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<document id="196CFC6E3220E11D95A9E421A498EC39" ID-DOI="10.37828/em.2019.20.18" ID-ISSN="2336-9744" ID-Zenodo-Dep="13359824" ID-ZooBank="urn:lsid:zoobank.org:pub:58760F6A-D4DA-4844-A373-E93E622EDE4C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1720660543355" checkinUser="felipe" docAuthor="Volynkin, Anton V. &amp; László, Gyula M." docDate="2019" docId="A237E31AFFC1FFFDA096FDF9D0EEF81A" docLanguage="en" docName="EcolMontenegrina.20.215-221.pdf" docOrigin="Ecologica Montenegrina 20 (2019)" docSource="http://dx.doi.org/10.37828/em.2019.20.18" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.3:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="3" docTitle="Cyana (Louisia) smithi Volynkin &amp; Laszlo 2019, sp. nov." docType="treatment" docUuid="C0FD5783-9C23-47DC-ACD0-22DA8752D329" docUuidSource="ZooBank" docVersion="2" lastPageNumber="220" masterDocId="5E0E9B62FFC0FFF8A001FFA0D44BFFA2" masterDocTitle="Cyana smithi, a new species from Liberia (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="221" masterPageNumber="215" pageNumber="216" updateTime="1724326691582" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="F9A36323EB40C99DC7ABC88119E73549" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D4FE9B68FB99CCFAC9AD8CA364ED5EA6">
<mods:title id="2E1655BF4A59823E98A23758C3BD7246">Cyana smithi, a new species from Liberia (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
</mods:titleInfo>
<mods:name id="524CC32759CF806B0F431A0EECF4DD92" type="personal">
<mods:role id="0BC102432EE76CED976440648F5ABA11">
<mods:roleTerm id="4F596FB1CDF41C1371B9E49AF40B987C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8EC898C6FC1F51CFA78393F8BF20A398">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="889308B21DC482E1ED4971ECA9382265">Altai State University, Lenina Avenue 61, RF- 656049, Barnaul, Russia. E-mail: volynkin _ a @ mail. ru &amp; Tomsk State University, Lenina Avenue, 36, RF- 634050, Tomsk, Russia &amp; Corresponding author</mods:affiliation>
</mods:name>
<mods:name id="FC285F6B10331FBFCDDC73BD3688DEC3" type="personal">
<mods:role id="B8B03202B04B9D08D39528256FCE0B87">
<mods:roleTerm id="F2F3A9718000405597095E1C54328850">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="11806EDA2BA9EA63EEEC025B7700B488">László, Gyula M.</mods:namePart>
<mods:affiliation id="A07665C18A7C5B15EEAFCF1158F67733">The African Natural History Research Trust (ANHRT), Street Court Leominster-Kingsland, HR 6 9 QA, United Kingdom. E-mails: gyula @ anhrt. org. uk, gyula. m. laszlo @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5F0C05D346EB15596AB7EF9C0B767A4E">text</mods:typeOfResource>
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<mods:titleInfo id="4DB8BA0DD7D97D86B141CC400B7BC736">
<mods:title id="46630B0BFEF1DBA602C09B1777764B36">Ecologica Montenegrina</mods:title>
</mods:titleInfo>
<mods:part id="3C4FDFED6646F8DE7045D2863511C17F">
<mods:date id="EBF87AFC3B1AB1D8225A86EBE8B1F5FE">2019</mods:date>
<mods:detail id="6D7725F3B574BB5EFF91D1F1FA38F753" type="pubDate">
<mods:number id="68334A16767312C15E110166FCB10EF5">2019-03-29</mods:number>
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<mods:number id="7908EA8977DCBBDD78EB7A6CB902FFF9">20</mods:number>
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<mods:detail id="9733505BEDF1FA988A2590D7459A8D3E" type="issue">
<mods:number id="0D6489E27131E2F511DFE8B8C6BED6DE">2019</mods:number>
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<mods:start id="98074099B0A2D9AF83B09D659359A120">215</mods:start>
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<mods:url id="D0B7B261F32EC3430B9F8244A37CFA88">http://dx.doi.org/10.37828/em.2019.20.18</mods:url>
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<mods:classification id="B5D731B3E35F08CEA77D4703C431ADD0">journal article</mods:classification>
<mods:identifier id="2EBD0A381E73FF37461873C07C600939" type="DOI">10.37828/em.2019.20.18</mods:identifier>
<mods:identifier id="1C5D47DFA55614E1ACF0241C4713274D" type="ISSN">2336-9744</mods:identifier>
<mods:identifier id="2CF0E62D2F9F40254364CC27AFC8FCC8" type="Zenodo-Dep">13359824</mods:identifier>
<mods:identifier id="74BF267F3FCBC1E823F5A4734A885EC8" type="ZooBank">urn:lsid:zoobank.org:pub:58760F6A-D4DA-4844-A373-E93E622EDE4C</mods:identifier>
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<treatment id="A237E31AFFC1FFFDA096FDF9D0EEF81A" ID-DOI="http://doi.org/10.5281/zenodo.12717246" ID-Zenodo-Dep="12717246" LSID="urn:lsid:zoobank.org:act:C0FD5783-9C23-47DC-ACD0-22DA8752D329" httpUri="http://treatment.plazi.org/id/A237E31AFFC1FFFDA096FDF9D0EEF81A" lastPageId="5" lastPageNumber="220" pageId="1" pageNumber="216">
<subSubSection id="62840187FFC1FFF9A096FDF9D74DFDD1" box="[151,774,600,627]" pageId="1" pageNumber="216" type="nomenclature">
<paragraph id="2A21520CFFC1FFF9A096FDF9D74DFDD1" blockId="1.[151,774,600,661]" box="[151,774,600,627]" pageId="1" pageNumber="216">
<heading id="7169E560FFC1FFF9A096FDF9D74DFDD1" box="[151,774,600,627]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="216" reason="2">
<taxonomicName id="ED9E298FFFC1FFF9A096FDF9D6D1FDD1" ID-CoL="BFNVS" authority="Volynkin &amp; Laszlo" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[151,666,600,627]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi" status="sp. nov." subGenus="Louisia">
<emphasis id="18EA8E1EFFC1FFF9A096FDF9D5E5FDD1" bold="true" box="[151,430,601,627]" pageId="1" pageNumber="216">
<emphasis id="18EA8E1EFFC1FFF9A096FDF9D4AEFDD1" bold="true" box="[151,229,601,627]" italics="true" pageId="1" pageNumber="216">Cyana</emphasis>
(
<emphasis id="18EA8E1EFFC1FFF9A0F6FDF9D519FDD1" bold="true" box="[247,338,601,627]" italics="true" pageId="1" pageNumber="216">Louisia</emphasis>
)
<emphasis id="18EA8E1EFFC1FFF9A162FDF9D5E5FDD1" bold="true" box="[355,430,601,627]" italics="true" pageId="1" pageNumber="216">smithi</emphasis>
</emphasis>
Volynkin &amp; László
</taxonomicName>
,
<emphasis id="18EA8E1EFFC1FFF9A2A6FDF9D74DFDD1" bold="true" box="[679,774,601,627]" pageId="1" pageNumber="216">
<taxonomicNameLabel id="03D93365FFC1FFF9A2A6FDF9D74DFDD1" box="[679,774,601,627]" pageId="1" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="62840187FFC1FFF9A096FDDAD595FD37" box="[151,478,634,661]" pageId="1" pageNumber="216" type="description">
<paragraph id="2A21520CFFC1FFF9A096FDDAD595FD37" blockId="1.[151,774,600,661]" box="[151,478,634,661]" pageId="1" pageNumber="216">
(
<figureCitation id="B2A54E89FFC1FFF9A09EFDDAD54CFD37" box="[159,263,634,661]" captionStart="Figures 1–8" captionStartId="2.[151,237,1933,1957]" captionTargetBox="[169,1399,190,1903]" captionTargetId="figure-17@2.[157,1431,189,1914]" captionTargetPageId="2" captionText="Figures 1–8. Cyana spp.: adults. 1, C. smithi, holotype male, Liberia (ANHRT); 2, C. smithi, paratype male, Liberia (ANHRT); 3, C. smithi, paratype male, Liberia (ANHRT); 4, C. smithi, paratype female, Liberia (ANHRT); 5, C. quentini, male, DR Congo (MWM/ZSM); 6, C. quentini, male, Cameroon (ANHRT); 7, C. quentini, female, Cameroon (ANHRT); 8, C. yaseminae, paratype male, Uganda (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13359826" httpUri="https://zenodo.org/record/13359826/files/figure.png" pageId="1" pageNumber="216">Figs 1–4</figureCitation>
,
<figureCitation id="B2A54E89FFC1FFF9A117FDDAD51BFD37" box="[278,336,634,661]" captionStart="Figures 9–12" captionStartId="3.[151,237,1949,1973]" captionTargetBox="[208,1413,226,1924]" captionTargetId="figure-8@3.[151,1413,223,1931]" captionTargetPageId="3" captionText="Figures 9–12. Cyana spp.: male genitalia. 9, C. smithi, holotype, Liberia, slide AV4482 Volynkin; 10, C. smithi, paratype, Liberia, slide AV4453 Volynkin; 11, C. quentini, DR Congo, slide MWM 35737 Volynkin; 12, C. quentini, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359828" httpUri="https://zenodo.org/record/13359828/files/figure.png" pageId="1" pageNumber="216">9, 10</figureCitation>
,
<figureCitation id="B2A54E89FFC1FFF9A15EFDDAD537FD37" box="[351,380,634,661]" captionStart="Figures 13–16" captionStartId="4.[151,237,1552,1576]" captionTargetBox="[151,1436,197,1523]" captionTargetId="figure-17@4.[151,1436,189,1534]" captionTargetPageId="4" captionText="Figures 13–16. Cyana spp.: male genitalia. 13, C. yaseminae, paratype, Uganda, slide BMNH(E) Arct-6196 Karisch (©NHMUK); 14, C. yaseminae, paratype, Uganda, slide 3300 Karisch (photo by T. Karisch); 15, C. smithi: magnified juxta, paratype, Liberia, slide AV4990 Volynkin; 16, C. quentini: magnified juxta, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359832" httpUri="https://zenodo.org/record/13359832/files/figure.png" pageId="1" pageNumber="216">15</figureCitation>
,
<figureCitation id="B2A54E89FFC1FFF9A18AFDDAD59FFD37" box="[395,468,634,661]" captionStart="Figures 17–19" captionStartId="5.[151,237,1689,1713]" captionTargetBox="[151,1436,897,1656]" captionTargetId="figure-283@5.[151,1436,897,1669]" captionTargetPageId="5" captionText="Figures 17–19. Cyana spp.: female genitalia. 17, C. smithi, paratype, Liberia, slide AV4992 Volynkin; 18, C. quentini, Cameroon, slide AV5078 Voynkin; 19, C. smithi: magnified posterior part of corpus bursae, paratype, Liberia, slide AV4992 Volynkin." figureDoi="http://doi.org/10.5281/zenodo.13359834" httpUri="https://zenodo.org/record/13359834/files/figure.png" pageId="1" pageNumber="216">17, 19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="62840187FFC1FFF9A096FD1ED616FAB4" pageId="1" pageNumber="216" type="materials_examined">
<paragraph id="2A21520CFFC1FFF9A096FD1ED545FC9F" blockId="1.[151,1436,701,829]" pageId="1" pageNumber="216">
<emphasis id="18EA8E1EFFC1FFF9A096FD1ED51AFD7A" bold="true" box="[151,337,702,728]" pageId="1" pageNumber="216">Type material.</emphasis>
<materialsCitation id="9AF65851FFC1FFF9A15BFD1ED545FC9F" collectingDate="2017-11-20" collectingDateMax="2017-12-01" collectingDateMin="2017-11-20" collectionCode="MV, ANHRT" collectorName="Aristophanous, M. &amp; Safian, Sz. &amp; Simonics, G. &amp; Smith, L." country="Liberia" elevation="611" latitude="8.121389" location="Light Trap" longLatPrecision="21" longitude="-9.961666" municipality="Wologizi Mts." pageId="1" pageNumber="216" specimenCount="1" specimenCount-male="1" stateProvince="Lofa" typeStatus="holotype">
<emphasis id="18EA8E1EFFC1FFF9A15BFD1ED587FD7A" bold="true" box="[346,460,702,728]" pageId="1" pageNumber="216">
<typeStatus id="F525ECAEFFC1FFF9A15BFD1ED587FD7A" box="[346,460,702,728]" pageId="1" pageNumber="216" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="B2A54E89FFC1FFF9A1DFFD1DD660FD7A" box="[478,555,701,728]" captionStart="Figures 1–8" captionStartId="2.[151,237,1933,1957]" captionTargetBox="[169,1399,190,1903]" captionTargetId="figure-17@2.[157,1431,189,1914]" captionTargetPageId="2" captionText="Figures 1–8. Cyana spp.: adults. 1, C. smithi, holotype male, Liberia (ANHRT); 2, C. smithi, paratype male, Liberia (ANHRT); 3, C. smithi, paratype male, Liberia (ANHRT); 4, C. smithi, paratype female, Liberia (ANHRT); 5, C. quentini, male, DR Congo (MWM/ZSM); 6, C. quentini, male, Cameroon (ANHRT); 7, C. quentini, female, Cameroon (ANHRT); 8, C. yaseminae, paratype male, Uganda (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13359826" httpUri="https://zenodo.org/record/13359826/files/figure.png" pageId="1" pageNumber="216">Figs 1</figureCitation>
,
<figureCitation id="B2A54E89FFC1FFF9A23AFD1DD601FD7A" box="[571,586,701,728]" captionStart="Figures 9–12" captionStartId="3.[151,237,1949,1973]" captionTargetBox="[208,1413,226,1924]" captionTargetId="figure-8@3.[151,1413,223,1931]" captionTargetPageId="3" captionText="Figures 9–12. Cyana spp.: male genitalia. 9, C. smithi, holotype, Liberia, slide AV4482 Volynkin; 10, C. smithi, paratype, Liberia, slide AV4453 Volynkin; 11, C. quentini, DR Congo, slide MWM 35737 Volynkin; 12, C. quentini, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359828" httpUri="https://zenodo.org/record/13359828/files/figure.png" pageId="1" pageNumber="216">9</figureCitation>
): male,
<collectingCountry id="5289129CFFC1FFF9A2AEFD1DD74CFD7A" box="[687,775,701,728]" name="Liberia" pageId="1" pageNumber="216">Liberia</collectingCountry>
,
<quantity id="ED66FFE9FFC1FFF9A315FD1DD711FD7A" box="[788,858,701,728]" metricMagnitude="2" metricUnit="m" metricValue="6.11" pageId="1" pageNumber="216" unit="m" value="611.0">
<elevation id="A1B3B53FFFC1FFF9A315FD1DD711FD7A" box="[788,858,701,728]" metricMagnitude="2" metricUnit="m" metricValue="6.11" pageId="1" pageNumber="216" unit="m" value="611.0">611m</elevation>
</quantity>
,
<collectingRegion id="E85A9CEEFFC1FFF9A366FD1DD04AFD7A" box="[871,1025,701,728]" country="Liberia" name="Lofa" pageId="1" pageNumber="216">Lofa County</collectingRegion>
,
<collectingMunicipality id="CA45C876FFC1FFF9A40EFD1DD0F6FD7A" box="[1039,1213,701,728]" pageId="1" pageNumber="216">Wologizi Mts.</collectingMunicipality>
, base camp forest,
<geoCoordinate id="4FAA34CBFFC1FFF9A096FD7FD56BFD58" box="[151,288,735,762]" degrees="8" direction="north" minutes="07" orientation="latitude" pageId="1" pageNumber="216" precision="15" seconds="17" value="8.121389">8°07’17’’N</geoCoordinate>
,
<geoCoordinate id="4FAA34CBFFC1FFF9A134FD7FD58DFD58" box="[309,454,735,762]" degrees="9" direction="west" minutes="57" orientation="longitude" pageId="1" pageNumber="216" precision="15" seconds="42" value="-9.961666">9°57’42’’W</geoCoordinate>
,
<date id="5E2074CCFFC1FFF9A1D8FD7FD68FFD58" box="[473,708,735,762]" pageId="1" pageNumber="216" value="2017-11-20" valueMax="2017-12-01" valueMin="2017-11-20">
<collectingDate id="4E648D24FFC1FFF9A1D8FD7FD68FFD58" box="[473,708,735,762]" pageId="1" pageNumber="216" value="2017-11-20" valueMax="2017-12-01" valueMin="2017-11-20">20.XI.–01.XII.2017</collectingDate>
</date>
,
<collectionCode id="4C8FCAC9FFC1FFF9A2D9FD40D743FD58" box="[728,776,736,762]" country="0" name="University of Montana Museum" pageId="1" pageNumber="216">MV</collectionCode>
<location id="2F4104D7FFC1FFF9A317FD7FD7D6FD58" LSID="urn:lsid:plazi:treatment:A237E31AFFC1FFFDA096FDF9D0EEF81A:2F4104D7FFC1FFF9A317FD7FD7D6FD58" box="[790,925,735,762]" country="Liberia" latitude="8.121389" longLatPrecision="21" longitude="-9.961666" municipality="Wologizi Mts." name="Light Trap" pageId="1" pageNumber="216" stateProvince="Lofa">Light Trap</location>
(125w),
<collectorName id="876B37DAFFC1FFF9A417FD7FD0B0FD58" box="[1046,1275,735,762]" pageId="1" pageNumber="216">Aristophanous, M.</collectorName>
,
<collectorName id="876B37DAFFC1FFF9A50EFD7FD1DDFD58" box="[1295,1430,735,762]" pageId="1" pageNumber="216">Sáfián, Sz.</collectorName>
,
<collectorName id="876B37DAFFC1FFF9A096FCA1D57AFCBE" box="[151,305,769,796]" pageId="1" pageNumber="216">Simonics, G.</collectorName>
,
<collectorName id="876B37DAFFC1FFF9A13FFCA1D5E7FCBE" box="[318,428,769,796]" pageId="1" pageNumber="216">Smith, L.</collectorName>
leg.,
<collectionCode id="4C8FCAC9FFC1FFF9A1EFFCA1D61CFCBE" box="[494,599,769,796]" pageId="1" pageNumber="216">ANHRT</collectionCode>
:2017.33 / ANHRTUK 00019056, slide No.: AV4482
<specimenCount id="3C989985FFC1FFF9A4CFFCA1D0AAFCBE" box="[1230,1249,769,796]" count="1" pageId="1" pageNumber="216" type="male">♂</specimenCount>
Volynkin (coll.
<collectionCode id="4C8FCAC9FFC1FFF9A096FC82D549FC9F" box="[151,258,802,829]" pageId="1" pageNumber="216">ANHRT</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="2A21520CFFC1FFF9A096FCC6D616FAB4" blockId="1.[151,1437,870,1302]" pageId="1" pageNumber="216">
<materialsCitation id="9AF65851FFC1FFF9A096FCC6D09CFC01" collectingDate="2017-11-20" collectingDateMax="2017-12-01" collectingDateMin="2017-11-20" collectionCode="ANHRT" collectorName="No." country="Liberia" elevation="611" latitude="8.121389" location="Light Trap" longLatPrecision="21" longitude="-9.961666" municipality="Wologizi Mts." pageId="1" pageNumber="216" specimenCount="7" specimenCount-male="7" stateProvince="Lofa" typeStatus="paratype">
<emphasis id="18EA8E1EFFC1FFF9A096FCC6D55EFC22" bold="true" box="[151,277,870,896]" pageId="1" pageNumber="216">
<typeStatus id="F525ECAEFFC1FFF9A096FCC6D55EFC22" box="[151,277,870,896]" pageId="1" pageNumber="216" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="3C989985FFC1FFF9A126FCC7D5C3FC23" box="[295,392,870,897]" count="7" pageId="1" pageNumber="216" type="male">7 males</specimenCount>
, with same data as the holotype (
<collectionCode id="4C8FCAC9FFC1FFF9A320FCC6D7C3FC23" box="[801,904,870,897]" pageId="1" pageNumber="216">ANHRT</collectionCode>
unique id numbers: ANHRTUK 00020212, 00026543, 00020215, 00020305, 00020472, 00020207, 00020214), slide
<collectorName id="876B37DAFFC1FFF9A42CFC29D017FC01" box="[1069,1116,905,931]" pageId="1" pageNumber="216">No.</collectorName>
: AV4990
</materialsCitation>
<specimenCount id="3C989985FFC1FFF9A4D6FC28D0A1FC01" box="[1239,1258,904,931]" count="1" pageId="1" pageNumber="216" type="male">♂</specimenCount>
Volynkin;
<materialsCitation id="9AF65851FFC1FFF9A57EFC28D52DFC44" collectingDate="2017-11-10" collectingDateMax="2017-11-19" collectingDateMin="2017-11-10" collectionCode="MV" country="Liberia" elevation="530" location="Foya Proposed Protected Area" longitude="-10.276667" pageId="1" pageNumber="216" specimenCount="16" specimenCount-male="16" stateProvince="Lofa" typeStatus="paratype">
<specimenCount id="3C989985FFC1FFF9A57EFC28D495FC66" count="16" pageId="1" pageNumber="216" type="male">16 males</specimenCount>
,
<collectingCountry id="5289129CFFC1FFF9A0F7FC09D505FC66" box="[246,334,937,964]" name="Liberia" pageId="1" pageNumber="216">Liberia</collectingCountry>
,
<quantity id="ED66FFE9FFC1FFF9A164FC09D5E0FC66" box="[357,427,937,964]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="216" unit="m" value="530.0">
<elevation id="A1B3B53FFFC1FFF9A164FC09D5E0FC66" box="[357,427,937,964]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="216" unit="m" value="530.0">530m</elevation>
</quantity>
,
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, 7°56’36’N,
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, G.,
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leg.,
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2017.33 (
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unique id numbers: ANHRTUK00019046, 00019047, 00020088, 00026301, 00026465, 00026294, 00026282, 00026320, 00026408, 00026407, 00026406, 00026479, 00026480, 00020084, 00025905, 00009091), slide
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Volynkin;
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,
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(blended bulb 250W),
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leg.,
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:2017.33 / ANHRTUK 00020164, slide No.: AV4992
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Volynkin;
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<materialsCitation id="9AF65851FFC1FFF9A4CAFB17D6C6FB56" collectionCode="ANHRT" collectorName="Sz. &amp; Simonics, G." country="Liberia" location="Safian" municipality="Light Trap" pageId="1" pageNumber="216" specimenCount="1" stateProvince="Lofa" typeStatus="paratype">
<collectingMunicipality id="CA45C876FFC1FFF9A4CAFB17D11BFB70" box="[1227,1360,1207,1234]" pageId="1" pageNumber="216">Light Trap</collectingMunicipality>
(8w),
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,
<collectorName id="876B37DAFFC1FFF9A0F8FB79D554FB56" box="[249,287,1241,1268]" pageId="1" pageNumber="216">Sz.</collectorName>
,
<collectorName id="876B37DAFFC1FFF9A135FB79D59FFB56" box="[308,468,1241,1268]" pageId="1" pageNumber="216">Simonics, G.</collectorName>
leg.,
<collectionCode id="4C8FCAC9FFC1FFF9A224FB79D6C6FB56" box="[549,653,1241,1268]" pageId="1" pageNumber="216">ANHRT</collectionCode>
</materialsCitation>
:2018.43 (
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unique id numbers: ANHRTUK 00060192, 00060193, 00061769) (coll.
<collectionCode id="4C8FCAC9FFC1FFF9A1E7FB5BD61AFAB4" box="[486,593,1275,1302]" pageId="1" pageNumber="216">ANHRT</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="62840187FFC1FFF9A096FA9FD547F81B" pageId="1" pageNumber="216" type="diagnosis">
<paragraph id="2A21520CFFC1FFF9A096FA9FD547F81B" blockId="1.[151,1437,1342,1977]" pageId="1" pageNumber="216">
<emphasis id="18EA8E1EFFC1FFF9A096FA9FD55BFAFB" bold="true" box="[151,272,1343,1369]" pageId="1" pageNumber="216">Diagnosis</emphasis>
.
<taxonomicName id="ED9E298FFFC1FFF9A11FFA9FD5F0FAFB" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[286,443,1342,1369]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A11FFA9FD5F0FAFB" box="[286,443,1342,1369]" italics="true" pageId="1" pageNumber="216">Cyana smithi</emphasis>
</taxonomicName>
(
<figureCitation id="B2A54E89FFC1FFF9A1CBFA9ED67FFAFB" box="[458,564,1342,1369]" captionStart="Figures 1–8" captionStartId="2.[151,237,1933,1957]" captionTargetBox="[169,1399,190,1903]" captionTargetId="figure-17@2.[157,1431,189,1914]" captionTargetPageId="2" captionText="Figures 1–8. Cyana spp.: adults. 1, C. smithi, holotype male, Liberia (ANHRT); 2, C. smithi, paratype male, Liberia (ANHRT); 3, C. smithi, paratype male, Liberia (ANHRT); 4, C. smithi, paratype female, Liberia (ANHRT); 5, C. quentini, male, DR Congo (MWM/ZSM); 6, C. quentini, male, Cameroon (ANHRT); 7, C. quentini, female, Cameroon (ANHRT); 8, C. yaseminae, paratype male, Uganda (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13359826" httpUri="https://zenodo.org/record/13359826/files/figure.png" pageId="1" pageNumber="216">Figs 1–4</figureCitation>
) is the third member of the subgenus
<taxonomicName id="ED9E298FFFC1FFF9A3F0FAE0D002FAFB" authorityName="Karisch" authorityYear="2013" box="[1009,1097,1344,1369]" pageId="1" pageNumber="216" rank="subGenus" subGenus="Louisia">
<emphasis id="18EA8E1EFFC1FFF9A3F0FAE0D002FAFB" box="[1009,1097,1344,1369]" italics="true" pageId="1" pageNumber="216">Louisia</emphasis>
</taxonomicName>
. It does not show significant external differences from
<taxonomicName id="ED9E298FFFC1FFF9A1DCFAC1D62CFAD8" authorityName="Karisch" authorityYear="2003" box="[477,615,1377,1403]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A1DCFAC1D62CFAD8" box="[477,615,1377,1403]" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
(
<figureCitation id="B2A54E89FFC1FFF9A27FFAC0D6A5FAD9" box="[638,750,1376,1403]" captionStart="Figures 1–8" captionStartId="2.[151,237,1933,1957]" captionTargetBox="[169,1399,190,1903]" captionTargetId="figure-17@2.[157,1431,189,1914]" captionTargetPageId="2" captionText="Figures 1–8. Cyana spp.: adults. 1, C. smithi, holotype male, Liberia (ANHRT); 2, C. smithi, paratype male, Liberia (ANHRT); 3, C. smithi, paratype male, Liberia (ANHRT); 4, C. smithi, paratype female, Liberia (ANHRT); 5, C. quentini, male, DR Congo (MWM/ZSM); 6, C. quentini, male, Cameroon (ANHRT); 7, C. quentini, female, Cameroon (ANHRT); 8, C. yaseminae, paratype male, Uganda (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13359826" httpUri="https://zenodo.org/record/13359826/files/figure.png" pageId="1" pageNumber="216">Figs 5–7</figureCitation>
) and can be distinguished by genitalia structure only. Compared to
<taxonomicName id="ED9E298FFFC1FFF9A13AFA23D590FA3E" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[315,475,1411,1437]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A13AFA23D590FA3E" box="[315,475,1411,1437]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
(
<figureCitation id="B2A54E89FFC1FFF9A1ECFA22D67DFA3F" box="[493,566,1410,1437]" captionStart="Figures 1–8" captionStartId="2.[151,237,1933,1957]" captionTargetBox="[169,1399,190,1903]" captionTargetId="figure-17@2.[157,1431,189,1914]" captionTargetPageId="2" captionText="Figures 1–8. Cyana spp.: adults. 1, C. smithi, holotype male, Liberia (ANHRT); 2, C. smithi, paratype male, Liberia (ANHRT); 3, C. smithi, paratype male, Liberia (ANHRT); 4, C. smithi, paratype female, Liberia (ANHRT); 5, C. quentini, male, DR Congo (MWM/ZSM); 6, C. quentini, male, Cameroon (ANHRT); 7, C. quentini, female, Cameroon (ANHRT); 8, C. yaseminae, paratype male, Uganda (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13359826" httpUri="https://zenodo.org/record/13359826/files/figure.png" pageId="1" pageNumber="216">Fig. 8</figureCitation>
),
<taxonomicName id="ED9E298FFFC1FFF9A24EFA23D6F0FA3F" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[591,699,1410,1437]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A24EFA23D6F0FA3F" box="[591,699,1410,1437]" italics="true" pageId="1" pageNumber="216">C. smithi</emphasis>
</taxonomicName>
is somewhat smaller in size (male’s wingspan of
<taxonomicName id="ED9E298FFFC1FFF9A515FA23D1CBFA3F" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[1300,1408,1410,1437]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A515FA23D1CBFA3F" box="[1300,1408,1410,1437]" italics="true" pageId="1" pageNumber="216">C. smithi</emphasis>
</taxonomicName>
is
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, while that of
<taxonomicName id="ED9E298FFFC1FFF9A1CEFA04D63BFA1C" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[463,624,1444,1470]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A1CEFA04D63BFA1C" box="[463,624,1444,1470]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
is
<quantity id="ED66FFE9FFC1FFF9A298FA04D757FA1D" box="[665,796,1444,1471]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="1" pageNumber="216" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18–19 mm</quantity>
). In addition, the forewing costa of
<taxonomicName id="ED9E298FFFC1FFF9A4D6FA04D10FFA1D" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[1239,1348,1444,1471]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A4D6FA04D10FFA1D" box="[1239,1348,1444,1471]" italics="true" pageId="1" pageNumber="216">C. smithi</emphasis>
</taxonomicName>
is pale orange in the underside (similar to that of
<taxonomicName id="ED9E298FFFC1FFF9A28AFA66D745FA7D" authorityName="Karisch" authorityYear="2003" box="[651,782,1478,1504]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A28AFA66D745FA7D" box="[651,782,1478,1504]" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
), whereas that area is dark brown in
<taxonomicName id="ED9E298FFFC1FFF9A4C6FA66D12DFA7D" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[1223,1382,1478,1504]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A4C6FA66D12DFA7D" box="[1223,1382,1478,1504]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="4E0F2FFDFFC1FFF9A096FA48D5E1F9A0" author="Baron, T. R. &amp; Karisch, T." box="[151,426,1511,1538]" pageId="1" pageNumber="216" pagination="31 - 33" refId="ref3076" refString="Baron, T. R. &amp; Karisch, T. (2016) A new species of Cyana Walker, 1854 from Uganda (Lepidoptera: Erebidae: Arctiinae). Metamorphosis, 27, 31 - 33." type="journal article" year="2016">Baron &amp; Karisch 2016</bibRefCitation>
: fig. 4). In the male clasping apparatus,
<taxonomicName id="ED9E298FFFC1FFF9A38BFA48D7BDF9A0" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[906,1014,1511,1538]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A38BFA48D7BDF9A0" box="[906,1014,1511,1538]" italics="true" pageId="1" pageNumber="216">C. smithi</emphasis>
</taxonomicName>
(
<figureCitation id="B2A54E89FFC1FFF9A407FA47D0CBF9A0" box="[1030,1152,1511,1538]" captionStart="Figures 9–12" captionStartId="3.[151,237,1949,1973]" captionTargetBox="[208,1413,226,1924]" captionTargetId="figure-8@3.[151,1413,223,1931]" captionTargetPageId="3" captionText="Figures 9–12. Cyana spp.: male genitalia. 9, C. smithi, holotype, Liberia, slide AV4482 Volynkin; 10, C. smithi, paratype, Liberia, slide AV4453 Volynkin; 11, C. quentini, DR Congo, slide MWM 35737 Volynkin; 12, C. quentini, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359828" httpUri="https://zenodo.org/record/13359828/files/figure.png" pageId="1" pageNumber="216">Figs 9, 10</figureCitation>
) differs clearly from
<taxonomicName id="ED9E298FFFC1FFF9A580FA48D4BCF981" authorityName="Karisch" authorityYear="2003" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A580FA48D4BCF981" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
(
<figureCitation id="B2A54E89FFC1FFF9A109F9A9D5DBF986" box="[264,400,1545,1572]" captionStart="Figures 9–12" captionStartId="3.[151,237,1949,1973]" captionTargetBox="[208,1413,226,1924]" captionTargetId="figure-8@3.[151,1413,223,1931]" captionTargetPageId="3" captionText="Figures 9–12. Cyana spp.: male genitalia. 9, C. smithi, holotype, Liberia, slide AV4482 Volynkin; 10, C. smithi, paratype, Liberia, slide AV4453 Volynkin; 11, C. quentini, DR Congo, slide MWM 35737 Volynkin; 12, C. quentini, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359828" httpUri="https://zenodo.org/record/13359828/files/figure.png" pageId="1" pageNumber="216">Figs 11, 12</figureCitation>
) and
<taxonomicName id="ED9E298FFFC1FFF9A1D5F9AAD638F981" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[468,627,1546,1572]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A1D5F9AAD638F981" box="[468,627,1546,1572]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
(
<figureCitation id="B2A54E89FFC1FFF9A285F9A9D740F986" box="[644,779,1545,1572]" captionStart="Figures 13–16" captionStartId="4.[151,237,1552,1576]" captionTargetBox="[151,1436,197,1523]" captionTargetId="figure-17@4.[151,1436,189,1534]" captionTargetPageId="4" captionText="Figures 13–16. Cyana spp.: male genitalia. 13, C. yaseminae, paratype, Uganda, slide BMNH(E) Arct-6196 Karisch (©NHMUK); 14, C. yaseminae, paratype, Uganda, slide 3300 Karisch (photo by T. Karisch); 15, C. smithi: magnified juxta, paratype, Liberia, slide AV4990 Volynkin; 16, C. quentini: magnified juxta, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359832" httpUri="https://zenodo.org/record/13359832/files/figure.png" pageId="1" pageNumber="216">Figs 13, 14</figureCitation>
) by its heavily setose valva apex (that is weakly setose in
<taxonomicName id="ED9E298FFFC1FFF9A0B7F98BD572F9E7" authorityName="Karisch" authorityYear="2003" box="[182,313,1579,1605]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A0B7F98BD572F9E7" box="[182,313,1579,1605]" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
and
<taxonomicName id="ED9E298FFFC1FFF9A170F98BD65BF9E7" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[369,528,1579,1605]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A170F98BD65BF9E7" box="[369,528,1579,1605]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
). In addition, in
<taxonomicName id="ED9E298FFFC1FFF9A2D7F98BD70BF9E4" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[726,832,1579,1606]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A2D7F98BD70BF9E4" box="[726,832,1579,1606]" italics="true" pageId="1" pageNumber="216">C. smithi</emphasis>
</taxonomicName>
the postero-lateral lobes of juxta have significantly finer dentation (
<figureCitation id="B2A54E89FFC1FFF9A155F9ECD590F9C5" box="[340,475,1612,1639]" captionStart="Figures 13–16" captionStartId="4.[151,237,1552,1576]" captionTargetBox="[151,1436,197,1523]" captionTargetId="figure-17@4.[151,1436,189,1534]" captionTargetPageId="4" captionText="Figures 13–16. Cyana spp.: male genitalia. 13, C. yaseminae, paratype, Uganda, slide BMNH(E) Arct-6196 Karisch (©NHMUK); 14, C. yaseminae, paratype, Uganda, slide 3300 Karisch (photo by T. Karisch); 15, C. smithi: magnified juxta, paratype, Liberia, slide AV4990 Volynkin; 16, C. quentini: magnified juxta, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359832" httpUri="https://zenodo.org/record/13359832/files/figure.png" pageId="1" pageNumber="216">Figs 15, 16</figureCitation>
) than in the related species, and the distal saccular process is basally broader and less curved than in
<taxonomicName id="ED9E298FFFC1FFF9A187F9CFD640F92A" authorityName="Karisch" authorityYear="2003" box="[390,523,1647,1673]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A187F9CFD640F92A" box="[390,523,1647,1673]" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
. In comparison with
<taxonomicName id="ED9E298FFFC1FFF9A310F9CFD7F9F92A" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[785,946,1647,1673]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A310F9CFD7F9F92A" box="[785,946,1647,1673]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
, the male genitalia of
<taxonomicName id="ED9E298FFFC1FFF9A4C9F9CFD17EF92B" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[1224,1333,1646,1673]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A4C9F9CFD17EF92B" box="[1224,1333,1646,1673]" italics="true" pageId="1" pageNumber="216">C. smithi</emphasis>
</taxonomicName>
has less sclerotized medial part of juxta, narrower valva with less curved costa, and shorter, slenderer and less curved distal saccular process. The aedeagus of
<taxonomicName id="ED9E298FFFC1FFF9A27AF912D6ADF96E" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[635,742,1713,1740]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A27AF912D6ADF96E" box="[635,742,1713,1740]" italics="true" pageId="1" pageNumber="216">C. smithi</emphasis>
</taxonomicName>
is much longer and narrower (that is short and very broad in
<taxonomicName id="ED9E298FFFC1FFF9A0B6F974D571F94F" authorityName="Karisch" authorityYear="2003" box="[183,314,1748,1774]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A0B6F974D571F94F" box="[183,314,1748,1774]" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
and
<taxonomicName id="ED9E298FFFC1FFF9A177F974D65DF94F" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[374,534,1748,1774]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A177F974D65DF94F" box="[374,534,1748,1774]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
), with a large dorsal process of carina (in
<taxonomicName id="ED9E298FFFC1FFF9A410F974D0DEF94F" authorityName="Karisch" authorityYear="2003" box="[1041,1173,1748,1774]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A410F974D0DEF94F" box="[1041,1173,1748,1774]" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
and
<taxonomicName id="ED9E298FFFC1FFF9A4D0F974D13BF94F" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[1233,1392,1748,1774]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A4D0F974D13BF94F" box="[1233,1392,1748,1774]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
the carina is represented by a sclerotised dentate crest). The vesica of the new species has only one small subbasal diverticulum situated laterally, whereas in
<taxonomicName id="ED9E298FFFC1FFF9A309F8B7D7C6F893" authorityName="Karisch" authorityYear="2003" box="[776,909,1815,1841]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A309F8B7D7C6F893" box="[776,909,1815,1841]" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
there are two subbasal diverticula: a small, short and narrow dorsal one, and a large ventral diverticulum, which is broadly conical with rounded apex and heavily granulated. The female genitalia of
<taxonomicName id="ED9E298FFFC1FFF9A2D7F8FBD70AF8D7" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[726,833,1882,1909]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC1FFF9A2D7F8FBD70AF8D7" box="[726,833,1882,1909]" italics="true" pageId="1" pageNumber="216">C. smithi</emphasis>
</taxonomicName>
(
<figureCitation id="B2A54E89FFC1FFF9A352F8FAD797F8D7" box="[851,988,1882,1909]" captionStart="Figures 17–19" captionStartId="5.[151,237,1689,1713]" captionTargetBox="[151,1436,897,1656]" captionTargetId="figure-283@5.[151,1436,897,1669]" captionTargetPageId="5" captionText="Figures 17–19. Cyana spp.: female genitalia. 17, C. smithi, paratype, Liberia, slide AV4992 Volynkin; 18, C. quentini, Cameroon, slide AV5078 Voynkin; 19, C. smithi: magnified posterior part of corpus bursae, paratype, Liberia, slide AV4992 Volynkin." figureDoi="http://doi.org/10.5281/zenodo.13359834" httpUri="https://zenodo.org/record/13359834/files/figure.png" pageId="1" pageNumber="216">Figs 17, 19</figureCitation>
) is very similar to that of
<taxonomicName id="ED9E298FFFC1FFF9A518F8FBD1D6F8D6" authorityName="Karisch" authorityYear="2003" box="[1305,1437,1883,1909]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC1FFF9A518F8FBD1D6F8D6" box="[1305,1437,1883,1909]" italics="true" pageId="1" pageNumber="216">C. quentini</emphasis>
</taxonomicName>
(
<figureCitation id="B2A54E89FFC1FFF9A0A1F8DCD4BEF835" box="[160,245,1916,1943]" captionStart="Figures 17–19" captionStartId="5.[151,237,1689,1713]" captionTargetBox="[151,1436,897,1656]" captionTargetId="figure-283@5.[151,1436,897,1669]" captionTargetPageId="5" captionText="Figures 17–19. Cyana spp.: female genitalia. 17, C. smithi, paratype, Liberia, slide AV4992 Volynkin; 18, C. quentini, Cameroon, slide AV5078 Voynkin; 19, C. smithi: magnified posterior part of corpus bursae, paratype, Liberia, slide AV4992 Volynkin." figureDoi="http://doi.org/10.5281/zenodo.13359834" httpUri="https://zenodo.org/record/13359834/files/figure.png" pageId="1" pageNumber="216">Fig. 18</figureCitation>
) and differs by its larger signum only. The vesica structure and female genitalia of
<taxonomicName id="ED9E298FFFC1FFF9A4D3F8DDD13AF834" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[1234,1393,1917,1943]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC1FFF9A4D3F8DDD13AF834" box="[1234,1393,1917,1943]" italics="true" pageId="1" pageNumber="216">C. yaseminae</emphasis>
</taxonomicName>
are unknown.
</paragraph>
</subSubSection>
<caption id="7EE10284FFC2FFFAA096F82DD712F7A2" ID-DOI="http://doi.org/10.5281/zenodo.13359826" ID-Zenodo-Dep="13359826" httpUri="https://zenodo.org/record/13359826/files/figure.png" pageId="2" pageNumber="217" startId="2.[151,237,1933,1957]" targetBox="[169,1399,190,1903]" targetPageId="2" targetType="figure">
<paragraph id="2A21520CFFC2FFFAA096F82DD712F7A2" blockId="2.[151,1437,1932,2049]" pageId="2" pageNumber="217">
<emphasis id="18EA8E1EFFC2FFFAA096F82DD56DF807" bold="true" box="[151,294,1933,1957]" pageId="2" pageNumber="217">Figures 1–8.</emphasis>
<taxonomicName id="ED9E298FFFC2FFFAA12FF82DD5E6F806" box="[302,429,1933,1956]" class="Insecta" family="Erebidae" genus="Cyana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="18EA8E1EFFC2FFFAA12FF82DD538F806" box="[302,371,1933,1956]" italics="true" pageId="2" pageNumber="217">Cyana</emphasis>
spp.
</taxonomicName>
: adults. 1,
<taxonomicName id="ED9E298FFFC2FFFAA227F82DD6C3F806" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[550,648,1932,1956]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC2FFFAA227F82DD6C3F806" box="[550,648,1932,1956]" italics="true" pageId="2" pageNumber="217">C. smithi</emphasis>
</taxonomicName>
, holotype male, Liberia (ANHRT); 2,
<taxonomicName id="ED9E298FFFC2FFFAA435F82DD0DCF806" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[1076,1175,1932,1956]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC2FFFAA435F82DD0DCF806" box="[1076,1175,1932,1956]" italics="true" pageId="2" pageNumber="217">C. smithi</emphasis>
</taxonomicName>
, paratype male, Liberia (ANHRT); 3,
<taxonomicName id="ED9E298FFFC2FFFAA136F80CD5D7F861" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[311,412,1963,1987]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC2FFFAA136F80CD5D7F861" box="[311,412,1963,1987]" italics="true" pageId="2" pageNumber="217">C. smithi</emphasis>
</taxonomicName>
, paratype male, Liberia (ANHRT); 4,
<taxonomicName id="ED9E298FFFC2FFFAA352F80CD7F3F861" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[851,952,1963,1987]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC2FFFAA352F80CD7F3F861" box="[851,952,1963,1987]" italics="true" pageId="2" pageNumber="217">C. smithi</emphasis>
</taxonomicName>
, paratype female, Liberia (ANHRT); 5,
<taxonomicName id="ED9E298FFFC2FFFAA585F80CD4A4F840" authorityName="Karisch" authorityYear="2003" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC2FFFAA585F80CD4A4F840" italics="true" pageId="2" pageNumber="217">C. quentini</emphasis>
</taxonomicName>
, male, DR Congo (MWM/ZSM); 6,
<taxonomicName id="ED9E298FFFC2FFFAA27DF86AD6BFF840" authorityName="Karisch" authorityYear="2003" box="[636,756,1994,2018]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC2FFFAA27DF86AD6BFF840" box="[636,756,1994,2018]" italics="true" pageId="2" pageNumber="217">C. quentini</emphasis>
</taxonomicName>
, male, Cameroon (ANHRT); 7,
<taxonomicName id="ED9E298FFFC2FFFAA453F86AD082F840" authorityName="Karisch" authorityYear="2003" box="[1106,1225,1994,2018]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC2FFFAA453F86AD082F840" box="[1106,1225,1994,2018]" italics="true" pageId="2" pageNumber="217">C. quentini</emphasis>
</taxonomicName>
, female, Cameroon (ANHRT); 8,
<taxonomicName id="ED9E298FFFC2FFFAA12FF849D5F5F7A2" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[302,446,2025,2048]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC2FFFAA12FF849D5F5F7A2" box="[302,446,2025,2048]" italics="true" pageId="2" pageNumber="217">C. yaseminae</emphasis>
</taxonomicName>
, paratype male, Uganda (©NHMUK).
</paragraph>
</caption>
<caption id="7EE10284FFC3FFFBA096F83DD658F850" ID-DOI="http://doi.org/10.5281/zenodo.13359828" ID-Zenodo-Dep="13359828" httpUri="https://zenodo.org/record/13359828/files/figure.png" pageId="3" pageNumber="218" startId="3.[151,237,1949,1973]" targetBox="[208,1413,226,1924]" targetPageId="3" targetType="figure">
<paragraph id="2A21520CFFC3FFFBA096F83DD658F850" blockId="3.[151,1437,1949,2034]" pageId="3" pageNumber="218">
<emphasis id="18EA8E1EFFC3FFFBA096F83DD57EF817" bold="true" box="[151,309,1949,1973]" pageId="3" pageNumber="218">Figures 9–12.</emphasis>
<taxonomicName id="ED9E298FFFC3FFFBA140F83DD589F816" box="[321,450,1949,1972]" class="Insecta" family="Erebidae" genus="Cyana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="18EA8E1EFFC3FFFBA140F83DD5CDF816" box="[321,390,1949,1972]" italics="true" pageId="3" pageNumber="218">Cyana</emphasis>
spp.
</taxonomicName>
: male genitalia. 9,
<taxonomicName id="ED9E298FFFC3FFFBA29EF83DD74FF817" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[671,772,1949,1973]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC3FFFBA29EF83DD74FF817" box="[671,772,1949,1973]" italics="true" pageId="3" pageNumber="218">C. smithi</emphasis>
</taxonomicName>
, holotype, Liberia, slide AV4482 Volynkin; 10,
<taxonomicName id="ED9E298FFFC3FFFBA530F83DD1DDF817" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[1329,1430,1949,1973]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC3FFFBA530F83DD1DDF817" box="[1329,1430,1949,1973]" italics="true" pageId="3" pageNumber="218">C. smithi</emphasis>
</taxonomicName>
, paratype, Liberia, slide AV4453 Volynkin; 11,
<taxonomicName id="ED9E298FFFC3FFFBA29DF81CD702F871" authority=", DR" authorityName="DR" box="[668,841,1980,2003]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC3FFFBA29DF81CD75FF871" box="[668,788,1980,2003]" italics="true" pageId="3" pageNumber="218">C. quentini</emphasis>
, DR
</taxonomicName>
Congo, slide MWM 35737 Volynkin; 12,
<taxonomicName id="ED9E298FFFC3FFFBA51CF81CD1DEF871" box="[1309,1429,1980,2003]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC3FFFBA51CF81CD1DEF871" box="[1309,1429,1980,2003]" italics="true" pageId="3" pageNumber="218">C. quentini</emphasis>
</taxonomicName>
, Cameroon, slide AV5077 Voynkin.
</paragraph>
</caption>
<caption id="7EE10284FFC4FFFCA096F9B0D1C0F9C9" ID-DOI="http://doi.org/10.5281/zenodo.13359832" ID-Zenodo-Dep="13359832" httpUri="https://zenodo.org/record/13359832/files/figure.png" pageId="4" pageNumber="219" startId="4.[151,237,1552,1576]" targetBox="[151,1436,197,1523]" targetPageId="4" targetType="figure">
<paragraph id="2A21520CFFC4FFFCA096F9B0D1C0F9C9" blockId="4.[151,1419,1552,1643]" pageId="4" pageNumber="219">
<emphasis id="18EA8E1EFFC4FFFCA096F9B0D576F98A" bold="true" box="[151,317,1552,1576]" pageId="4" pageNumber="219">Figures 13–16.</emphasis>
<taxonomicName id="ED9E298FFFC4FFFCA145F9B0D58BF98A" box="[324,448,1552,1576]" class="Insecta" family="Erebidae" genus="Cyana" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="18EA8E1EFFC4FFFCA145F9B0D5C1F985" box="[324,394,1552,1575]" italics="true" pageId="4" pageNumber="219">Cyana</emphasis>
spp.
</taxonomicName>
: male genitalia. 13,
<taxonomicName id="ED9E298FFFC4FFFCA297F9B0D76DF98A" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[662,806,1552,1576]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC4FFFCA297F9B0D76DF98A" box="[662,806,1552,1576]" italics="true" pageId="4" pageNumber="219">C. yaseminae</emphasis>
</taxonomicName>
, paratype, Uganda, slide BMNH(E) Arct-6196 Karisch (©NHMUK); 14,
<taxonomicName id="ED9E298FFFC4FFFCA159F98FD5A3F9E4" authorityName="Baron &amp; Karisch" authorityYear="2016" box="[344,488,1583,1606]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="yaseminae">
<emphasis id="18EA8E1EFFC4FFFCA159F98FD5A3F9E4" box="[344,488,1583,1606]" italics="true" pageId="4" pageNumber="219">C. yaseminae</emphasis>
</taxonomicName>
, paratype, Uganda, slide 3300 Karisch (photo by T. Karisch); 15,
<taxonomicName id="ED9E298FFFC4FFFCA4AAF98FD140F9E4" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[1195,1291,1582,1606]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC4FFFCA4AAF98FD140F9E4" box="[1195,1291,1582,1606]" italics="true" pageId="4" pageNumber="219">C. smithi</emphasis>
</taxonomicName>
: magnified juxta, paratype, Liberia, slide AV4990 Volynkin; 16,
<taxonomicName id="ED9E298FFFC4FFFCA2D5F9F3D701F9C8" box="[724,842,1619,1642]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC4FFFCA2D5F9F3D701F9C8" box="[724,842,1619,1642]" italics="true" pageId="4" pageNumber="219">C. quentini</emphasis>
</taxonomicName>
: magnified juxta, Cameroon, slide AV5077 Voynkin.
</paragraph>
</caption>
<subSubSection id="62840187FFC4FFFDA096F917D52EFCFD" lastPageId="5" lastPageNumber="220" pageId="4" pageNumber="219" type="description">
<paragraph id="2A21520CFFC4FFFCA096F917D137F87D" blockId="4.[151,1436,1718,2015]" pageId="4" pageNumber="219">
<emphasis id="18EA8E1EFFC4FFFCA096F917D563F973" bold="true" box="[151,296,1719,1745]" pageId="4" pageNumber="219">Description</emphasis>
.
<emphasis id="18EA8E1EFFC4FFFCA139F917D5CBF973" bold="true" box="[312,384,1719,1745]" pageId="4" pageNumber="219">Adult</emphasis>
(
<figureCitation id="B2A54E89FFC4FFFCA18EF916D5B1F973" box="[399,506,1718,1745]" captionStart="Figures 1–8" captionStartId="2.[151,237,1933,1957]" captionTargetBox="[169,1399,190,1903]" captionTargetId="figure-17@2.[157,1431,189,1914]" captionTargetPageId="2" captionText="Figures 1–8. Cyana spp.: adults. 1, C. smithi, holotype male, Liberia (ANHRT); 2, C. smithi, paratype male, Liberia (ANHRT); 3, C. smithi, paratype male, Liberia (ANHRT); 4, C. smithi, paratype female, Liberia (ANHRT); 5, C. quentini, male, DR Congo (MWM/ZSM); 6, C. quentini, male, Cameroon (ANHRT); 7, C. quentini, female, Cameroon (ANHRT); 8, C. yaseminae, paratype male, Uganda (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13359826" httpUri="https://zenodo.org/record/13359826/files/figure.png" pageId="4" pageNumber="219">Figs 1–4</figureCitation>
). Forewing length 7.5–8.0 mm in males (8.0 mm in
<typeStatus id="F525ECAEFFC4FFFCA465F916D087F973" box="[1124,1228,1718,1745]" pageId="4" pageNumber="219" type="holotype">holotype</typeStatus>
) and 8.0–8.5 mm in females. Wingspan
<quantity id="ED66FFE9FFC4FFFCA1B0F978D67EF951" box="[433,565,1752,1779]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="4" pageNumber="219" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">15–16 mm</quantity>
in males and 17.5–
<quantity id="ED66FFE9FFC4FFFCA334F978D7C6F951" box="[821,909,1752,1779]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="219" unit="mm" value="18.0">18 mm</quantity>
in females. Male antennae ciliate, female antennae filiform. The sexual dimorphism well expressed: besides presence of forewing androconial lobes on the underside of forewing, male characterized by its slightly less elongated forewing and black spots of forewing situated obliquely to the costal margin (female has black spots parallel to the costal margin). In males, size of black spots and width of transverse lines vary. Body of both sexes white. Forewing base with a black dot. Ante- and postmedial lines shortly and irregularly dentate, pale pinkish red from cell to inner forewing margin and yellowish at wing costa. Subterminal line pale pinkish red, short, absent from tornus to CuA2 vein, interrupted at veins M3 and CuA1. Cilia white. Hindwing and cilia white, wing pattern absent.
</paragraph>
<paragraph id="2A21520CFFC5FFFDA096FF64D4BEFD92" blockId="5.[151,1436,195,560]" pageId="5" pageNumber="220">
<emphasis id="18EA8E1EFFC5FFFDA096FF64D506FF7C" bold="true" box="[151,333,196,222]" pageId="5" pageNumber="220">Male genitalia</emphasis>
(
<figureCitation id="B2A54E89FFC5FFFDA160FF63D5AAFF7C" box="[353,481,195,222]" captionStart="Figures 9–12" captionStartId="3.[151,237,1949,1973]" captionTargetBox="[208,1413,226,1924]" captionTargetId="figure-8@3.[151,1413,223,1931]" captionTargetPageId="3" captionText="Figures 9–12. Cyana spp.: male genitalia. 9, C. smithi, holotype, Liberia, slide AV4482 Volynkin; 10, C. smithi, paratype, Liberia, slide AV4453 Volynkin; 11, C. quentini, DR Congo, slide MWM 35737 Volynkin; 12, C. quentini, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359828" httpUri="https://zenodo.org/record/13359828/files/figure.png" pageId="5" pageNumber="220">Figs 9, 10</figureCitation>
,
<figureCitation id="B2A54E89FFC5FFFDA1F3FF63D659FF7C" box="[498,530,195,222]" captionStart="Figures 13–16" captionStartId="4.[151,237,1552,1576]" captionTargetBox="[151,1436,197,1523]" captionTargetId="figure-17@4.[151,1436,189,1534]" captionTargetPageId="4" captionText="Figures 13–16. Cyana spp.: male genitalia. 13, C. yaseminae, paratype, Uganda, slide BMNH(E) Arct-6196 Karisch (©NHMUK); 14, C. yaseminae, paratype, Uganda, slide 3300 Karisch (photo by T. Karisch); 15, C. smithi: magnified juxta, paratype, Liberia, slide AV4990 Volynkin; 16, C. quentini: magnified juxta, Cameroon, slide AV5077 Voynkin." figureDoi="http://doi.org/10.5281/zenodo.13359832" httpUri="https://zenodo.org/record/13359832/files/figure.png" pageId="5" pageNumber="220">15</figureCitation>
). Uncus short, narrow, dorso-ventrally flattened, connected to tuba analis. Tuba analis distally broadened, setose. Tegumen short, broadened apically and narrow laterally. Juxta large, shield-like, with two lateral, rounded and inflated lobes densely covered with fine dentation apically. Vinculum short, broad, U-shaped. Valva elongated, slightly dilated medially; costa narrow; distal lobe of valva short, narrow, apically rounded, its distal half densely covered with long and strong setae. Sacculus broad, its distal process elongated trigonal, rather thorn-like, broad at basal with arcuate margins, its distal half slightly curved inwards, apically pointed. Aedeagus short, curved, strongly dilated distally; carina large, heavily sclerotised, more or less trapezoid, directed ventrally. Vesica thick, with a small, globular, granulated subbasal diverticulum situated laterally; a large, broad and globular distal diverticulum with two short but broad granulated subdiverticula apically; and a long and narrow, granulated dorsal diverticulum directed distally.
</paragraph>
<paragraph id="2A21520CFFC5FFFDA096FDF9D52EFCFD" blockId="5.[151,1437,600,863]" pageId="5" pageNumber="220">
<emphasis id="18EA8E1EFFC5FFFDA096FDF9D52CFDD1" bold="true" box="[151,359,601,627]" pageId="5" pageNumber="220">Female genitalia</emphasis>
(
<figureCitation id="B2A54E89FFC5FFFDA179FDF8D648FDD1" box="[376,515,600,627]" captionStart="Figures 17–19" captionStartId="5.[151,237,1689,1713]" captionTargetBox="[151,1436,897,1656]" captionTargetId="figure-283@5.[151,1436,897,1669]" captionTargetPageId="5" captionText="Figures 17–19. Cyana spp.: female genitalia. 17, C. smithi, paratype, Liberia, slide AV4992 Volynkin; 18, C. quentini, Cameroon, slide AV5078 Voynkin; 19, C. smithi: magnified posterior part of corpus bursae, paratype, Liberia, slide AV4992 Volynkin." figureDoi="http://doi.org/10.5281/zenodo.13359834" httpUri="https://zenodo.org/record/13359834/files/figure.png" pageId="5" pageNumber="220">Figs 17, 19</figureCitation>
). Ovipositor broad and short, papillae anales large, rectangular, with rounded corners, setose. Apophyses thin, apophyses posteriores ca. twice longer than apophyses anteriores. Ostium bursae broad. Ductus bursae as broad as ostium bursae, very short, rugose. Corpus bursae sack-like, its anterior third membranous, weakly rugose, with a well-developed rounded signum; medial and posterior thirds of corpus bursae strongly rugose, with weakly sclerotised longitudinal wrinkles. Posterior third of corpus bursae with a broad oblique fold and a short, tuber-like process covered with hair-like setae at the base of appendix bursae. Appendix bursae short, narrowly conical, membranous, situated postero-laterally, directed laterally.
</paragraph>
</subSubSection>
<caption id="7EE10284FFC5FFFDA096F939D52EF94C" ID-DOI="http://doi.org/10.5281/zenodo.13359834" ID-Zenodo-Dep="13359834" httpUri="https://zenodo.org/record/13359834/files/figure.png" pageId="5" pageNumber="220" startId="5.[151,237,1689,1713]" targetBox="[151,1436,897,1656]" targetPageId="5" targetType="figure">
<paragraph id="2A21520CFFC5FFFDA096F939D52EF94C" blockId="5.[151,1437,1688,1774]" pageId="5" pageNumber="220">
<emphasis id="18EA8E1EFFC5FFFDA096F939D575F913" bold="true" box="[151,318,1689,1713]" pageId="5" pageNumber="220">Figures 17–19.</emphasis>
<taxonomicName id="ED9E298FFFC5FFFDA144F939D589F912" box="[325,450,1689,1712]" class="Insecta" family="Erebidae" genus="Cyana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="18EA8E1EFFC5FFFDA144F939D5C0F912" box="[325,395,1689,1712]" italics="true" pageId="5" pageNumber="220">Cyana</emphasis>
spp.
</taxonomicName>
: female genitalia. 17,
<taxonomicName id="ED9E298FFFC5FFFDA2B1F939D75AF912" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[688,785,1688,1712]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC5FFFDA2B1F939D75AF912" box="[688,785,1688,1712]" italics="true" pageId="5" pageNumber="220">C. smithi</emphasis>
</taxonomicName>
, paratype, Liberia, slide AV4992 Volynkin; 18,
<taxonomicName id="ED9E298FFFC5FFFDA51FF939D1DDF912" box="[1310,1430,1689,1712]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="quentini">
<emphasis id="18EA8E1EFFC5FFFDA51FF939D1DDF912" box="[1310,1430,1689,1712]" italics="true" pageId="5" pageNumber="220">C. quentini</emphasis>
</taxonomicName>
, Cameroon, slide AV5078 Voynkin; 19,
<taxonomicName id="ED9E298FFFC5FFFDA252F917D6FDF96D" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[595,694,1719,1743]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC5FFFDA252F917D6FDF96D" box="[595,694,1719,1743]" italics="true" pageId="5" pageNumber="220">C. smithi</emphasis>
</taxonomicName>
: magnified posterior part of corpus bursae, paratype, Liberia, slide AV4992 Volynkin.
</paragraph>
</caption>
<subSubSection id="62840187FFC5FFFDA096F899D0EEF81A" pageId="5" pageNumber="220" type="distribution">
<paragraph id="2A21520CFFC5FFFDA096F899D0ADF8F1" blockId="5.[151,1254,1848,1875]" box="[151,1254,1848,1875]" pageId="5" pageNumber="220">
<heading id="7169E560FFC5FFFDA096F899D0ADF8F1" box="[151,1254,1848,1875]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="220" reason="2">
<emphasis id="18EA8E1EFFC5FFFDA096F899D57AF8F1" bold="true" box="[151,305,1849,1875]" pageId="5" pageNumber="220">Distribution</emphasis>
.
<taxonomicName id="ED9E298FFFC5FFFDA13EF899D590F8F1" ID-CoL="BFNVS" authorityName="Volynkin &amp; Laszlo" authorityYear="2019" box="[319,475,1848,1875]" class="Insecta" family="Erebidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="220" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="18EA8E1EFFC5FFFDA13EF899D590F8F1" box="[319,475,1848,1875]" italics="true" pageId="5" pageNumber="220">Cyana smithi</emphasis>
</taxonomicName>
is currently only known from northeastern
<collectingCountry id="5289129CFFC5FFFDA3DBF898D064F8F1" box="[986,1071,1848,1875]" name="Liberia" pageId="5" pageNumber="220">Liberia</collectingCountry>
(
<collectingRegion id="E85A9CEEFFC5FFFDA43CF898D091F8F1" box="[1085,1242,1848,1875]" country="Liberia" name="Lofa" pageId="5" pageNumber="220">Lofa County</collectingRegion>
).
</heading>
</paragraph>
<paragraph id="2A21520CFFC5FFFDA096F8DCD0EEF81A" blockId="5.[151,1435,1916,1976]" pageId="5" pageNumber="220">
<emphasis id="18EA8E1EFFC5FFFDA096F8DCD554F834" bold="true" box="[151,287,1916,1942]" pageId="5" pageNumber="220">Etymology</emphasis>
. The species is dedicated to Mr Richard Smith, founder of the African Natural History Research Trust, organiser of extensive entomological exploratory program in Sub-Saharan Africa.
</paragraph>
</subSubSection>
</treatment>
</document>
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