452 lines
52 KiB
XML
452 lines
52 KiB
XML
<document id="A1A90BF1046588396AF00FA1D72211D1" ID-CLB-Dataset="53364" ID-DOI="10.11646/zootaxa.5293.1.9" ID-GBIF-Dataset="d0c9563a-e785-4f66-9acf-19184bc9babd" ID-ISSN="1175-5326" ID-Zenodo-Dep="7959901" ID-ZooBank="1954C1DE-4A18-4760-99C7-F4E1AA87F704" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1684796984625" checkinUser="plazi" docAuthor="Duran, Daniel P., Chambers, Aaron W., Nelson, Kristie N. & Roman, Stephen J." docDate="2023" docId="A23F87AB5C2CFFC4FF61CE85FA59FCA0" docLanguage="en" docName="zootaxa.5293.1.9.pdf" docOrigin="Zootaxa 5293 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5293.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cicindela timbisha Duran, Chambers, Nelson & Roman 2023, n. sp." docType="treatment" docVersion="4" lastPageNumber="183" masterDocId="5E06FFD35C2DFFC0FFF6CC0BFFDFFFB5" masterDocTitle="A new species of tiger beetle (Coleoptera: Cicindelidae) from the Death Valley ecosystem" masterLastPageNumber="184" masterPageNumber="179" pageNumber="180" updateTime="1685034626501" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="DCCA209AD7DD069DDB263949BD94E853" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="ADE72504A0719124837837FA8E7B0CE3">
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<mods:title id="A7EC51B98A13CFDC2012C5D8958F99D3">A new species of tiger beetle (Coleoptera: Cicindelidae) from the Death Valley ecosystem</mods:title>
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</mods:titleInfo>
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<mods:name id="AC06103CF67ABCA655CD68ECE1A56344" type="personal">
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<mods:role id="A092670C4A72B9B9008E3775932A3FC1">
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<mods:roleTerm id="648949BA8487077384CB528DDFA676D5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="320CB4AB622E4848A66AA5D37807D082">Duran, Daniel P.</mods:namePart>
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<mods:affiliation id="9BDA1C4A93DCFBFAB482DF152ECC5EF3">Department of Environmental Science, Rowan University, Glassboro, NJ 08028, USA.</mods:affiliation>
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</mods:name>
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<mods:name id="32DA1FD363B21D818E9E3D22924D41D9" type="personal">
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<mods:role id="AA0D9FEA3DA75934259EC93BAEB65145">
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<mods:roleTerm id="007C51B1BB62A3289F53CAD55272D3ED">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D203A714311F4FDA315EC99BFF1E2337">Chambers, Aaron W.</mods:namePart>
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<mods:affiliation id="71176D6D3C59338E1D0A3DE7283D6571">Deserticolus Bioconsulting, Green Valley, AZ 86622 USA.</mods:affiliation>
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</mods:name>
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<mods:name id="D4D1D3CD8017B6FEF31C241C804D5511" type="personal">
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<mods:roleTerm id="7E874D78F703D907AC719E5CA24C57B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="99AE342DA0024595C97D69EDD8C01B33">Nelson, Kristie N.</mods:namePart>
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<mods:affiliation id="2943D06DC57ABA13B0E9281D97BFFE05">Missouri Department of Natural Resources, Division of State Parks, Jefferson City, MO 65101, USA.</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="A852488B72101909BF94AF5FC7AB595E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A77CA638A500AA48266454B067D70CDA">Roman, Stephen J.</mods:namePart>
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<mods:affiliation id="6DF2173181357A488494435F71161633">178 Winecup Way, Austin, TX 78737, USA.</mods:affiliation>
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<mods:typeOfResource id="8DF103847B80D3220F9EC5EE9695F87A">text</mods:typeOfResource>
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<mods:title id="98C90514E991564E486E666B50760319">Zootaxa</mods:title>
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<mods:part id="3179EE12286AAA96BE22D80590817914">
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<mods:date id="361BE9AE6E488999CFADE4E8A492E96A">2023</mods:date>
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<mods:number id="31DFFB4AB8FF67F892E77C5A6D7C6356">2023-05-19</mods:number>
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<mods:number id="84475F04B8578B0E9466C263865C2B8A">5293</mods:number>
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<mods:url id="BA803C2E540DB830CAC35A0932103491">http://dx.doi.org/10.11646/zootaxa.5293.1.9</mods:url>
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<mods:classification id="867DAFE2508010D8ABDBD47D206EEB97">journal article</mods:classification>
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<mods:identifier id="0313887C4BFD068B3C1AA3070D2B0003" type="DOI">10.11646/zootaxa.5293.1.9</mods:identifier>
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<treatment id="A23F87AB5C2CFFC4FF61CE85FA59FCA0" ID-DOI="http://doi.org/10.5281/zenodo.7971626" ID-Zenodo-Dep="7971626" LSID="urn:lsid:plazi:treatment:A23F87AB5C2CFFC4FF61CE85FA59FCA0" httpUri="http://treatment.plazi.org/id/A23F87AB5C2CFFC4FF61CE85FA59FCA0" lastPageId="4" lastPageNumber="183" pageId="1" pageNumber="180">
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<subSubSection id="628C65365C2CFFC1FF61CE85FC44FD1C" box="[151,923,654,681]" pageId="1" pageNumber="180" type="nomenclature">
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<paragraph id="2A2936BD5C2CFFC1FF61CE85FC44FD1C" blockId="1.[151,923,654,717]" box="[151,923,654,681]" pageId="1" pageNumber="180">
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<heading id="716181D15C2CFFC1FF61CE85FC44FD1C" bold="true" box="[151,923,654,681]" fontSize="11" level="1" pageId="1" pageNumber="180" reason="1">
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<emphasis id="18E2EAAF5C2CFFC1FF61CE85FC44FD1C" bold="true" box="[151,923,654,681]" pageId="1" pageNumber="180">
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<taxonomicName id="ED964D3E5C2CFFC1FF61CE85FC93FD1C" authority="Duran, Chambers, Nelson & Roman" authorityName="Duran, Chambers, Nelson & Roman" authorityYear="2023" box="[151,844,654,681]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
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||
<emphasis id="18E2EAAF5C2CFFC1FF61CE85FEA5FD1C" bold="true" box="[151,378,654,681]" italics="true" pageId="1" pageNumber="180">Cicindela timbisha</emphasis>
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||
Duran, Chambers, Nelson & Roman
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||
</taxonomicName>
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||
,
|
||
<taxonomicNameLabel id="03D157D45C2CFFC1FCAFCE84FC44FD1C" box="[857,923,655,681]" pageId="1" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="628C65365C2CFFC1FF61CEB8FE86FD78" box="[151,345,691,717]" pageId="1" pageNumber="180" type="description">
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<paragraph id="2A2936BD5C2CFFC1FF61CEB8FE86FD78" blockId="1.[151,923,654,717]" box="[151,345,691,717]" pageId="1" pageNumber="180">
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<figureCitation id="B2AD2A385C2CFFC1FF61CEB8FEC3FD78" box="[151,284,691,717]" captionStart="FIGURE 1" captionStartId="2.[152,255,1487,1511]" captionTargetBox="[168,1424,195,1452]" captionTargetId="figure-20@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of C. timbisha, n. sp. (top row) males (A–B), females (C–D) (USA: California: Inyo Co.); C. senilis (bottom row), males (E–F), females (G–H) (USA: California: Solano Co.)" figureDoi="http://doi.org/10.5281/zenodo.7959903" httpUri="https://zenodo.org/record/7959903/files/figure.png" pageId="1" pageNumber="180">Figs. 1A–D</figureCitation>
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,
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||
<figureCitation id="B2AD2A385C2CFFC1FEDECEB8FE8AFD78" box="[296,341,691,717]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[152,255,751,775]" captionStartId-1="3.[152,255,2017,2041]" captionStartId-2="4.[152,255,1572,1596]" captionTargetBox-0="[218,1371,181,719]" captionTargetBox-2="[257,1330,813,1547]" captionTargetId-0="figure-18@3.[217,1371,181,727]" captionTargetId-1="figure-40@3.[264,1323,799,1993]" captionTargetId-2="figure-310@4.[257,1330,813,1547]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Frontal habitus of C. timbisha, n. sp., A) male, B) female." captionText-1="FIGURE 3. Aedeagus of C. timbisha, n. sp. (left) and C. senilis (right)." captionText-2="FIGURE 4. Cicindela timbisha n. sp. in situ in dark muddy soil near a freshwater spring." figureDoi-0="http://doi.org/10.5281/zenodo.7959905" figureDoi-1="http://doi.org/10.5281/zenodo.7971624" figureDoi-2="http://doi.org/10.5281/zenodo.7959907" httpUri-0="https://zenodo.org/record/7959905/files/figure.png" httpUri-1="https://zenodo.org/record/7971624/files/figure.png" httpUri-2="https://zenodo.org/record/7959907/files/figure.png" pageId="1" pageNumber="180">2–4</figureCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="628C65365C2CFFC1FF61CEF0FAEBFC10" pageId="1" pageNumber="180" type="materials_examined">
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<paragraph id="2A2936BD5C2CFFC1FF61CEF0FAEBFC10" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="180">
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<emphasis id="18E2EAAF5C2CFFC1FF61CEF0FE9BFCA1" bold="true" box="[151,324,762,789]" pageId="1" pageNumber="180">Type material.</emphasis>
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||
<materialsCitation id="9AFE3CE05C2CFFC1FEBFCEF0FBE4FCA0" box="[329,1083,763,791]" collectionCode="USNM" county="Inyo Co." location="Inyo Co." pageId="1" pageNumber="180" specimenCount="1" stateProvince="California" typeStatus="holotype">
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||
<typeStatus id="F52D881F5C2CFFC1FEBFCEF0FE3EFCA0" box="[329,481,763,789]" pageId="1" pageNumber="180">HOLOTYPE</typeStatus>
|
||
: 1 ♁,
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||
<collectingRegion id="E852F85F5C2CFFC1FDD6CEF0FD49FCA0" box="[544,662,763,789]" country="United States of America" name="California" pageId="1" pageNumber="180">California</collectingRegion>
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||
:
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||
<collectingCounty id="C3484E315C2CFFC1FD56CEF0FCDEFCA0" box="[672,769,763,789]" pageId="1" pageNumber="180">Inyo Co.</collectingCounty>
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||
// 20.April.2022 // (
|
||
<collectionCode id="4C87AE785C2CFFC1FC2ACEF0FBECFCA0" box="[988,1075,763,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="180" type="Museum">USNM</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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<materialsCitation id="9AFE3CE05C2CFFC1FBB3CEF0FDE2FC8C" collectionCode="USNM" collectorName="April." county="Inyo Co." location="Inyo Co." pageId="1" pageNumber="180" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
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||
<typeStatus id="F52D881F5C2CFFC1FBB3CEF0FB3BFCA0" box="[1093,1252,763,789]" pageId="1" pageNumber="180">PARATYPES</typeStatus>
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||
:
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||
<specimenCount id="3C90FD345C2CFFC1FB1BCEF0FAC9FCA0" box="[1261,1302,763,789]" pageId="1" pageNumber="180" type="female">1 ♀</specimenCount>
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||
,
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<collectingRegion id="E852F85F5C2CFFC1FAD5CEF0FA46FCA0" box="[1315,1433,763,789]" country="United States of America" name="California" pageId="1" pageNumber="180">California</collectingRegion>
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||
:
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||
<collectingCounty id="C3484E315C2CFFC1FF61CF14FF25FC8C" box="[151,250,799,825]" pageId="1" pageNumber="180">Inyo Co.</collectingCounty>
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||
// 20.
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||
<collectorName id="8763536B5C2CFFC1FECFCF14FEA3FC8C" box="[313,380,799,825]" pageId="1" pageNumber="180">April.</collectorName>
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||
2022 // (
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<collectionCode id="4C87AE785C2CFFC1FE28CF14FDEAFC8C" box="[478,565,799,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="180" type="Museum">USNM</collectionCode>
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||
)
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||
</materialsCitation>
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||
; 3 ♁,
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<materialsCitation id="9AFE3CE05C2CFFC1FD75CF14FB02FC8C" box="[643,1245,799,825]" collectionCode="DPDC" collectorName="April." county="Inyo Co." location="Inyo Co." pageId="1" pageNumber="180" specimenCount="3" specimenCount-female="3" stateProvince="California" typeStatus="paratype">
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||
<specimenCount id="3C90FD345C2CFFC1FD75CF14FD71FC8C" box="[643,686,799,825]" pageId="1" pageNumber="180" type="female">3 ♀</specimenCount>
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||
,
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||
<collectingRegion id="E852F85F5C2CFFC1FD4ACF14FCEDFC8C" box="[700,818,799,825]" country="United States of America" name="California" pageId="1" pageNumber="180">California</collectingRegion>
|
||
:
|
||
<collectingCounty id="C3484E315C2CFFC1FCC8CF14FC7EFC8C" box="[830,929,799,825]" pageId="1" pageNumber="180">Inyo Co.</collectingCounty>
|
||
// 20.
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||
<collectorName id="8763536B5C2CFFC1FC16CF14FBFCFC8C" box="[992,1059,799,825]" pageId="1" pageNumber="180">April.</collectorName>
|
||
2022 // (
|
||
<collectionCode id="4C87AE785C2CFFC1FB72CF14FB0AFC8C" box="[1156,1237,799,825]" pageId="1" pageNumber="180">DPDC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AFE3CE05C2CFFC1FB1FCF14FDE2FCE8" collectionCode="AWCC" collectorName="April." county="Inyo Co." location="Inyo Co." pageId="1" pageNumber="180" specimenCount="1" stateProvince="California" typeStatus="paratype">
|
||
1 ♁,
|
||
<collectingRegion id="E852F85F5C2CFFC1FAD5CF14FA46FC8C" box="[1315,1433,799,825]" country="United States of America" name="California" pageId="1" pageNumber="180">California</collectingRegion>
|
||
:
|
||
<collectingCounty id="C3484E315C2CFFC1FF61CF48FF25FCE8" box="[151,250,835,861]" pageId="1" pageNumber="180">Inyo Co.</collectingCounty>
|
||
// 20.
|
||
<collectorName id="8763536B5C2CFFC1FECECF48FEA4FCE8" box="[312,379,835,861]" pageId="1" pageNumber="180">April.</collectorName>
|
||
2022 // (
|
||
<collectionCode id="4C87AE785C2CFFC1FE2ACF48FDEAFCE8" box="[476,565,835,861]" pageId="1" pageNumber="180">AWCC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="9AFE3CE05C2CFFC1FDBECF48FB7AFCE8" box="[584,1189,835,861]" collectionCode="KNNC" collectorName="April." county="Inyo Co." location="Inyo Co." pageId="1" pageNumber="180" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
|
||
<specimenCount id="3C90FD345C2CFFC1FDBECF48FDACFCE8" box="[584,627,835,861]" pageId="1" pageNumber="180" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="E852F85F5C2CFFC1FD77CF48FD28FCE8" box="[641,759,835,861]" country="United States of America" name="California" pageId="1" pageNumber="180">California</collectingRegion>
|
||
:
|
||
<collectingCounty id="C3484E315C2CFFC1FCF4CF48FCBAFCE8" box="[770,869,835,861]" pageId="1" pageNumber="180">Inyo Co.</collectingCounty>
|
||
// 20.
|
||
<collectorName id="8763536B5C2CFFC1FC55CF48FC39FCE8" box="[931,998,835,861]" pageId="1" pageNumber="180">April.</collectorName>
|
||
2022 // (
|
||
<collectionCode id="4C87AE785C2CFFC1FBB1CF48FB42FCE8" box="[1095,1181,835,861]" pageId="1" pageNumber="180">KNNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
; 1 ♁,
|
||
<materialsCitation id="9AFE3CE05C2CFFC1FB1CCF48FDFEFC34" collectionCode="SJRC" collectorName="April." county="Inyo Co." location="Inyo Co." pageId="1" pageNumber="180" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
|
||
<specimenCount id="3C90FD345C2CFFC1FB1CCF48FACBFCE8" box="[1258,1300,835,861]" pageId="1" pageNumber="180" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="E852F85F5C2CFFC1FAD5CF48FA46FCE8" box="[1315,1433,835,861]" country="United States of America" name="California" pageId="1" pageNumber="180">California</collectingRegion>
|
||
:
|
||
<collectingCounty id="C3484E315C2CFFC1FF61CF6CFF27FC34" box="[151,248,871,897]" pageId="1" pageNumber="180">Inyo Co.</collectingCounty>
|
||
// 20.
|
||
<collectorName id="8763536B5C2CFFC1FEC5CF6CFEA9FC34" box="[307,374,871,897]" pageId="1" pageNumber="180">April.</collectorName>
|
||
2022 // (
|
||
<collectionCode id="4C87AE785C2CFFC1FE22CF6CFDC5FC34" box="[468,538,871,897]" pageId="1" pageNumber="180">SJRC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
; 2 ♁,
|
||
<materialsCitation id="9AFE3CE05C2CFFC1FD97CF6CFB72FC34" box="[609,1197,871,897]" collectionCode="DEKC" collectorName="April." county="Inyo Co." location="Inyo Co." pageId="1" pageNumber="180" specimenCount="2" specimenCount-female="2" stateProvince="California" typeStatus="paratype">
|
||
<specimenCount id="3C90FD345C2CFFC1FD97CF6CFD55FC34" box="[609,650,871,897]" pageId="1" pageNumber="180" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="E852F85F5C2CFFC1FD61CF6CFCD2FC34" box="[663,781,871,897]" country="United States of America" name="California" pageId="1" pageNumber="180">California</collectingRegion>
|
||
:
|
||
<collectingCounty id="C3484E315C2CFFC1FCE0CF6CFCA7FC34" box="[790,888,871,897]" pageId="1" pageNumber="180">Inyo Co.</collectingCounty>
|
||
// 20.
|
||
<collectorName id="8763536B5C2CFFC1FC44CF6CFC2AFC34" box="[946,1013,871,897]" pageId="1" pageNumber="180">April.</collectorName>
|
||
2022 // (
|
||
<collectionCode id="4C87AE785C2CFFC1FBA5CF6CFB7AFC34" box="[1107,1189,871,897]" pageId="1" pageNumber="180">DEKC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
; 1 ♁,
|
||
<materialsCitation id="9AFE3CE05C2CFFC1FB1BCF6CFDF6FC10" collectionCode="JASC" collectorName="April." county="Inyo Co." location="Inyo Co." pageId="1" pageNumber="180" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="paratype">
|
||
<specimenCount id="3C90FD345C2CFFC1FB1BCF6CFAC9FC34" box="[1261,1302,871,897]" pageId="1" pageNumber="180" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="E852F85F5C2CFFC1FAD5CF6CFA46FC34" box="[1315,1433,871,897]" country="United States of America" name="California" pageId="1" pageNumber="180">California</collectingRegion>
|
||
:
|
||
<collectingCounty id="C3484E315C2CFFC1FF61CF80FF25FC10" box="[151,250,907,933]" pageId="1" pageNumber="180">Inyo Co.</collectingCounty>
|
||
// 20.
|
||
<collectorName id="8763536B5C2CFFC1FEC1CF80FEA5FC10" box="[311,378,907,933]" pageId="1" pageNumber="180">April.</collectorName>
|
||
2022 // (
|
||
<collectionCode id="4C87AE785C2CFFC1FE2FCF80FDFFFC10" box="[473,544,907,933]" pageId="1" pageNumber="180">JASC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="9AFE3CE05C2CFFC1FDC5CF80FC95FC10" box="[563,842,907,933]" location="Type" pageId="1" pageNumber="180" specimenCount="1" stateProvince="California" typeStatus="paratype">
|
||
<location id="2F4960665C2CFFC1FDC5CF80FDB4FC10" LSID="urn:lsid:plazi:treatment:A23F87AB5C2CFFC4FF61CE85FA59FCA0:2F4960665C2CFFC1FDC5CF80FDB4FC10" box="[563,619,907,933]" name="Type" pageId="1" pageNumber="180" stateProvince="California">Type</location>
|
||
specimens labelled
|
||
</materialsCitation>
|
||
:
|
||
<typeStatus id="F52D881F5C2CFFC1FCA3CF80FC36FC10" box="[853,1001,907,933]" pageId="1" pageNumber="180">HOLOTYPE</typeStatus>
|
||
or
|
||
<typeStatus id="F52D881F5C2CFFC1FBF8CF80FB43FC10" box="[1038,1180,907,933]" pageId="1" pageNumber="180">PARATYPE</typeStatus>
|
||
, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="628C65365C2CFFC1FF31CFA5FEB1FACC" pageId="1" pageNumber="180" type="diagnosis">
|
||
<paragraph id="2A2936BD5C2CFFC1FF31CFA5FDA4FB14" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="180">
|
||
<emphasis id="18E2EAAF5C2CFFC1FF31CFA5FE9DFC7D" bold="true" box="[199,322,942,968]" pageId="1" pageNumber="180">Diagnosis.</emphasis>
|
||
<taxonomicName id="ED964D3E5C2CFFC1FEBECFA4FDC3FC7C" authority="Duran & Chambers & Nelson & Roman, 2023" authorityName="Duran & Chambers & Nelson & Roman" authorityYear="2023" box="[328,540,943,969]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
|
||
<emphasis id="18E2EAAF5C2CFFC1FEBECFA4FDC3FC7C" box="[328,540,943,969]" italics="true" pageId="1" pageNumber="180">Cicindela timbisha</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="18E2EAAF5C2CFFC1FDD5CFA4FDBDFC7C" bold="true" box="[547,610,943,969]" pageId="1" pageNumber="180">
|
||
<taxonomicNameLabel id="03D157D45C2CFFC1FDD5CFA4FDBDFC7C" box="[547,610,943,969]" pageId="1" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar to
|
||
<taxonomicName id="ED964D3E5C2CFFC1FCC7CFA4FC45FC7C" authorityName="G. Horn" authorityYear="1866" box="[817,922,943,969]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2CFFC1FCC7CFA4FC45FC7C" box="[817,922,943,969]" italics="true" pageId="1" pageNumber="180">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
and could only be confused with that species. In the dichotomous key by
|
||
<bibRefCitation id="4E074B4C5C2CFFC1FE33CFD8FD77FC58" author="Pearson, D. L. & Knisley, C. B. & Duran, D. P. & Kazilek, C. J." box="[453,680,978,1005]" pageId="1" pageNumber="180" refId="ref3073" refString="Pearson, D. L., Knisley, C. B., Duran, D. P. & Kazilek, C. J. (2015) A field guide to the tiger beetles of the United States and Canada: Identification, natural history, and distribution of the Cicindelids. 2 nd Edition. Oxford University Press, New York, New York, 251 pp." type="book" year="2015">
|
||
Pearson
|
||
<emphasis id="18E2EAAF5C2CFFC1FDD2CFD8FD8AFC58" box="[548,597,979,1005]" italics="true" pageId="1" pageNumber="180">et al</emphasis>
|
||
(2015)
|
||
</bibRefCitation>
|
||
, this new species would key to
|
||
<taxonomicName id="ED964D3E5C2CFFC1FBF2CFD8FBB3FC58" authorityName="G. Horn" authorityYear="1866" box="[1028,1132,979,1005]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2CFFC1FBF2CFD8FBB3FC58" box="[1028,1132,979,1005]" italics="true" pageId="1" pageNumber="180">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
, but is distinguished by the following combination of characters. This species lacks a sub-humeral spot (
|
||
<figureCitation id="B2AD2A385C2CFFC1FC04CFFCFB59FBA4" box="[1010,1158,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[152,255,1487,1511]" captionTargetBox="[168,1424,195,1452]" captionTargetId="figure-20@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of C. timbisha, n. sp. (top row) males (A–B), females (C–D) (USA: California: Inyo Co.); C. senilis (bottom row), males (E–F), females (G–H) (USA: California: Solano Co.)" figureDoi="http://doi.org/10.5281/zenodo.7959903" httpUri="https://zenodo.org/record/7959903/files/figure.png" pageId="1" pageNumber="180">Fig 1A–B, D</figureCitation>
|
||
) or in a small number of individuals may possess an atrophied remnant of a sub-humeral spot (
|
||
<figureCitation id="B2AD2A385C2CFFC1FC58C810FC20FB80" box="[942,1023,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[152,255,1487,1511]" captionTargetBox="[168,1424,195,1452]" captionTargetId="figure-20@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of C. timbisha, n. sp. (top row) males (A–B), females (C–D) (USA: California: Inyo Co.); C. senilis (bottom row), males (E–F), females (G–H) (USA: California: Solano Co.)" figureDoi="http://doi.org/10.5281/zenodo.7959903" httpUri="https://zenodo.org/record/7959903/files/figure.png" pageId="1" pageNumber="180">Fig 1C</figureCitation>
|
||
), unlike
|
||
<taxonomicName id="ED964D3E5C2CFFC1FB92C810FB11FB80" authorityName="G. Horn" authorityYear="1866" box="[1124,1230,1051,1077]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2CFFC1FB92C810FB11FB80" box="[1124,1230,1051,1077]" italics="true" pageId="1" pageNumber="180">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
, which has a bold sub-humeral maculation (
|
||
<figureCitation id="B2AD2A385C2CFFC1FE40C834FDF6FBEC" box="[438,553,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[152,255,1487,1511]" captionTargetBox="[168,1424,195,1452]" captionTargetId="figure-20@2.[151,1436,181,1463]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of C. timbisha, n. sp. (top row) males (A–B), females (C–D) (USA: California: Inyo Co.); C. senilis (bottom row), males (E–F), females (G–H) (USA: California: Solano Co.)" figureDoi="http://doi.org/10.5281/zenodo.7959903" httpUri="https://zenodo.org/record/7959903/files/figure.png" pageId="1" pageNumber="180">Fig 1E–H</figureCitation>
|
||
), connected laterally to the humeral spot. The antennal scape of
|
||
<taxonomicName id="ED964D3E5C2CFFC1FB0BC834FAA1FBEC" authority="Duran & Chambers & Nelson & Roman, 2023" authorityName="Duran & Chambers & Nelson & Roman" authorityYear="2023" box="[1277,1406,1087,1113]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
|
||
<emphasis id="18E2EAAF5C2CFFC1FB0BC834FACFFBEC" box="[1277,1296,1087,1113]" italics="true" pageId="1" pageNumber="180">C</emphasis>
|
||
.
|
||
<emphasis id="18E2EAAF5C2CFFC1FAE8C834FAA1FBEC" box="[1310,1406,1087,1113]" italics="true" pageId="1" pageNumber="180">timbisha</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="18E2EAAF5C2CFFC1FA70C834FF67FBC8" bold="true" pageId="1" pageNumber="180">
|
||
<taxonomicNameLabel id="03D157D45C2CFFC1FA70C834FF67FBC8" pageId="1" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
possesses 4–8 setae (often partially abraded, but still indicated by visible setigerous punctures), compared to>10 scape setae (typically 12–16) for
|
||
<taxonomicName id="ED964D3E5C2CFFC1FDFAC88CFDABFB14" authorityName="G. Horn" authorityYear="1866" box="[524,628,1159,1185]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2CFFC1FDFAC88CFDABFB14" box="[524,628,1159,1185]" italics="true" pageId="1" pageNumber="180">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2A2936BD5C2CFFC1FF31C8A0FA96FA84" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="180">
|
||
Other characteristics that differ include the following. In most specimens (16 of 18) of
|
||
<taxonomicName id="ED964D3E5C2CFFC1FB6AC8A0FAC2FB70" authority="Duran & Chambers & Nelson & Roman, 2023" authorityName="Duran & Chambers & Nelson & Roman" authorityYear="2023" box="[1180,1309,1195,1221]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
|
||
<emphasis id="18E2EAAF5C2CFFC1FB6AC8A0FAC2FB70" box="[1180,1309,1195,1221]" italics="true" pageId="1" pageNumber="180">C. timbisha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="18E2EAAF5C2CFFC1FADAC8A0FAACFB70" bold="true" box="[1324,1395,1195,1221]" pageId="1" pageNumber="180">
|
||
<taxonomicNameLabel id="03D157D45C2CFFC1FADAC8A0FAB3FB70" box="[1324,1388,1195,1221]" pageId="1" pageNumber="180" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
the middle band maculations do not reach the lateral margin of the elytra, whereas they always do in
|
||
<taxonomicName id="ED964D3E5C2CFFC1FB1CC8C4FA8BFB5C" authorityName="G. Horn" authorityYear="1866" box="[1258,1364,1231,1257]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2CFFC1FB1CC8C4FA8BFB5C" box="[1258,1364,1231,1257]" italics="true" pageId="1" pageNumber="180">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
. Even when the maculations do reach the lateral margin in the new species they connect with the margin less broadly than in
|
||
<taxonomicName id="ED964D3E5C2CFFC1FF42C91CFEC2FA84" authorityName="G. Horn" authorityYear="1866" box="[180,285,1303,1329]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2CFFC1FF42C91CFEC2FA84" box="[180,285,1303,1329]" italics="true" pageId="1" pageNumber="180">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
. The new species also possesses fewer setae on the frons (15–25) compared to
|
||
<taxonomicName id="ED964D3E5C2CFFC1FB62C91CFB22FA84" authorityName="G. Horn" authorityYear="1866" box="[1172,1277,1303,1329]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2CFFC1FB62C91CFB22FA84" box="[1172,1277,1303,1329]" italics="true" pageId="1" pageNumber="180">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
(>30).
|
||
</paragraph>
|
||
<paragraph id="2A2936BD5C2CFFC1FF31C930FEB1FACC" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="180">
|
||
In addition, the new species is statistically smaller in size from
|
||
<taxonomicName id="ED964D3E5C2CFFC1FC60C930FBDFFAE0" authorityName="G. Horn" authorityYear="1866" box="[918,1024,1339,1365]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2CFFC1FC60C930FBDFFAE0" box="[918,1024,1339,1365]" italics="true" pageId="1" pageNumber="180">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
(two-tailed t-test,
|
||
<emphasis id="18E2EAAF5C2CFFC1FB27C930FB3DFAE1" box="[1233,1250,1339,1364]" italics="true" pageId="1" pageNumber="180">P</emphasis>
|
||
<0.001) for both males and females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="628C65365C2CFFC4FF31C989FDF3FDAC" lastPageId="4" lastPageNumber="183" pageId="1" pageNumber="180" type="description">
|
||
<paragraph id="2A2936BD5C2CFFC1FF31C989FE5FF8B0" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="180">
|
||
<emphasis id="18E2EAAF5C2CFFC1FF31C989FE86FA28" bold="true" box="[199,345,1410,1437]" pageId="1" pageNumber="180">Description.</emphasis>
|
||
Small-sized
|
||
<taxonomicName id="ED964D3E5C2CFFC1FE1CC988FD88FA28" authorityName="Linne" authorityYear="1758" box="[490,599,1411,1437]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="180" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="18E2EAAF5C2CFFC1FE1CC988FD88FA28" box="[490,599,1411,1437]" italics="true" pageId="1" pageNumber="180">Cicindela</emphasis>
|
||
</taxonomicName>
|
||
. Body length
|
||
<quantity id="ED6E9B585C2CFFC1FD02C988FC4AFA28" box="[756,917,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.11" metricValueMax="1.19" metricValueMin="1.03" pageId="1" pageNumber="180" unit="mm" value="11.1" valueMax="11.9" valueMin="10.3">10.3–11.9 mm</quantity>
|
||
<specimenCount id="3C90FD345C2CFFC1FC6DC988FC6FFA28" box="[923,944,1411,1437]" pageId="1" pageNumber="180" type="female">♀</specimenCount>
|
||
mean
|
||
<quantity id="ED6E9B585C2CFFC1FC0CC989FB81FA28" box="[1018,1118,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="180" unit="mm" value="11.5">11.5 mm</quantity>
|
||
, ♁ mean 10.7. Head slightly wider than pronotum, width
|
||
<quantity id="ED6E9B585C2CFFC1FE2FC9ACFDBDFA74" box="[473,610,1447,1473]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="1" pageNumber="180" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7–3.1 mm</quantity>
|
||
, mean
|
||
<specimenCount id="3C90FD345C2CFFC1FD44C9ACFD18FA74" box="[690,711,1447,1473]" pageId="1" pageNumber="180" type="female">♀</specimenCount>
|
||
3.0, mean ♁ 2.8, vertex dark bronze with brassy cupreous highlights especially in supraorbital region; frons dark bronze with metallic blue reflections present in sulci along lateral areas; frons possessing numerous setae (15–25); two supraorbital setae present near each eye. Frons slightly convex, clearly delimited from clypeus, gradually blending into vertex. Frons surface with distinct longitudinal striae especially in lateral areas bordering eyes. Genae bright polished green to blue with deep longitudinal striae abruptly ending at border of vertex. Clypeus color ranging from cupreous to green, blending to other metallic colors along margins, irregularly wrinkled to finely vermiculate. Labrum with 4–8 setae, ochre-yellow to pale yellow with thin dark brown to black border; labrum unidentate, length
|
||
<quantity id="ED6E9B585C2CFFC1FCF2CAA9FC51F908" box="[772,910,1698,1725]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="180" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.5–0.8 mm</quantity>
|
||
, width 1.0–
|
||
<quantity id="ED6E9B585C2CFFC1FBE3CAA8FBB2F908" box="[1045,1133,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="180" unit="mm" value="1.3">1.3 mm</quantity>
|
||
. Mandibles medium-sized, ochraceous, dark testaceous along edges. Maxillary palpi dark shiny metallic green to violet. Labial palpi dark metallic green to violet.
|
||
</paragraph>
|
||
<paragraph id="2A2936BD5C2CFFC1FF31CB04FA86F80C" blockId="1.[151,1437,762,1977]" pageId="1" pageNumber="180">Antennae of normal length, reaching humerus to basal third of elytra, scape dark testaceous with metallic reflections of cupreous, gold, and violet, with 4–8 setae; pedicel dark testaceous with metallic reflections similar to scape, lacking any setae; flagellum antennomeres 3‒4 dark testaceous with metallic cupreous and violet reflections, with ring of apical setae and additional sparse setae throughout, antennomeres 5‒11 ochre-brown, dull-textured without metallic reflections and possessing erect setae in apical rings only, covered with fine pubescence throughout.</paragraph>
|
||
<caption id="7EE966355C2FFFC2FF6EC9C4FC58F9BE" ID-DOI="http://doi.org/10.5281/zenodo.7959903" ID-Zenodo-Dep="7959903" httpUri="https://zenodo.org/record/7959903/files/figure.png" pageId="2" pageNumber="181" startId="2.[152,255,1487,1511]" targetBox="[168,1424,195,1452]" targetPageId="2" targetType="figure">
|
||
<paragraph id="2A2936BD5C2FFFC2FF6EC9C4FC58F9BE" blockId="2.[151,1436,1487,1547]" pageId="2" pageNumber="181">
|
||
<emphasis id="18E2EAAF5C2FFFC2FF6EC9C4FEC7FA52" bold="true" box="[152,280,1487,1511]" pageId="2" pageNumber="181">FIGURE 1.</emphasis>
|
||
Dorsal habitus of
|
||
<taxonomicName id="ED964D3E5C2FFFC2FE39C9C4FD9CFA52" authority="Duran & Chambers & Nelson & Roman, 2023" authorityName="Duran & Chambers & Nelson & Roman" authorityYear="2023" box="[463,579,1487,1511]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
|
||
<emphasis id="18E2EAAF5C2FFFC2FE39C9C4FD9CFA52" box="[463,579,1487,1511]" italics="true" pageId="2" pageNumber="181">C. timbisha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="18E2EAAF5C2FFFC2FDB9C9C4FD58FA52" bold="true" box="[591,647,1487,1511]" pageId="2" pageNumber="181">
|
||
<taxonomicNameLabel id="03D157D45C2FFFC2FDB9C9C4FD58FA52" box="[591,647,1487,1511]" pageId="2" pageNumber="181" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(top row) males (A–B), females (C–D) (USA: California: Inyo Co.);
|
||
<taxonomicName id="ED964D3E5C2FFFC2FAC9C9C4FA43FA52" authorityName="G. Horn" authorityYear="1866" box="[1343,1436,1487,1511]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="181" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2FFFC2FAC9C9C4FA43FA52" box="[1343,1436,1487,1511]" italics="true" pageId="2" pageNumber="181">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
(bottom row), males (E–F), females (G–H) (USA: California: Solano Co.)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2A2936BD5C2FFFC2FF31CA31FB83F956" blockId="2.[151,1437,1593,1980]" pageId="2" pageNumber="181">
|
||
Pronotum 2.7–3.0 mm wide,
|
||
<specimenCount id="3C90FD345C2FFFC2FDE2CA31FDF6F9E1" box="[532,553,1594,1620]" pageId="2" pageNumber="181" type="female">♀</specimenCount>
|
||
mean
|
||
<quantity id="ED6E9B585C2FFFC2FD81CA32FD0FF9E1" box="[631,720,1593,1620]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="181" unit="mm" value="2.9">2.9 mm</quantity>
|
||
, ♁ mean 2.8. mm, length
|
||
<quantity id="ED6E9B585C2FFFC2FC0DCA32FB59F9E1" box="[1019,1158,1593,1620]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="2" pageNumber="181" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
|
||
,
|
||
<specimenCount id="3C90FD345C2FFFC2FB65CA31FB77F9E1" box="[1171,1192,1594,1620]" pageId="2" pageNumber="181" type="female">♀</specimenCount>
|
||
<quantity id="ED6E9B585C2FFFC2FB46CA32FAD5F9E1" box="[1200,1290,1593,1620]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="181" unit="mm" value="2.8">2.8 mm</quantity>
|
||
, mean ♁
|
||
<quantity id="ED6E9B585C2FFFC2FA8FCA32FF1AF9CD" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="181" unit="mm" value="2.7">2.7 mm</quantity>
|
||
, dark bronze to dark cupreous, sparse white setae present along marginal areas of dorsal surface; disc finely rugose to vermiculate with thin but distinct median line and most strongly impressed anterior and posterior sulci; sulci often with faint to distinct green-blue metallic reflections; notopleural sutures clearly defined, not visible from dorsal view; proepisternum (
|
||
<figureCitation id="B2AD2A385C2FFFC2FE2ECAC2FDEDF956" box="[472,562,1737,1763]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="3.[152,255,2017,2041]" captionStartId-1="4.[152,255,1572,1596]" captionTargetBox-1="[257,1330,813,1547]" captionTargetId-0="figure-40@3.[264,1323,799,1993]" captionTargetId-1="figure-310@4.[257,1330,813,1547]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 3. Aedeagus of C. timbisha, n. sp. (left) and C. senilis (right)." captionText-1="FIGURE 4. Cicindela timbisha n. sp. in situ in dark muddy soil near a freshwater spring." figureDoi-0="http://doi.org/10.5281/zenodo.7971624" figureDoi-1="http://doi.org/10.5281/zenodo.7959907" httpUri-0="https://zenodo.org/record/7971624/files/figure.png" httpUri-1="https://zenodo.org/record/7959907/files/figure.png" pageId="2" pageNumber="181">Fig 3–4</figureCitation>
|
||
) polished cupreous, setae throughout the surface.
|
||
</paragraph>
|
||
<paragraph id="2A2936BD5C2FFFC2FF31CAE6FB69F80E" blockId="2.[151,1437,1593,1980]" pageId="2" pageNumber="181">
|
||
Elytra elongate,
|
||
<quantity id="ED6E9B585C2FFFC2FE89CAE6FDD9F8BD" box="[383,518,1773,1800]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.3" metricValueMin="6.3" pageId="2" pageNumber="181" unit="mm" value="6.8" valueMax="7.3" valueMin="6.3">6.3–7.3 mm</quantity>
|
||
length,
|
||
<specimenCount id="3C90FD345C2FFFC2FD96CAE5FDAAF8BD" box="[608,629,1774,1800]" pageId="2" pageNumber="181" type="female">♀</specimenCount>
|
||
mean
|
||
<quantity id="ED6E9B585C2FFFC2FD49CAE6FCC9F8BD" box="[703,790,1773,1800]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="2" pageNumber="181" unit="mm" value="6.9">6.9 mm</quantity>
|
||
, mean ♁
|
||
<quantity id="ED6E9B585C2FFFC2FC77CAE6FC07F8BD" box="[897,984,1773,1800]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="181" unit="mm" value="6.6">6.6 mm</quantity>
|
||
, shape similar in both sexes, but slightly wider in female, especially toward apical third; sutural spine small, very weak microserrations present on elytral apices; elytral dorsal texture granulate-punctate throughout; faint subsutural foveae present and possessing metallic green reflections. Elytral coloration mostly dark brown with fine green punctures throughout, visible under magnification. Elytral maculations present, consisting of a humeral spot, bold middle band that typically does not meet the lateral margin, bold humeral lunule; sub-humeral spot is generally absent or greatly atrophied.
|
||
</paragraph>
|
||
<caption id="7EE966355C2EFFC3FF6ECEE4FCB5FCB2" ID-DOI="http://doi.org/10.5281/zenodo.7959905" ID-Zenodo-Dep="7959905" box="[152,874,751,775]" httpUri="https://zenodo.org/record/7959905/files/figure.png" pageId="3" pageNumber="182" startId="3.[152,255,751,775]" targetBox="[218,1371,181,719]" targetPageId="3" targetType="figure">
|
||
<paragraph id="2A2936BD5C2EFFC3FF6ECEE4FCB5FCB2" blockId="3.[152,874,751,775]" box="[152,874,751,775]" pageId="3" pageNumber="182">
|
||
<emphasis id="18E2EAAF5C2EFFC3FF6ECEE4FEC7FCB2" bold="true" box="[152,280,751,775]" pageId="3" pageNumber="182">FIGURE 2.</emphasis>
|
||
Frontal habitus of
|
||
<taxonomicName id="ED964D3E5C2EFFC3FE2FCEFBFD91FCB2" authority="Duran & Chambers & Nelson & Roman, 2023" authorityName="Duran & Chambers & Nelson & Roman" authorityYear="2023" box="[473,590,751,775]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
|
||
<emphasis id="18E2EAAF5C2EFFC3FE2FCEFBFD91FCB2" box="[473,590,751,775]" italics="true" pageId="3" pageNumber="182">C. timbisha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="18E2EAAF5C2EFFC3FDADCEE4FD4BFCB2" bold="true" box="[603,660,751,775]" pageId="3" pageNumber="182">
|
||
<taxonomicNameLabel id="03D157D45C2EFFC3FDADCEE4FD4BFCB2" box="[603,660,751,775]" pageId="3" pageNumber="182" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A) male, B) female.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7EE966355C2EFFC3FF6ECBEAFCABF84C" ID-DOI="http://doi.org/10.5281/zenodo.7971624" ID-Zenodo-Dep="7971624" box="[152,884,2017,2042]" httpUri="https://zenodo.org/record/7971624/files/figure.png" pageId="3" pageNumber="182" startId="3.[152,255,2017,2041]" targetBox="[320,1270,832,1945]" targetPageId="3" targetType="figure">
|
||
<paragraph id="2A2936BD5C2EFFC3FF6ECBEAFCABF84C" blockId="3.[152,884,2017,2042]" box="[152,884,2017,2042]" pageId="3" pageNumber="182">
|
||
<emphasis id="18E2EAAF5C2EFFC3FF6ECBEAFEC7F84C" bold="true" box="[152,280,2017,2041]" pageId="3" pageNumber="182">FIGURE 3.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="ED964D3E5C2EFFC3FE55CBE9FDC7F84C" authority="Duran & Chambers & Nelson & Roman, 2023" authorityName="Duran & Chambers & Nelson & Roman" authorityYear="2023" box="[419,536,2017,2041]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
|
||
<emphasis id="18E2EAAF5C2EFFC3FE55CBE9FDC7F84C" box="[419,536,2017,2041]" italics="true" pageId="3" pageNumber="182">C. timbisha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="18E2EAAF5C2EFFC3FDD2CBE9FD81F84F" bold="true" box="[548,606,2018,2042]" pageId="3" pageNumber="182">
|
||
<taxonomicNameLabel id="03D157D45C2EFFC3FDD2CBE9FD81F84F" box="[548,606,2018,2042]" pageId="3" pageNumber="182" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(left) and
|
||
<taxonomicName id="ED964D3E5C2EFFC3FD3ECBE9FCF9F84C" authorityName="G. Horn" authorityYear="1866" box="[712,806,2017,2041]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="182" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C2EFFC3FD3ECBE9FCF9F84C" box="[712,806,2017,2041]" italics="true" pageId="3" pageNumber="182">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
(right).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2A2936BD5C29FFC4FF31CC9CFD0EFED0" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="183">Procoxae and mesocoxae dark metallic green to blue-violet, setae present throughout; metacoxae dark metallic green to blue-violet, with setae present throughout; pro- and meso- trochanters with a single subapical seta; femora dark metallic green with blue and violet reflections, femoral surface with rows of erect white setae dorsally and ventrally; tibiae violet to dark cupreous with dark green reflections near the apices, clothed with white setae that are sparser and shorter than those of the femora; tarsi violet with blue reflections dorsally, first three dilated protarsomeres in male with dense greyish-white setal pads.</paragraph>
|
||
<paragraph id="2A2936BD5C29FFC4FF31CD64FDF3FDAC" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="183">
|
||
Abdominal ventrites 1‒6 with metallic blue, dark blue and violet reflections; fine white setae present mostly along lateral third of each ventrite. Aedeagus (
|
||
<figureCitation id="B2AD2A385C29FFC4FD40CD98FCCCFE18" box="[694,787,403,429]" captionStart="FIGURE 3" captionStartId="3.[152,255,2017,2041]" captionTargetId="figure-40@3.[264,1323,799,1993]" captionTargetPageId="3" captionText="FIGURE 3. Aedeagus of C. timbisha, n. sp. (left) and C. senilis (right)." figureDoi="http://doi.org/10.5281/zenodo.7971624" httpUri="https://zenodo.org/record/7971624/files/figure.png" pageId="4" pageNumber="183">Fig. 3A</figureCitation>
|
||
), possessing a well-developed flagellum and ear-shaped internal sac, similar to
|
||
<taxonomicName id="ED964D3E5C29FFC4FE6ACDBCFDDAFE64" authorityName="G. Horn" authorityYear="1866" box="[412,517,439,465]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C29FFC4FE6ACDBCFDDAFE64" box="[412,517,439,465]" italics="true" pageId="4" pageNumber="183">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B2AD2A385C29FFC4FDE3CDBCFDB1FE64" box="[533,622,439,465]" captionStart="FIGURE 3" captionStartId="3.[152,255,2017,2041]" captionTargetId="figure-40@3.[264,1323,799,1993]" captionTargetPageId="3" captionText="FIGURE 3. Aedeagus of C. timbisha, n. sp. (left) and C. senilis (right)." figureDoi="http://doi.org/10.5281/zenodo.7971624" httpUri="https://zenodo.org/record/7971624/files/figure.png" pageId="4" pageNumber="183">Fig. 3B</figureCitation>
|
||
), characteristics which are similar to most
|
||
<taxonomicName id="ED964D3E5C29FFC4FBA0CDBCFA47FE64" authority="(Rivalier 1954)" baseAuthorityName="Rivalier" baseAuthorityYear="1954" box="[1110,1432,438,465]" class="Insecta" family="Carabidae" genus="Cicindelidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="18E2EAAF5C29FFC4FBA0CDBCFB3FFE64" box="[1110,1248,439,465]" italics="true" pageId="4" pageNumber="183">Cicindelidia</emphasis>
|
||
(
|
||
<bibRefCitation id="4E074B4C5C29FFC4FB06CDBCFA4FFE64" author="Rivalier, E." box="[1264,1424,438,465]" pageId="4" pageNumber="183" pagination="249 - 268" refId="ref3145" refString="Rivalier, E. (1954) Demembrement du genre Cicindela Linne. II. Faune americaine. Revue Francaise d'Entomologie, 21 (4), 249 - 268." type="journal article" year="1954">Rivalier 1954</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, although it has been established by
|
||
<bibRefCitation id="4E074B4C5C29FFC4FDD1CDD0FC9DFE40" author="Duran, D. P. & Gough, H. M." box="[551,834,475,501]" pageId="4" pageNumber="183" pagination="1 - 12" refId="ref2770" refString="Duran, D. P. & Gough, H. M. (2019) Unifying systematics and taxonomy: Nomenclatural changes to Nearctic tiger beetles (Coleoptera: Carabidae: Cicindelinae) based on phylogenetics, morphology and life history. Insecta Mundi, 0727, 1 - 12." type="book chapter" year="2019">Duran and Gough (2019)</bibRefCitation>
|
||
that the
|
||
<taxonomicName id="ED964D3E5C29FFC4FC52CDD0FBD2FE40" authorityName="G. Horn" authorityYear="1866" box="[932,1037,475,501]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="senilis">
|
||
<emphasis id="18E2EAAF5C29FFC4FC52CDD0FBD2FE40" box="[932,1037,475,501]" italics="true" pageId="4" pageNumber="183">C. senilis</emphasis>
|
||
</taxonomicName>
|
||
species group belongs in
|
||
<taxonomicName id="ED964D3E5C29FFC4FAD9CDD0FA43FE40" authorityName="Linne" authorityYear="1758" box="[1327,1436,475,501]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="18E2EAAF5C29FFC4FAD9CDD0FA43FE40" box="[1327,1436,475,501]" italics="true" pageId="4" pageNumber="183">Cicindela</emphasis>
|
||
</taxonomicName>
|
||
based on DNA and life-history data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="628C65365C29FFC4FF31CE28FC7FFDD4" pageId="4" pageNumber="183" type="etymology">
|
||
<paragraph id="2A2936BD5C29FFC4FF31CE28FC7FFDD4" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="183">
|
||
<emphasis id="18E2EAAF5C29FFC4FF31CE28FE91FD88" bold="true" box="[199,334,547,573]" pageId="4" pageNumber="183">Etymology.</emphasis>
|
||
<taxonomicName id="ED964D3E5C29FFC4FEA3CE28FDF5FD88" authority="Duran & Chambers & Nelson & Roman, 2023" authorityName="Duran & Chambers & Nelson & Roman" authorityYear="2023" box="[341,554,547,573]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
|
||
<emphasis id="18E2EAAF5C29FFC4FEA3CE28FDF5FD88" box="[341,554,547,573]" italics="true" pageId="4" pageNumber="183">Cicindela timbisha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="18E2EAAF5C29FFC4FDCFCE28FDA7FD88" bold="true" box="[569,632,547,573]" pageId="4" pageNumber="183">
|
||
<taxonomicNameLabel id="03D157D45C29FFC4FDCFCE28FDA7FD88" box="[569,632,547,573]" pageId="4" pageNumber="183" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is named for the Timbisha-Shoshone Tribe of Death Valley, as the new species was found at a locality that falls within their aboriginal lands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="628C65365C29FFC4FF31CE61FA59FCA0" pageId="4" pageNumber="183" type="distribution">
|
||
<paragraph id="2A2936BD5C29FFC4FF31CE61FA59FCA0" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="183">
|
||
<emphasis id="18E2EAAF5C29FFC4FF31CE61FE31FD31" bold="true" box="[199,494,618,645]" pageId="4" pageNumber="183">Distribution and habitat</emphasis>
|
||
.
|
||
<taxonomicName id="ED964D3E5C29FFC4FE0BCE60FD0DFD30" authority="Duran & Chambers & Nelson & Roman, 2023" authorityName="Duran & Chambers & Nelson & Roman" authorityYear="2023" box="[509,722,619,645]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="183" phylum="Arthropoda" rank="species" species="timbisha" status="n. sp.">
|
||
<emphasis id="18E2EAAF5C29FFC4FE0BCE60FD0DFD30" box="[509,722,619,645]" italics="true" pageId="4" pageNumber="183">Cicindela timbisha</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="18E2EAAF5C29FFC4FD2DCE60FCC4FD30" bold="true" box="[731,795,619,645]" pageId="4" pageNumber="183">
|
||
<taxonomicNameLabel id="03D157D45C29FFC4FD2DCE60FCC4FD30" box="[731,795,619,645]" pageId="4" pageNumber="183" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from a single locality along a freshwater spring in Inyo County,
|
||
<collectingRegion id="E852F85F5C29FFC4FE6ECE84FDD3FD1C" box="[408,524,655,681]" country="United States of America" name="California" pageId="4" pageNumber="183">California</collectingRegion>
|
||
, east of the Sierra
|
||
<collectingRegion id="E852F85F5C29FFC4FD2CCE84FCEFFD1C" box="[730,816,655,681]" country="United States of America" name="Nevada" pageId="4" pageNumber="183">Nevada</collectingRegion>
|
||
Mountains. Beetles were observed ovipositing in damp dark soils along the freshwater spring (
|
||
<figureCitation id="B2AD2A385C29FFC4FDB3CEB8FD5CFD78" box="[581,643,691,717]" captionStart="FIGURE 4" captionStartId="4.[152,255,1572,1596]" captionTargetBox="[257,1330,813,1547]" captionTargetId="figure-310@4.[257,1330,813,1547]" captionTargetPageId="4" captionText="FIGURE 4. Cicindela timbisha n. sp. in situ in dark muddy soil near a freshwater spring." figureDoi="http://doi.org/10.5281/zenodo.7959907" httpUri="https://zenodo.org/record/7959907/files/figure.png" pageId="4" pageNumber="183">Fig 4</figureCitation>
|
||
). No additional areas of suitable habitat could be located in the vicinity. Due to the highly restricted geographic distribution of the species (<5 ha of available habitat), we cannot provide any additional details about the location. The species is at great risk of extinction due to potential overcollection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |