treatments-xml/data/A2/44/87/A24487FE3E606A6C6D3FFDB94DB0F9BA.xml

<document id="2497D299A68ED637EC9723FF90CBBF0D" ID-DOI="10.5281/zenodo.158165" ID-GBIF-Dataset="fb07d896-f75a-4f51-aeb0-2700da02c70e" ID-ISSN="1175­5326" ID-Zenodo-Dep="158165" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038221360" checkinUser="plazi" docAuthor="Short, A. E. Z. &amp; Perkins, P. D." docDate="2004" docId="A24487FE3E606A6C6D3FFDB94DB0F9BA" docLanguage="en" docName="zt00783.pdf" docOrigin="Zootaxa 783" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Oocyclus sharpi Short &amp; Perkins, 2004, n. sp." docType="treatment" docVersion="10" lastPageNumber="19" masterDocId="5E7DFF863E716A7E6C37FFE54909FF8C" masterDocTitle="A revision of Oocyclus Sharp of Mexico and Central America (Coleoptera: Hydrophilidae)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="18" updateTime="1698574933192" updateUser="plazi">
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<mods:title id="C5FD2F2C78F951C246FE8AD660E7ED4B">A revision of Oocyclus Sharp of Mexico and Central America (Coleoptera: Hydrophilidae)</mods:title>
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<mods:namePart id="E582F81C5432863370F8B35BFAB3C087">Short, A. E. Z.</mods:namePart>
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<mods:date id="AF8723ED91DD45DDAC2331F32706B81F">2004</mods:date>
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<taxonomicName id="EDED4D6B3E606A6F6D3FFDB948C2FDFA" ID-CoL="49RWR" box="[264,459,604,630]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sharpi" status="sp. nov.">
<emphasis id="1899EAFA3E606A6F6D3FFDB948C2FDFA" bold="true" box="[264,459,604,630]" italics="true" pageId="17" pageNumber="18">Oocyclus sharpi</emphasis>
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(
<figureCitation id="B2D62A6D3E606A6F6D26FD9A487BFD15" box="[273,370,639,665]" captionStart="FIGURE 12" captionStartId="35.[264,368,1849,1873]" captionTargetBox="[327,1290,318,1789]" captionTargetId="figure@35.[264,1323,308,1825]" captionTargetPageId="35" captionText="FIGURE 12. Oocyclus sharpi n sp. holotype. Dorsal and lateral habitus." httpUri="https://zenodo.org/record/169621/files/figure.png" pageId="17" pageNumber="18">Figs. 12</figureCitation>
, 
<figureCitation id="B2D62A6D3E606A6F6D48FD9A4894FD15" box="[383,413,639,665]" captionStart="FIGURES 24 – 27" captionStartId="41.[264,382,1801,1825]" captionTargetBox="[318,1252,289,1788]" captionTargetId="figure@41.[264,1323,288,1797]" captionTargetPageId="41" captionText="FIGURES 24 – 27. Oocyclus aedeagi, dorsal views. Scale bars = 0.2 mm. – 24. O. sharpi (holotype). – 25. O. substillus n. sp. (holotype). – 26. O. tapanti n. sp. (holotype). – 27. O. vestitus Sharp (holotype)." httpUri="https://zenodo.org/record/158183/files/figure.png" pageId="17" pageNumber="18">24</figureCitation>
)
</paragraph>
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<taxonomicName id="EDED4D6B3E606A6F6D3FFD2A4B0EFD69" ID-CoL="49RX3" authority="Sharp" authorityName="Sharp" box="[264,519,718,742]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="1899EAFA3E606A6F6D3FFD2A48CBFD6A" box="[264,450,719,742]" italics="true" pageId="17" pageNumber="18">Oocyclus vestitus</emphasis>
<treatmentCitation id="AB4C10F93E606A6F6DFEFD2B4B0EFD69" author="Sharp" box="[457,519,718,741]" page="764" pageId="17" pageNumber="18" year="1887">
<bibRefCitation id="4E7C4B193E606A6F6DFEFD2B4B0EFD69" author="Sharp" box="[457,519,718,741]" pageId="17" pageNumber="24" refString="Sharp, D. (1887). Insecta, Coleoptera, Vol. 1, Part 2. In: Godman, F. D, Schvin, O.: Biologia Centrali-Americana (16): XV + 824 pp. London. (pp i - xi, 673 - 824 issued in 1887)." type="book chapter" year="1887">Sharp</bibRefCitation>
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–Sharp, 1887: 764 (nec. 
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: 62
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)
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</paragraph>
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<typeStatus id="F556884A3E606A6F6D3FFCFC484CFCBF" box="[264,325,793,819]" pageId="17" pageNumber="18">Type</typeStatus>
Locality.
</emphasis>
Rock face seep at Cerro Azul, on road between Carmona and Bella Vista, Guanacaste Province, 
<collectingCountry id="52FA76783E606A6F6E3AFCA44B98FCD7" box="[525,657,833,859]" name="Costa Rica" pageId="17" pageNumber="18">Costa Rica</collectingCountry>
, 
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, 
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, 
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elevation.
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Material. 
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.
</emphasis>
Male: “ 
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: Guanacaste Prov./ Cerro Azul, road btwn Carmona/ &amp; Bella Vista, 
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elev./ waterfall/seep; 
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/ A. Short, R. Roughley, W. Porras”, “ 
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/ 
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/ 
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/ Short &amp; Perkins” (
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). 
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<typeStatus id="F556884A3E606A6F6D3FFC04488CFC77" box="[264,389,993,1019]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
(213). 
<collectingCountry id="52FA76783E606A6F6DE8FC044B92FC77" box="[479,667,993,1019]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
: Alajuela Prov.,
</emphasis>
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N. Los Lagos on Los Lagos­ Colonia Rd., 
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elev., seeps along Rio San Lorenzo, A. E. Z. Short &amp; R. Roughley leg., CR­03­04 (7 
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, 13 
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); 
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N. Los Lagos, small stream nr. Rio San Lorenzo, 
<date id="5E5310283E606A6F6D3FFBBC4881FBFF" box="[264,392,1113,1139]" pageId="17" pageNumber="18" value="2003-06-12">12.vi.2003</date>
, A. E. Z. Short leg. (1 
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); 
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S. Vara Blanca on rt. 126, 
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, rock wall seeps, A. E. Z. Short leg. (10 
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, 25 
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); 
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S. Cariblanco, 
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elev., 
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, wall seep, A. E. Z. Short &amp; D. J. Lebbin leg., AS­04­055 (3 
<collectionCode id="4CFCAE2D3E606A6F68A3FB4C4DE3FB4F" box="[1172,1258,1193,1219]" pageId="17" pageNumber="18">INBIO</collectionCode>
); San Ramón, Alberto Brenes Biological Station, Rio San Lorencito, 
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elev., (
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)­ (
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).1999, E. Ulate &amp; A. Mora leg., “Tp. Berlesse” (1 
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); 
<emphasis id="1899EAFA3E606A6F6832FB1C4DE1FA9F" bold="true" box="[1029,1256,1273,1299]" pageId="17" pageNumber="18">Guanacaste Prov.,</emphasis>
same data as 
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(30 
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, 60 
<collectionCode id="4CFCAE2D3E606A6F6E67FAC44BAFFAB7" box="[592,678,1313,1339]" pageId="17" pageNumber="18">INBIO</collectionCode>
); 
<quantity id="ED159B0D3E606A6F6E8BFAC44BF3FAB7" box="[700,762,1313,1339]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="km" value="2.0">2 km</quantity>
N of Carmona, 
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elev., 
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, waterfall margin, A. E. Z. Short, R. Roughley, &amp; W. Porras leg, CR­03­06 (16 
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, 48 
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). Representative specimens deposited in 
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, 
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, 
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, 
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, 
<collectionCode id="4CFCAE2D3E606A6F685AFA944DC9FA07" box="[1133,1216,1393,1419]" pageId="17" pageNumber="18">ESMC</collectionCode>
, 
<collectionCode id="4CFCAE2D3E606A6F68F9FA944C2FFA07" LSID="urn:lsid:biocol.org:col:34795" box="[1230,1318,1393,1419]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18">FMNH</collectionCode>
, 
<collectionCode id="4CFCAE2D3E606A6F6D3FFA7C4843FA3F" LSID="urn:lsid:biocol.org:col:33791" box="[264,330,1433,1459]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="17" pageNumber="18">MCZ</collectionCode>
, 
<collectionCode id="4CFCAE2D3E606A6F6D61FA7C48A8FA3F" box="[342,417,1433,1459]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="17" pageNumber="18">NMW</collectionCode>
&amp; 
<collectionCode id="4CFCAE2D3E606A6F6DF0FA7C4B16FA3F" LSID="urn:lsid:biocol.org:col:34871" box="[455,543,1433,1459]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
.
</paragraph>
<paragraph id="2A5236E83E606A6F6D0FFA244D7CF9A7" blockId="17.[264,1324,793,1899]" pageId="17" pageNumber="18">
<emphasis id="1899EAFA3E606A6F6D0FFA244B59FA57" bold="true" box="[312,592,1473,1499]" pageId="17" pageNumber="18">Differential Diagnosis.</emphasis>
The dorsal setae arising from the general punctation separate this species from all others in the region except the Mexican 
<taxonomicName id="EDED4D6B3E606A6F6FE9FA0C4D53F98E" box="[990,1114,1513,1538]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="1899EAFA3E606A6F6FE9FA0C4D53F98E" box="[990,1114,1513,1538]" italics="true" pageId="17" pageNumber="18">O. vestitus</emphasis>
</taxonomicName>
, which possesses dorsal punctation of multiple sizes and pale pronotal anterolateral margins.
</paragraph>
</subSubSection>
<subSubSection id="62F765633E606A6C6D0FF9DC4A82FA6A" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="2A5236E83E606A6C6D0FF9DC4A82FA6A" blockId="17.[264,1324,793,1899]" lastBlockId="18.[264,1323,284,1590]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="1899EAFA3E606A6F6D0FF9DC4BBCF9DF" bold="true" box="[312,693,1593,1619]" pageId="17" pageNumber="18">Description. Size and Form</emphasis>
. Total Length=3.0– 
<quantity id="ED159B0D3E606A6F6FF6F9DC4D1DF9DF" box="[961,1044,1593,1619]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="mm" value="3.5">3.5mm</quantity>
. EL/EW=1.07. Oval, slightly convex. Elytra slightly longer than wide. 
<emphasis id="1899EAFA3E606A6F6F4EF9844AB6F9F7" bold="true" box="[889,959,1633,1659]" pageId="17" pageNumber="18">Color</emphasis>
. Dorsum black; each elytron with 3–4 iridescent bronze patches. Maxillary and labial palpi yellow. Mentum and stipes yellow to, lighter than venter of head. Pronotum entirely black, without pale spots anterolaterally. Sternum, ventrites, epipleura and pseudoepipleura yellow to light brown. Legs yellow, with femora slightly darkened basally and along anterior edge. 
<emphasis id="1899EAFA3E606A6F6858F8E44DB8F897" bold="true" box="[1135,1201,1793,1819]" pageId="17" pageNumber="18">Head</emphasis>
. Labrum and clypeus with general fine punctation, distance between punctures 1.5–2.5x width of one puncture. Maxillary palpi ¾ width of labrum; segments 2 and 3 subequal with apical segment slightly longer. Labial palpi almost equal to length of mentum; apical two segments subequal in length. Scape subequal in length as segments 2–5, segment 2 subequal or longer in length than segments 3–5, segments 3 and 4 subequal in length, apical segment at least 1.5x longer then penultimate segment. Labrum with 2–3 setiferous systematic punctures centrally; anterior margin straight, not sinuate. Clypeus with 4–6 setiferous systematic punctures along anterior margin. Frons with irregular row of setiferous systematic punctures mesad of each eye; top of head with impressed setiferous punctures 2– 
<date id="5E5310283E636A6C6938FDE94C15FDAA" box="[1295,1308,524,550]" pageId="18" pageNumber="19">3</date>
x larger than and mixed with general punctation. Mentum quadrate, anterior margin slightly convex and with small indentation anteromedially; with several setiferous punctures on anterior half. 
<emphasis id="1899EAFA3E636A6C6D92FD614B0BFD12" bold="true" box="[421,514,644,670]" pageId="18" pageNumber="19">Thorax</emphasis>
. Pronotal and elytral punctation of pronotum composed of very fine, nearly obsolete punctation and distinctly larger punctures bearing erect or recumbent long setae; larger setiferous punctures sparser on pronotal disc. Lateral margins of pronotum and elytra with row of punctures bearing long yellow seta, appearing as a yellow fringe. Systematic punctures of pronotum obscured by other setiferous punctures. Posterolateral corners of pronotum rounded. Band of irregular punctures bearing erect setae present along elytral suture; these punctures slightly smaller than other setiferous elytral punctures. Sutural punctation of elytra composed of irregular row of fine punctures bearing usually erect setae. First row of punctures on elytra composed of clusters of setiferous punctures anteriorly, bearing erect setae, forming a single row of setiferous punctures in posterior half. Second row of punctures composed of series of two aggregated setiferous punctures anteriorly. Prosternum carinate medially, without spines or long setae anteromedially. Mesosternal process 1.5x longer than wide, bearing 3–5 long setae at apex of process. Metasternum with oval glabrous area posteromedially, about as long as wide; less than half the total length of metasternum. Procoxae densely pubescent, without spines. Protibiae with 9–11 spines on dorsal face. Protarsal segments 1–4 small, subequal in length; segment 5 as long as segments 1–4. 
<collectingCountry id="52FA76783E636A6C6F20FAE14A5BFA92" box="[791,850,1284,1310]" name="India" pageId="18" pageNumber="19">Hind</collectingCountry>
femora glabrous. 
<emphasis id="1899EAFA3E636A6C6806FAE14DA2FA92" bold="true" box="[1073,1195,1284,1310]" pageId="18" pageNumber="19">Abdomen</emphasis>
. Ventrites sparsely pubescent; setae usually suberect; longer than longest setae around margin of metasternal glabrous area. Fifth exposed ventrite entire. Aedeagus (
<figureCitation id="B2D62A6D3E636A6C6801FAB14D84FAE2" box="[1078,1165,1364,1390]" captionStart="FIGURES 24 – 27" captionStartId="41.[264,382,1801,1825]" captionTargetBox="[318,1252,289,1788]" captionTargetId="figure@41.[264,1323,288,1797]" captionTargetPageId="41" captionText="FIGURES 24 – 27. Oocyclus aedeagi, dorsal views. Scale bars = 0.2 mm. – 24. O. sharpi (holotype). – 25. O. substillus n. sp. (holotype). – 26. O. tapanti n. sp. (holotype). – 27. O. vestitus Sharp (holotype)." httpUri="https://zenodo.org/record/158183/files/figure.png" pageId="18" pageNumber="19">Fig. 24</figureCitation>
) with ventral sides of parameres narrowly but distinctly wider than dorsal sides; tips curved inward and slightly pointed on inner margin. Tip of median lobe bifid, rising to nearly the apex of the parameres. Basal piece narrowed and rounded basally.
</paragraph>
</subSubSection>
<subSubSection id="62F765633E636A6C6D0FFA114BDBF982" box="[312,722,1524,1550]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="2A5236E83E636A6C6D0FFA114BDBF982" blockId="18.[264,1323,284,1590]" box="[312,722,1524,1550]" pageId="18" pageNumber="19">
<emphasis id="1899EAFA3E636A6C6D0FFA1148D8F982" bold="true" box="[312,465,1524,1550]" pageId="18" pageNumber="19">Distribution</emphasis>
. 
<collectingCountry id="52FA76783E636A6C6DE8FA114B6BF982" box="[479,610,1524,1550]" name="Costa Rica" pageId="18" pageNumber="19">Costa Rica</collectingCountry>
, 
<collectingCountry id="52FA76783E636A6C6E59FA114BC6F982" box="[622,719,1524,1550]" name="Panama" pageId="18" pageNumber="19">Panamá</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="62F765633E636A6C6D0FF9F94DB0F9BA" box="[312,1209,1564,1590]" pageId="18" pageNumber="19" type="etymology">
<paragraph id="2A5236E83E636A6C6D0FF9F94DB0F9BA" blockId="18.[264,1323,284,1590]" box="[312,1209,1564,1590]" pageId="18" pageNumber="19">
<emphasis id="1899EAFA3E636A6C6D0FF9F948B4F9BA" bold="true" box="[312,445,1564,1590]" pageId="18" pageNumber="19">Etymology</emphasis>
. Named in honor of David Sharp, who first described the genus.
</paragraph>
</subSubSection>
</treatment>
</document>