M. E. &amp; P. D. Perkins leg. (38
<collectionCode id="4CFCAE2D3E7B6A746E59F9F24BC1F9BD" LSID="urn:lsid:biocol.org:col:34871" box="[622,712,1559,1585]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746EEBF9F34A98F9BC" bold="true" box="[732,913,1558,1584]" pageId="10" pageNumber="11">Suchitepequez</emphasis>
,
<quantity id="ED159B0D3E7B6A746F90F9F24AE9F9BD" box="[935,992,1559,1585]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="10" pageNumber="11" unit="mi" value="1.0">1 mi</quantity>
. W. Cocales,
<date id="5E5310283E7B6A74684AF9F24DF4F9BD" box="[1149,1277,1559,1585]" pageId="10" pageNumber="11" value="1974-06-26">26.vi.1974</date>
, M. E. &amp; P. D. Perkins leg (19
<collectionCode id="4CFCAE2D3E7B6A746E71F9DA4BA9F9D5" LSID="urn:lsid:biocol.org:col:34871" box="[582,672,1599,1625]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746E8AF9DB4A6CF9D4" bold="true" box="[701,869,1598,1624]" pageId="10" pageNumber="11">
<collectingCountry id="52FA76783E7B6A746E8AF9DB4A6CF9D4" box="[701,869,1598,1624]" name="Honduras" pageId="10" pageNumber="11">HONDURAS</collectingCountry>
</emphasis>
:
<emphasis id="1899EAFA3E7B6A746F42F9DB4D02F9D4" bold="true" box="[885,1035,1598,1624]" pageId="10" pageNumber="11">Comayagua</emphasis>
,
<quantity id="ED159B0D3E7B6A74682CF9DA4D5DF9D5" box="[1051,1108,1599,1625]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="10" pageNumber="11" unit="mi" value="6.0">6 mi</quantity>
. N. Siguatepeque,
<date id="5E5310283E7B6A746D3FF9824881F90D" box="[264,392,1639,1665]" pageId="10" pageNumber="11" value="1974-06-20">20.vi.1974</date>
, M. E. &amp; P. D. Perkins leg. (17
<collectionCode id="4CFCAE2D3E7B6A746F35F9824A55F90D" LSID="urn:lsid:biocol.org:col:34871" box="[770,860,1639,1665]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746F47F9834D0DF90C" bold="true" box="[880,1028,1638,1664]" pageId="10" pageNumber="11">Ocotepeque</emphasis>
,
<quantity id="ED159B0D3E7B6A746823F9824D53F90D" box="[1044,1114,1639,1665]" metricMagnitude="4" metricUnit="m" metricValue="1.7702784" pageId="10" pageNumber="11" unit="mi" value="11.0">11 mi</quantity>
. S. Nueva Ocotepeque,
<date id="5E5310283E7B6A746D69F96A48D4F925" box="[350,477,1679,1705]" pageId="10" pageNumber="11" value="1974-06-17">17.vi.1974</date>
, M. E. &amp; P. D. Perkins leg. (22
<collectionCode id="4CFCAE2D3E7B6A746F6AF96A4ABEF925" LSID="urn:lsid:biocol.org:col:34871" box="[861,951,1679,1705]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746FFAF96B4D41F924" bold="true" box="[973,1096,1678,1704]" pageId="10" pageNumber="11">
<collectingCountry id="52FA76783E7B6A746FFAF96B4D41F924" box="[973,1096,1678,1704]" name="Mexico" pageId="10" pageNumber="11">MEXICO</collectingCountry>
</emphasis>
:
<emphasis id="1899EAFA3E7B6A74686EF96B4DA4F924" bold="true" box="[1113,1197,1678,1704]" pageId="10" pageNumber="11">Jalisco</emphasis>
,
<quantity id="ED159B0D3E7B6A746889F96A4C0EF925" box="[1214,1287,1679,1705]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="10" pageNumber="11" unit="mi" value="12.0">12 mi</quantity>
. S. Puerto Vallarta,
<date id="5E5310283E7B6A746DFDF9524B59F95D" box="[458,592,1719,1745]" pageId="10" pageNumber="11" value="1974-07-21">21.vii.1974</date>
, M. E. &amp; P. D. Perkins leg. (22
<collectionCode id="4CFCAE2D3E7B6A746FD9F9524D41F95D" LSID="urn:lsid:biocol.org:col:34871" box="[1006,1096,1719,1745]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<quantity id="ED159B0D3E7B6A746868F9524D93F95D" box="[1119,1178,1719,1745]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="10" pageNumber="11" unit="mi" value="5.0">5 mi</quantity>
. W. Plan de Barrancas,
<date id="5E5310283E7B6A746DA4F93A4B12F975" box="[403,539,1759,1785]" pageId="10" pageNumber="11" value="1954-07-21">21.vii.1954</date>
, J. MacSwain leg. (6
<collectionCode id="4CFCAE2D3E7B6A746F0CF93A4A9CF975" LSID="urn:lsid:biocol.org:col:34871" box="[827,917,1759,1785]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746F87F93B4D19F974" bold="true" box="[944,1040,1758,1784]" pageId="10" pageNumber="11">Nayarit</emphasis>
, Tepic,
<date id="5E5310283E7B6A74684BF93A4DF4F975" box="[1148,1277,1759,1785]" pageId="10" pageNumber="11" value="1953-09-15">15.ix.1953</date>
, B. Malkin leg. (24
<collectionCode id="4CFCAE2D3E7B6A746DF0F8E24B0AF8AD" LSID="urn:lsid:biocol.org:col:34700" box="[455,515,1799,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="10" pageNumber="11">CAS</collectionCode>
, 19
<collectionCode id="4CFCAE2D3E7B6A746E02F8E24B86F8AD" LSID="urn:lsid:biocol.org:col:34871" box="[565,655,1799,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
); same locality,
<date id="5E5310283E7B6A746F79F8E24AC7F8AD" box="[846,974,1799,1825]" pageId="10" pageNumber="11" value="1953-09-23">23.ix.1953</date>
, B. Malkin, leg. (35
<collectionCode id="4CFCAE2D3E7B6A7468F2F8E24C08F8AD" LSID="urn:lsid:biocol.org:col:34700" box="[1221,1281,1799,1825]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="10" pageNumber="11">CAS</collectionCode>
, 25
<collectionCode id="4CFCAE2D3E7B6A746D3FF8CA486BF8C5" LSID="urn:lsid:biocol.org:col:34871" box="[264,354,1839,1865]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746D4CF8CB48D0F8C4" bold="true" box="[379,473,1838,1864]" pageId="10" pageNumber="11">Oaxaca</emphasis>
,
<quantity id="ED159B0D3E7B6A746DDBF8CA4B20F8C5" box="[492,553,1839,1865]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="10" pageNumber="11" unit="mi" value="8.0">8 mi</quantity>
. S. Valle Nacional,
<date id="5E5310283E7B6A746F16F8CA4A93F8C5" box="[801,922,1839,1865]" pageId="10" pageNumber="11" value="1974-07-06">6.vii.1974</date>
, M. E. &amp; P. D. Perkins leg. (75
<collectionCode id="4CFCAE2D3E7B6A746D3FF8B2486BF8FD" LSID="urn:lsid:biocol.org:col:34871" box="[264,354,1879,1905]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11">USNM</collectionCode>
);
<emphasis id="1899EAFA3E7B6A746D4FF8B348E3F8FC" bold="true" box="[376,490,1878,1904]" pageId="10" pageNumber="11">Veracruz</emphasis>
, Catemaco (
<quantity id="ED159B0D3E7B6A746EB5F8B24BC9F8FD" box="[642,704,1879,1905]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="11" unit="km" value="8.0">8 km</quantity>
SE), El Salto Eyipantla,
<date id="5E5310283E7B6A746FDAF8B24D89F8FD" box="[1005,1152,1879,1905]" pageId="10" pageNumber="11" value="1990-03-null">30.iii. 1990</date>
, Paul J. Spangler leg. (1
<collectionCode id="4CFCAE2D3E7A6A756D93FEF948F7FEBA" LSID="urn:lsid:biocol.org:col:34871" box="[420,510,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12">USNM</collectionCode>
). Representative specimens deposited in
<collectionCode id="4CFCAE2D3E7A6A756837FEF94D45FEBA" box="[1024,1100,284,310]" pageId="11" pageNumber="12">AEZS</collectionCode>
,
<collectionCode id="4CFCAE2D3E7A6A756868FEF94DB3FEBA" box="[1119,1210,284,310]" pageId="11" pageNumber="12">BMNH</collectionCode>
,
<collectionCode id="4CFCAE2D3E7A6A7568FAFEF94C2EFEBA" box="[1229,1319,284,310]" pageId="11" pageNumber="12">CMNC</collectionCode>
,
<collectionCode id="4CFCAE2D3E7A6A756D3FFEA1486AFED2" box="[264,355,324,350]" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="11" pageNumber="12">CMNH</collectionCode>
,
<collectionCode id="4CFCAE2D3E7A6A756D59FEA148BEFED2" box="[366,439,324,350]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="11" pageNumber="12">CUIC</collectionCode>
,
<collectionCode id="4CFCAE2D3E7A6A756DF4FEA14B1FFED2" box="[451,534,324,350]" pageId="11" pageNumber="12">ESMC</collectionCode>
,
<collectionCode id="4CFCAE2D3E7A6A756E16FEA14B70FED2" LSID="urn:lsid:biocol.org:col:34795" box="[545,633,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="12">FMNH</collectionCode>
, and
<collectionCode id="4CFCAE2D3E7A6A756E81FEA14A0FFED2" box="[694,774,324,350]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="12">NMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="62F765633E7A6A726D0FFE8E4A73FE5A" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="2A5236E83E7A6A756D0FFE8E4D59FE5A" blockId="11.[264,1324,284,1870]" pageId="11" pageNumber="12">
<emphasis id="1899EAFA3E7A6A756D0FFE8E48BEFE09" bold="true" box="[312,439,363,389]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
The largest species of
<taxonomicName id="EDED4D6B3E7A6A756EE3FE894A48FE09" box="[724,833,364,389]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="1899EAFA3E7A6A756EE3FE894A48FE09" box="[724,833,364,389]" italics="true" pageId="11" pageNumber="12">Oocyclus</emphasis>
</taxonomicName>
in Central
<collectingCountry id="52FA76783E7A6A756FFFFE894D39FE0A" box="[968,1072,364,390]" name="United States of America" pageId="11" pageNumber="12">America</collectingCountry>
, only
<taxonomicName id="EDED4D6B3E7A6A75684DFE894C07FE09" box="[1146,1294,364,389]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="substillus">
<emphasis id="1899EAFA3E7A6A75684DFE894C07FE09" box="[1146,1294,364,389]" italics="true" pageId="11" pageNumber="12">O. substillus</emphasis>
</taxonomicName>
is similar in size but the latter has spinose posterior pronotal apices. The long mesosternal process, which bears 6–8 distinct spines, also characterizes this species.
</paragraph>
<paragraph id="2A5236E83E7A6A726D0FFE064A73FE5A" blockId="11.[264,1324,284,1870]" lastBlockId="12.[264,1323,284,550]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="1899EAFA3E7A6A756D0FFE0648F8FE71" bold="true" box="[312,497,483,509]" pageId="11" pageNumber="12">Size and Form</emphasis>
. Length=
<quantity id="ED159B0D3E7A6A756E59FE014BFFFE72" box="[622,758,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.1" metricValueMin="4.2" pageId="11" pageNumber="12" unit="mm" value="4.65" valueMax="5.1" valueMin="4.2">4.2–5.1mm</quantity>
. EL/EW=1.13. Elongate oval; moderately convex, elytra longer than wide; disc of elytra more convex transversely than longitudinally.
<emphasis id="1899EAFA3E7A6A756D3FFDD6485DFDC1" bold="true" box="[264,340,563,589]" pageId="11" pageNumber="12">Color.</emphasis>
Dorsum generally black; anterolateral corners of pronotum distinctly yellow to white. Elytra with irregular patches of bronze or green iridescence. Maxillary and labial palpi yellow. Mentum and stipes brown, slightly paler than venter of head. Sternum brown to dark brown. Base of femora dark brown, gradually becoming light brown distally; tibia and tarsi light brown. Epipleura dark brown; pseudoepileura yellow. Each ventrite brown with light brown patches laterally.
<emphasis id="1899EAFA3E7A6A756F7EFD194A85FC9A" bold="true" box="[841,908,764,790]" pageId="11" pageNumber="12">Head</emphasis>
. Punctation on frons, clypeus and frons uniformly fine; punctures separated by 2–3 widths of one puncture. Antennae with scape subequal in length to segments 2–5, segment 2 subequal to segments 3–5, segments 4–5 equal in length; first two segments of club equal in size, third segment 1.5x longer and moderately compressed laterally. Maxillary palpi slightly shorter than width of labrum; segments 2 and 3 subequal in length, apical segment slightly longer. Labial palpi ¾ length of mentum; apical two segments subequal in length. Labrum without detectable systematic punctures. Clypeus with 4–5 nearly indistinct systematic punctures near each anterolateral margin. Frons with an irregular row of systematic punctures mesad of each eye bearing short recumbent setae. Mentum quadrate, anterior margin slightly convex, with scattered spare punctures of mixed size; slightly rugose anteriorly.
<emphasis id="1899EAFA3E7A6A75682AFB694D73FB2A" bold="true" box="[1053,1146,1164,1190]" pageId="11" pageNumber="12">Thorax</emphasis>
. Dorsal punctation of pronotum and elytra extremely fine, almost obsolete. Posterolateral corners of pronotum angulate but not spinose; lateral margins of pronotum lined with a row of setiferous punctures. Pronotal systematic punctures bearing short recumbent setae, with anterior series forming an irregular row, posterior series forming a diffuse field. Sutural punctation of elytra extremely fine and irregular, not bearing setae. Elytra with six rows of setiferous punctures; first row with punctures very dense anteriorly and becoming more diffuse posteriorly, setae erect on anterior half; rows 2–4 with punctures more irregularly spaced, suberect to recumbent; row 5 irregular and diffuse; row six along lateral edge of elytra, usually recumbent. Lateral margins of elytra slightly explanate anteriorly. Prosternum carinate centrally, with two coarse spines anteromedially. Mesosternal process about
<date id="5E5310283E7A6A756D3FF9A1482CF9D2" box="[264,293,1604,1630]" pageId="11" pageNumber="12">3x</date>
longer than wide, with seven coarse spines (occasionally six or eight spines); anterior margin of process before spines thin. Metasternum with elongate­oval glabrous area posteromedially, about
<date id="5E5310283E7A6A756DCAF9714B12F922" box="[509,539,1684,1710]" pageId="11" pageNumber="12">2x</date>
longer than wide; about 2/3 width of metasternum. Procoxae pubescent and bearing a number of distinct spines, slightly smaller than those of prosternum. Protibia with 13–15 spines on dorsal face. Protarsal segments 1–4 small, subequal in length; segment 5 as long as segments 1–4; protarsal claws of males slightly enlarged.
<collectingCountry id="52FA76783E7A6A756D3FF8D1484AF8C2" box="[264,323,1844,1870]" name="India" pageId="11" pageNumber="12">Hind</collectingCountry>
femora glabrous.
<emphasis id="1899EAFA3E7A6A756E00F8D14BB8F8C2" bold="true" box="[567,689,1844,1870]" pageId="11" pageNumber="12">Abdomen</emphasis>
. Ventrites with dense, uniform pubescence, usually suberect to recumbent; longest setae shorter to subequal in length to longest setae around margin of metasternal glabrous area. Fifth exposed ventrite entire. Aedeagus (
<figureCitation id="B2D62A6D3E7D6A7268FDFEA14C2BFED2" box="[1226,1314,324,350]" captionStart="FIGURES 16 – 19" captionStartId="39.[264,382,1803,1827]" captionTargetBox="[383,1201,291,1793]" captionTargetId="figure@39.[264,1323,289,1798]" captionTargetPageId="39" captionText="FIGURES 16 – 19. Oocyclus aedeagi, dorsal views. Scale bars = 0.2 mm. – 16. O. brevicornis Sharp (specimen from Guatemala). – 17. O. catarata n. sp. (holotype). – 18. O. funestus n. sp. (holotype). – 19. O. grandis n. sp. (holotype)." httpUri="https://zenodo.org/record/158181/files/figure.png" pageId="12" pageNumber="13">Fig. 19</figureCitation>
) with both dorsal and ventral sides curved; ventral side moderately broader than dorsal side; tips curved inward. Tip of median lobe bifid, not reaching apex of parameres. Basal piece slightly narrowed, evenly and broadly rounded.
</paragraph>
</subSubSection>
<subSubSection id="62F765633E7D6A726D0FFE064AD7FE72" box="[312,990,483,510]" pageId="12" pageNumber="13" type="distribution">
<paragraph id="2A5236E83E7D6A726D0FFE064AD7FE72" blockId="12.[264,1323,284,550]" box="[312,990,483,510]" pageId="12" pageNumber="13">
<emphasis id="1899EAFA3E7D6A726D0FFE0648D1FE71" bold="true" box="[312,472,483,509]" pageId="12" pageNumber="13">Distribution.</emphasis>
<collectingCountry id="52FA76783E7D6A726DE8FE014B35FE72" box="[479,572,484,510]" name="Mexico" pageId="12" pageNumber="13">México</collectingCountry>
,
<collectingCountry id="52FA76783E7D6A726E7FFE014BC3FE72" box="[584,714,484,510]" name="Guatemala" pageId="12" pageNumber="13">Guatemala</collectingCountry>
,
<collectingCountry id="52FA76783E7D6A726EE1FE014A42FE72" box="[726,843,484,510]" name="Honduras" pageId="12" pageNumber="13">Honduras</collectingCountry>
,
<collectingCountry id="52FA76783E7D6A726F61FE014AD2FE72" box="[854,987,484,510]" name="Costa Rica" pageId="12" pageNumber="13">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="62F765633E7D6A726D0FFDEE4D1BFDAA" box="[312,1042,523,550]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="2A5236E83E7D6A726D0FFDEE4D1BFDAA" blockId="12.[264,1323,284,550]" box="[312,1042,523,550]" pageId="12" pageNumber="13">
<emphasis id="1899EAFA3E7D6A726D0FFDEE48B4FDA9" bold="true" box="[312,445,523,549]" pageId="12" pageNumber="13">Etymology</emphasis>
.
<taxonomicName id="EDED4D6B3E7D6A726DFBFDE94B25FDA9" box="[460,556,524,549]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="40" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
<emphasis id="1899EAFA3E7D6A726DFBFDE94B25FDA9" box="[460,556,524,549]" italics="true" pageId="12" pageNumber="13">Grandis</emphasis>
</taxonomicName>
, L. refers to the large size of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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