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<document id="EA24262A7B3D2012DCC1A8905E7E9C75" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki &amp; Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB72EFF9569F0F97946F75CE3" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Nematodes apicalis Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="3" lastPageNumber="33" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="32" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="865AE1EE17F62DB5D6D1294CEF3B5601" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EC310CE9ED9756755C61925D1C0B3A7E">
<mods:title id="1F969A151DC2A3AF4F500A183FCC705C">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
</mods:titleInfo>
<mods:name id="305086D6E9276395A197C09A4FC50395" type="personal">
<mods:role id="2447D8E445F345F457DFEECA4A267BC0">
<mods:roleTerm id="583144DF17867CD0C6E8A8284226DDDA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15A236A0CCBA29EA87E47F10A1389A80">Otto, Robert L.</mods:namePart>
<mods:affiliation id="499E4B89D045AB8D50038659FEE2869B">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
</mods:name>
<mods:name id="BB2C4E6364DF8C434FDBEE8AC31F01A8" type="personal">
<mods:role id="DDA06B973D53F239FF6757F0A17EDA2C">
<mods:roleTerm id="5E7BA3C05E956C8443C0F665DD4455FB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB2879ECA083E590ABD5CE92314C0153">Muona, Jyrki</mods:namePart>
<mods:affiliation id="BEFD7A176FF6535919C0A7E728718D0C">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
</mods:name>
<mods:name id="26317B994B3A77468E81C9B0ADC21146" type="personal">
<mods:role id="C6C472A7D021A0C4F6303CEE9DB97F66">
<mods:roleTerm id="4C753B25193CEEF9C30717E02C42EB41">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4956064E95F9FF21DDC14B68054E98CB">Córdoba-Alfaro, Jim</mods:namePart>
<mods:affiliation id="D897FC0B7FA44E80F22A85AF283868AC">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
</mods:name>
<mods:typeOfResource id="55832FF88B91D7F466FF00BDA0385385">text</mods:typeOfResource>
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<mods:titleInfo id="E336DCCCE730E79904134A29EDF06BCB">
<mods:title id="2679DCC31767E082D8BCC72D0E20554D">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="E658F7089B014CBB7DC50F104EDDF83E">
<mods:date id="AA1CBA60E62CA2F52EA90F62D3F6D1B2">2023</mods:date>
<mods:detail id="0FAB208F2C69B0A978C7E24384BAD472" type="pubDate">
<mods:number id="3B631F4B430C750D4D036C0B6014CC3F">2023-05-26</mods:number>
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<mods:detail id="C4AE722F43F665781E3FCF3951DD0A93" type="volume">
<mods:number id="8F8B4EF8E903B6930DF8F282480213CF">2023</mods:number>
</mods:detail>
<mods:detail id="1785D49966FF96FD4099283BB85F291D" type="issue">
<mods:number id="0CB8CCEDE50E812853540175426C1879">991</mods:number>
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<mods:start id="06CAAFA98E1AA7E2188BF9E273FA9BCB">1</mods:start>
<mods:end id="960AAFBF79D731DFDEC10D65BAFD0D9F">36</mods:end>
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<mods:classification id="0AA6B9BD991B0A34779D6D8AF69E158C">journal article</mods:classification>
<mods:identifier id="7098F5DD2599F7FE18D291306521B68B" type="DOI">10.5281/zenodo.10831085</mods:identifier>
<mods:identifier id="E1142E1F9A43B72DD9B207CEFC371A1B" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="46A79D6B7B682A140BC7E35432467DBA" type="Zenodo-Dep">10831085</mods:identifier>
<mods:identifier id="321394A4845C9E0BCC7AB60C2DA24F5D" type="ZooBank">C1D5B819-A964-4679-B090-84CDBBC59D6A</mods:identifier>
</mods:mods>
<treatment id="954287CDB72EFF9569F0F97946F75CE3" LSID="urn:lsid:plazi:treatment:954287CDB72EFF9569F0F97946F75CE3" httpUri="http://treatment.plazi.org/id/954287CDB72EFF9569F0F97946F75CE3" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32">
<subSubSection id="55F16550B72EFF9669F0F97947655C81" box="[192,1036,1711,1740]" pageId="33" pageNumber="32" type="nomenclature">
<paragraph id="1D5436DBB72EFF9669F0F97947655C81" blockId="33.[192,1036,1711,1774]" box="[192,1036,1711,1740]" pageId="33" pageNumber="32">
<heading id="461C81B7B72EFF9669F0F97947655C81" box="[192,1036,1711,1740]" fontSize="12" level="2" pageId="33" pageNumber="32" reason="2">
<emphasis id="2F9FEAC9B72EFF9669F0F97947655C81" bold="true" box="[192,1036,1711,1740]" pageId="33" pageNumber="32">
<taxonomicName id="DAEB4D58B72EFF9669F0F97940005C81" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,873,1711,1740]" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="apicalis" status="sp. nov.">
<emphasis id="2F9FEAC9B72EFF9669F0F97942DA5C81" bold="true" box="[192,435,1711,1740]" italics="true" pageId="33" pageNumber="32">Nematodes apicalis</emphasis>
Otto, Muona and Córdoba-Alfaro
</taxonomicName>
,
<taxonomicNameLabel id="34AC57B2B72EFF966A47F96647655C81" box="[887,1036,1711,1740]" pageId="33" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B72EFF9669F0F90242375CA3" box="[192,350,1748,1774]" pageId="33" pageNumber="32" type="description">
<paragraph id="1D5436DBB72EFF9669F0F90242375CA3" blockId="33.[192,1036,1711,1774]" box="[192,350,1748,1774]" pageId="33" pageNumber="32">
<figureCitation id="85D02A5EB72EFF9669F0F90242375CA3" box="[192,350,1748,1774]" captionStart="Figures 71–74" captionStartId="34.[192,279,732,758]" captionTargetBox="[321,1311,213,703]" captionTargetId="figure-461@34.[321,1311,213,703]" captionTargetPageId="34" captionText="Figures 71–74. Costa Rican Nematodes species. 71) Nematodes mannerheimi, dorsal habitus. 72) Nematodes mexicanus, dorsal habitus.73) Nematodes apicalis sp. nov., dorsal habitus. 74) Nematodes apicalis sp. nov., ventral habitus. (Scale: 72–74 = 1.0 mm; 71 = 5.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831118" httpUri="https://zenodo.org/record/10831118/files/figure.png" pageId="33" pageNumber="32">Figures 73–74</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B72EFF9669F0F92B40595D77" pageId="33" pageNumber="32" type="diagnosis">
<paragraph id="1D5436DBB72EFF9669F0F92B40595D77" blockId="33.[192,1440,1789,1850]" pageId="33" pageNumber="32">
<emphasis id="2F9FEAC9B72EFF9669F0F92B42555D5A" bold="true" box="[192,316,1789,1815]" pageId="33" pageNumber="32">Diagnosis.</emphasis>
The new eucnemid species is distinguished from all known Central American
<taxonomicName id="DAEB4D58B72EFF966D81F92846165D55" box="[1201,1407,1790,1816]" pageId="33" pageNumber="32">
<emphasis id="2F9FEAC9B72EFF966D81F92846435D55" box="[1201,1322,1790,1816]" italics="true" pageId="33" pageNumber="32">Nematodes</emphasis>
species
</taxonomicName>
by the presence of silver-colored setae on the elytral apices.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B72EFF9569F0F89C46F959C4" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32" type="materials_examined">
<paragraph id="1D5436DBB72EFF9669F0F89C46C95DCA" blockId="33.[192,1441,1866,1927]" pageId="33" pageNumber="32">
<emphasis id="2F9FEAC9B72EFF9669F0F89C41585D29" bold="true" box="[192,561,1866,1892]" pageId="33" pageNumber="32">
Type material. Female
<typeStatus id="C2508879B72EFF9668F9F89C41585D29" box="[457,561,1866,1892]" pageId="33" pageNumber="32" type="holotype">holotype</typeStatus>
</emphasis>
: “
<collectingCountry id="65FC764BB72EFF966B76F89C41815D29" box="[582,744,1866,1892]" name="Costa Rica" pageId="33" pageNumber="32">COSTA RICA</collectingCountry>
:
<collectingRegion id="DF2FF839B72EFF966BC1F89C40275D29" box="[753,846,1866,1892]" country="Costa Rica" name="Heredia" pageId="33" pageNumber="32">Heredia</collectingRegion>
; Est.Bio. La Selva, 50–;
<quantity id="DA139B3EB72EFF966D60F89C47FC5D29" box="[1104,1173,1866,1893]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="32" unit="m" value="150.0">150 m</quantity>
<geoCoordinate id="78DF501CB72EFF966DABF89C479F5D29" box="[1179,1270,1866,1892]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="33" pageNumber="32" precision="925" value="10.433333">10°26′N</geoCoordinate>
<geoCoordinate id="78DF501CB72EFF966DCBF89C46365D29" box="[1275,1375,1866,1892]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="33" pageNumber="32" precision="925" value="-84.01667">84°01′W</geoCoordinate>
; Proy.
<collectionCode id="7BFAAE1EB72EFF9669F0F8BA426F5DCB" box="[192,262,1900,1926]" pageId="33" pageNumber="32">ALAS</collectionCode>
, INBio-OET” / “M/07/337;
<date id="6955101BB72EFF966B74F8BA41945DCB" box="[580,765,1900,1926]" pageId="33" pageNumber="32" value="1994-02-01">01 Febrero 1994</date>
; Bosque primario” / “
<emphasis id="2F9FEAC9B72EFF966AC6F8BA47FD5DCB" bold="true" box="[1014,1172,1900,1926]" pageId="33" pageNumber="32">
<typeStatus id="C2508879B72EFF966AC6F8BA47E75DCB" box="[1014,1166,1900,1926]" pageId="33" pageNumber="32" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
;
<taxonomicName id="DAEB4D58B72EFF966D92F8BA46755DCB" authorityName="Berthold" authorityYear="1827" box="[1186,1308,1900,1926]" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="2F9FEAC9B72EFF966D92F8BA46755DCB" box="[1186,1308,1900,1926]" italics="true" pageId="33" pageNumber="32">Nematodes</emphasis>
</taxonomicName>
;
<taxonomicName id="DAEB4D58B72EFF966C1AF8BA46145DCB" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[1322,1405,1900,1926]" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="apicalis">
<emphasis id="2F9FEAC9B72EFF966C1AF8BA46145DCB" box="[1322,1405,1900,1926]" italics="true" pageId="33" pageNumber="32">apicalis</emphasis>
</taxonomicName>
<specimenCount id="0BEDFD52B72EFF966CB4F8A646F05DCB" box="[1412,1433,1904,1926]" pageId="33" pageNumber="32" type="female">♀</specimenCount>
;
</paragraph>
<caption id="49946653B72DFF9569F0FD0A41AE5977" ID-DOI="http://doi.org/10.5281/zenodo.10831118" ID-Zenodo-Dep="10831118" httpUri="https://zenodo.org/record/10831118/files/figure.png" pageId="34" pageNumber="33" startId="34.[192,279,732,758]" targetBox="[321,1311,213,703]" targetPageId="34" targetType="figure">
<paragraph id="1D5436DBB72DFF9569F0FD0A41AE5977" blockId="34.[191,1440,732,827]" pageId="34" pageNumber="33">
<emphasis id="2F9FEAC9B72DFF9569F0FD0A420758BB" bold="true" box="[192,366,732,758]" pageId="34" pageNumber="33">Figures 71–74.</emphasis>
Costa Rican
<taxonomicName id="DAEB4D58B72DFF956B3DFD0B418F58BA" box="[525,742,733,759]" class="Insecta" family="Eucnemidae" genus="Nematodes" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2F9FEAC9B72DFF956B3DFD0B41EF58BA" box="[525,646,733,759]" italics="true" pageId="34" pageNumber="33">Nematodes</emphasis>
species.
</taxonomicName>
<emphasis id="2F9FEAC9B72DFF956BDFFD0A407C58BB" bold="true" box="[751,789,732,758]" pageId="34" pageNumber="33">71)</emphasis>
<taxonomicName id="DAEB4D58B72DFF956A2FFD0B475C58BA" box="[799,1077,733,759]" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="mannerheimi">
<emphasis id="2F9FEAC9B72DFF956A2FFD0B475C58BA" box="[799,1077,733,759]" italics="true" pageId="34" pageNumber="33">Nematodes mannerheimi</emphasis>
</taxonomicName>
, dorsal habitus.
<emphasis id="2F9FEAC9B72DFF956DC7FD0A467458BB" bold="true" box="[1271,1309,732,758]" pageId="34" pageNumber="33">72)</emphasis>
<taxonomicName id="DAEB4D58B72DFF956C17FD0B425D5955" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="2F9FEAC9B72DFF956C17FD0B425D5955" italics="true" pageId="34" pageNumber="33">Nematodes mexicanus</emphasis>
</taxonomicName>
, dorsal habitus.
<emphasis id="2F9FEAC9B72DFF9568D6FD2841655955" bold="true" box="[486,524,766,792]" pageId="34" pageNumber="33">73)</emphasis>
<taxonomicName id="DAEB4D58B72DFF956B22FD28418B5955" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[530,738,766,792]" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="apicalis" status="sp. nov.">
<emphasis id="2F9FEAC9B72DFF956B22FD28418B5955" box="[530,738,766,792]" italics="true" pageId="34" pageNumber="33">Nematodes apicalis</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B72DFF956BD7FD2840545955" bold="true" box="[743,829,766,792]" pageId="34" pageNumber="33">
<taxonomicNameLabel id="34AC57B2B72DFF956BD7FD2840545955" box="[743,829,766,792]" pageId="34" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal habitus.
<emphasis id="2F9FEAC9B72DFF956ADFFD28477C5955" bold="true" box="[1007,1045,766,792]" pageId="34" pageNumber="33">74)</emphasis>
<taxonomicName id="DAEB4D58B72DFF956D2BFD2847825955" authority="Otto &amp; Muona &amp; Córdoba-Alfaro, 2023" authorityName="Otto &amp; Muona &amp; Córdoba-Alfaro" authorityYear="2023" box="[1051,1259,766,792]" class="Insecta" family="Eucnemidae" genus="Nematodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="apicalis" status="sp. nov.">
<emphasis id="2F9FEAC9B72DFF956D2BFD2847825955" box="[1051,1259,766,792]" italics="true" pageId="34" pageNumber="33">Nematodes apicalis</emphasis>
</taxonomicName>
<emphasis id="2F9FEAC9B72DFF956DC1FD28462E5955" bold="true" box="[1265,1351,766,792]" pageId="34" pageNumber="33">
<taxonomicNameLabel id="34AC57B2B72DFF956DC1FD28462E5955" box="[1265,1351,766,792]" pageId="34" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ventral habitus. (Scale: 72–74 = 1.0 mm; 71 = 5.0 mm)
</paragraph>
</caption>
<paragraph id="1D5436DBB72DFF9569F0FCB946F959C4" blockId="34.[192,1424,879,905]" box="[192,1424,879,905]" pageId="34" pageNumber="33">
<materialsCitation id="AD833C86B72DFF9569F0FCB946F959C4" box="[192,1424,879,905]" collectionCode="R, INBC" collectorName="Otto &amp; Muona &amp; Cordoba-Alfaro &amp; Det. &amp; L. Otto" pageId="34" pageNumber="33" specimenCount="1" typeStatus="holotype">
<collectorName id="B01E530DB72DFF9569F0FCB9439F59C4" box="[192,246,879,905]" pageId="34" pageNumber="33">Otto</collectorName>
,
<collectorName id="B01E530DB72DFF956831FCB9423B59C4" box="[257,338,879,905]" pageId="34" pageNumber="33">Muona</collectorName>
&amp;
<collectorName id="B01E530DB72DFF956843FCB9414259C4" box="[371,555,879,905]" pageId="34" pageNumber="33">Córdoba-Alfaro</collectorName>
;
<collectorName id="B01E530DB72DFF956B06FCB9410F59C4" box="[566,614,879,905]" pageId="34" pageNumber="33">Det.</collectorName>
<collectionCode id="7BFAAE1EB72DFF956B5CFCB941E959C4" box="[620,640,879,905]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="34" pageNumber="33">R</collectionCode>
.
<collectorName id="B01E530DB72DFF956BB4FCB941BE59C4" box="[644,727,879,905]" pageId="34" pageNumber="33">L. Otto</collectorName>
; 2019” (red printed label).
<typeStatus id="C2508879B72DFF956D36FCB9470659C4" box="[1030,1135,879,905]" pageId="34" pageNumber="33" type="holotype">Holotype</typeStatus>
to be deposited in
<collectionCode id="7BFAAE1EB72DFF956C76FCB946E559C4" box="[1350,1420,879,905]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="34" pageNumber="33">INBC</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="55F16550B72DFF9569F0FC4E42335C42" pageId="34" pageNumber="33" type="description">
<paragraph id="1D5436DBB72DFF9569F0FC4E42335C42" blockId="34.[191,1442,920,1551]" pageId="34" pageNumber="33">
<emphasis id="2F9FEAC9B72DFF9569F0FC4E414959FE" bold="true" box="[192,544,920,947]" pageId="34" pageNumber="33">
Description. Female
<typeStatus id="C2508879B72DFF956888FC4F414959FE" box="[440,544,921,947]" pageId="34" pageNumber="33" type="holotype">holotype</typeStatus>
</emphasis>
: Length, 6.0 mm. Width,
<quantity id="DA139B3EB72DFF956A7BFC4E40DA59FE" box="[843,947,920,947]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="34" pageNumber="33" unit="mm" value="1.25">1.25 mm</quantity>
. Body elongate; dark black; antennae black with apices of antennomere XI infuscate reddish; femur and tibiae black, tarsi dark brownish-black; head, pronotum and elytral humeri and elytral apices clothed with short, recumbent silver-colored setae (
<figureCitation id="85D02A5EB72DFF956C10FC0A46FD59BB" box="[1312,1428,988,1014]" captionStart="Figures 71–74" captionStartId="34.[192,279,732,758]" captionTargetBox="[321,1311,213,703]" captionTargetId="figure-461@34.[321,1311,213,703]" captionTargetPageId="34" captionText="Figures 71–74. Costa Rican Nematodes species. 71) Nematodes mannerheimi, dorsal habitus. 72) Nematodes mexicanus, dorsal habitus.73) Nematodes apicalis sp. nov., dorsal habitus. 74) Nematodes apicalis sp. nov., ventral habitus. (Scale: 72–74 = 1.0 mm; 71 = 5.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831118" httpUri="https://zenodo.org/record/10831118/files/figure.png" pageId="34" pageNumber="33">Figure 73</figureCitation>
).
<emphasis id="2F9FEAC9B72DFF9569F0FC2B43965E5A" bold="true" box="[192,255,1021,1047]" pageId="34" pageNumber="33">Head</emphasis>
: Surface densely punctate, somewhat dullish, subspherical; frons convex, with slightly depressed median fovea above frontoclypeal region; apical margin of frontoclypeal region rounded, about 2 times wider than base; interantennal carina absent; mandibles stout, bidentate, densely punctate.
<emphasis id="2F9FEAC9B72DFF956D64FB9647D15E17" bold="true" box="[1108,1208,1088,1114]" pageId="34" pageNumber="33">Antenna</emphasis>
: Filiform from flagellomeres I–VIII, attaining about 1/3 the length of the body, never beyond pronotal hind angles; flagellomere I longer than II; flagellomeres II–III sub-equal, quadrate; flagellomere IV larger than III, slightly shorter than V, quadrate; flagellomeres V–VIII each larger than IV, sub-equal, quadrate; flagellomere IX simple, slightly longer than VIII.
<emphasis id="2F9FEAC9B72DFF956809FB1142DB5EAC" bold="true" box="[313,434,1223,1249]" pageId="34" pageNumber="33">Pronotum</emphasis>
: Surface somewhat dullish, densely punctate to rugose; longer than wide, with moderate, sharp hind angles; lateral sides sinuous basally, apically arcuate; disc convex with pair of very shallow circular fovea; base sinuous.
<emphasis id="2F9FEAC9B72DFF956896FADC41325F69" bold="true" box="[422,603,1290,1316]" pageId="34" pageNumber="33">Scutellar shield</emphasis>
: Elongate, sub-triangular, setose, shallowly punctate and distally rounded.
<emphasis id="2F9FEAC9B72DFF9569F0FAFA42615F0B" bold="true" box="[192,264,1324,1350]" pageId="34" pageNumber="33">Elytra</emphasis>
: Striae present on humeri and elytral apices, indistinct elsewhere; interstices flattened; surfaces shiny, transversely rugose at basal 1/4, dense, crowded punctures present elsewhere.
<emphasis id="2F9FEAC9B72DFF956D28FA9B47245F2A" bold="true" box="[1048,1101,1357,1383]" pageId="34" pageNumber="33">Legs</emphasis>
: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I–III simple; metatarsomere IV short, excavate-emarginate; metatarsomere V elongate with simple claws.
<emphasis id="2F9FEAC9B72DFF95698FFA6442645F81" bold="true" box="[191,269,1458,1484]" pageId="34" pageNumber="33">Venter</emphasis>
(
<figureCitation id="85D02A5EB72DFF95682DFA6442E75F81" box="[285,398,1458,1484]" captionStart="Figures 71–74" captionStartId="34.[192,279,732,758]" captionTargetBox="[321,1311,213,703]" captionTargetId="figure-461@34.[321,1311,213,703]" captionTargetPageId="34" captionText="Figures 71–74. Costa Rican Nematodes species. 71) Nematodes mannerheimi, dorsal habitus. 72) Nematodes mexicanus, dorsal habitus.73) Nematodes apicalis sp. nov., dorsal habitus. 74) Nematodes apicalis sp. nov., ventral habitus. (Scale: 72–74 = 1.0 mm; 71 = 5.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831118" httpUri="https://zenodo.org/record/10831118/files/figure.png" pageId="34" pageNumber="33">Figure 74</figureCitation>
): Closely punctate, with short, recumbent silver-colored setae; hypomeron without antennal grooves; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially 3.0–6.0 times wider than laterally.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B72DFF9569F0F9C942545C17" pageId="34" pageNumber="33" type="distribution">
<paragraph id="1D5436DBB72DFF9569F0F9C942545C17" blockId="34.[192,1440,1567,1626]" pageId="34" pageNumber="33">
<emphasis id="2F9FEAC9B72DFF9569F0F9C942325C74" bold="true" box="[192,347,1567,1593]" pageId="34" pageNumber="33">Distribution.</emphasis>
This eucnemid species is known from a
<typeStatus id="C2508879B72DFF956A2EF9C940E95C74" box="[798,896,1567,1593]" pageId="34" pageNumber="33" type="holotype">holotype</typeStatus>
taken from a single location within the province of
<collectingRegion id="DF2FF839B72DFF9569EDF99642505C17" box="[221,313,1600,1626]" country="Costa Rica" name="Heredia" pageId="34" pageNumber="33">Heredia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B72DFF9569F0F9BC47F05CC9" box="[192,1177,1642,1669]" pageId="34" pageNumber="33" type="biology_ecology">
<paragraph id="1D5436DBB72DFF9569F0F9BC47F05CC9" blockId="34.[192,1177,1642,1669]" box="[192,1177,1642,1669]" pageId="34" pageNumber="33">
<emphasis id="2F9FEAC9B72DFF9569F0F9BC42495CC9" bold="true" box="[192,288,1642,1668]" pageId="34" pageNumber="33">Biology.</emphasis>
The
<typeStatus id="C2508879B72DFF956866F9BC42D15CC9" box="[342,440,1642,1668]" pageId="34" pageNumber="33" type="holotype">holotype</typeStatus>
was taken from a primary forest. Larvae and pupae are unknown.
</paragraph>
</subSubSection>
<subSubSection id="55F16550B72DFF9569F0F94246F75CE3" box="[192,1438,1684,1710]" pageId="34" pageNumber="33" type="etymology">
<paragraph id="1D5436DBB72DFF9569F0F94246F75CE3" blockId="34.[192,1438,1684,1710]" box="[192,1438,1684,1710]" pageId="34" pageNumber="33">
<emphasis id="2F9FEAC9B72DFF9569F0F942422D5CE3" bold="true" box="[192,324,1684,1710]" pageId="34" pageNumber="33">Etymology.</emphasis>
The specific epithet is derived from the presence of silver colored setae present on the elytral apices.
</paragraph>
</subSubSection>
</treatment>
</document>
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