262 lines
31 KiB
XML
262 lines
31 KiB
XML
<document id="2A2F79A70CF8C75261005B15CC8B28D1" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki & Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB707FFBE69F2FC3C460A5F0D" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Adelothyreus brevis Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="7" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="9040B6AABC649A82C169E027377977EC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B003673723A3B39C9F3F32BBC13BA6D7">
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<mods:title id="35B3BFDFFEEF1DCBF1F424472EEAB69B">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
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</mods:titleInfo>
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<mods:name id="3DEA7B253F482F97B35EEB7CD24DFDAD" type="personal">
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<mods:role id="8C0C5B7DD2F431D905A8092C96C3ED7E">
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<mods:roleTerm id="1B32D1C7E4744B43C100D3CE92526992">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7C0E217B057AF669E7F07715F6555737">Otto, Robert L.</mods:namePart>
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<mods:affiliation id="7D40D8102DA7FBD1A4F5CBBBC1A51F11">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:name id="17C7D46BEB0AC2F8742F06F8306306F5" type="personal">
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<mods:role id="5D8C2A5F20AC8E52E42B043A87C6DFCD">
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<mods:roleTerm id="3427E58635A4DFB9E926D72586888089">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FA610E05A57DD3C1165666E774DBEC42">Muona, Jyrki</mods:namePart>
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<mods:affiliation id="D31B58F278DEBBF1310F0F5F8907CF1D">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
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</mods:name>
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<mods:name id="34919D834BD5A0E079D0A7B8180D32A1" type="personal">
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<mods:role id="87287F342D219E99B78BD964D3FCE773">
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<mods:roleTerm id="1338BC7CD0623EE889C465A12BCC27C0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="398E38DE43100E59C647D2F0BAF062C1">Córdoba-Alfaro, Jim</mods:namePart>
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<mods:affiliation id="F6812855929AAA11AF9AFF2C98F4F71A">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="FBEFB96C980E6DD8A16D3E2C43D2DF16">text</mods:typeOfResource>
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<mods:titleInfo id="CE76FE54DCEC895EDCD6759912559263">
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<mods:title id="8A9331D49B47977779214C90463C8D35">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="83856F50BFB412CCA300686C24B0DD06">
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<mods:date id="880A16B9B0110953055A702FF10FF34B">2023</mods:date>
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<mods:detail id="29E95CD869A748E4E6CB245A82C4E8F8" type="pubDate">
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<mods:number id="6222912FAF529564AA39B3DD413D83D5">2023-05-26</mods:number>
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</mods:detail>
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<mods:detail id="A68D51CCA24A46ECB8F4B1DB40A4A286" type="volume">
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<mods:number id="134843ACDA617293EE00E170DE3B815E">2023</mods:number>
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</mods:detail>
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<mods:detail id="66B36D5D28AAB99E3A5AC7E5758497A1" type="issue">
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<mods:number id="F2E0810ACDD4BC48A249D8F5098B81C3">991</mods:number>
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</mods:detail>
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<mods:extent id="53B53A1C13B7F66EE4051C1708068AC4" unit="page">
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<mods:start id="AD257213E7D37DB810565CEE9E1BEBC4">1</mods:start>
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<mods:end id="7143B74B4ACBB2271DED545370F6C667">36</mods:end>
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<mods:classification id="320BFFCA2711730C0897BA91EE9233A3">journal article</mods:classification>
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<mods:identifier id="CF2B89566752D3DA5D8DDED7D55A7E10" type="DOI">10.5281/zenodo.10831085</mods:identifier>
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<mods:identifier id="6427175FF3619D592E8132F10377C3DC" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="B7A086899095211F453335C318F9C9F0" type="Zenodo-Dep">10831085</mods:identifier>
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<mods:identifier id="909B1CDE80F8066A199E4A24FA40BEEF" type="ZooBank">C1D5B819-A964-4679-B090-84CDBBC59D6A</mods:identifier>
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</mods:mods>
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<treatment id="954287CDB707FFBE69F2FC3C460A5F0D" LSID="urn:lsid:plazi:treatment:954287CDB707FFBE69F2FC3C460A5F0D" httpUri="http://treatment.plazi.org/id/954287CDB707FFBE69F2FC3C460A5F0D" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
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<subSubSection id="55F16550B707FFBF69F2FC3C477D5E4A" box="[194,1044,1001,1031]" pageId="8" pageNumber="7" type="nomenclature">
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<paragraph id="1D5436DBB707FFBF69F2FC3C477D5E4A" blockId="8.[194,1044,1001,1064]" box="[194,1044,1001,1031]" pageId="8" pageNumber="7">
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<heading id="461C81B7B707FFBF69F2FC3C477D5E4A" box="[194,1044,1001,1031]" fontSize="12" level="2" pageId="8" pageNumber="7" reason="2">
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||
<emphasis id="2F9FEAC9B707FFBF69F2FC3C477D5E4A" bold="true" box="[194,1044,1001,1031]" pageId="8" pageNumber="7">
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<taxonomicName id="DAEB4D58B707FFBF69F2FC3C40185E4B" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[194,881,1001,1031]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
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<emphasis id="2F9FEAC9B707FFBF69F2FC3C42D35E4A" bold="true" box="[194,442,1002,1031]" italics="true" pageId="8" pageNumber="7">Adelothyreus brevis</emphasis>
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Otto, Muona and Córdoba-Alfaro
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</taxonomicName>
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,
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<taxonomicNameLabel id="34AC57B2B707FFBF6A4FFC3C477D5E4A" box="[895,1044,1002,1031]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="55F16550B707FFBF69F2FBD842375E65" box="[194,350,1038,1064]" pageId="8" pageNumber="7" type="description">
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||
<paragraph id="1D5436DBB707FFBF69F2FBD842375E65" blockId="8.[194,1044,1001,1064]" box="[194,350,1038,1064]" pageId="8" pageNumber="7">
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||
<figureCitation id="85D02A5EB707FFBF69F2FBD842375E65" box="[194,350,1038,1064]" captionStart="Figures 10–16" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 10–16. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 10–16 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="8" pageNumber="7">Figures 10–12</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="55F16550B707FFBF69F2FBE1419D5E3E" pageId="8" pageNumber="7" type="diagnosis">
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<paragraph id="1D5436DBB707FFBF69F2FBE1419D5E3E" blockId="8.[194,1442,1079,1139]" pageId="8" pageNumber="7">
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<emphasis id="2F9FEAC9B707FFBF69F2FBE142575E1C" bold="true" box="[194,318,1079,1105]" pageId="8" pageNumber="7">Diagnosis.</emphasis>
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The presence of a very short anterior lateral pronotal ridge will distinguish the new species from all known
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<taxonomicName id="DAEB4D58B707FFBF6826FB8F42905E3E" box="[278,505,1113,1139]" pageId="8" pageNumber="7">
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<emphasis id="2F9FEAC9B707FFBF6826FB8F42CD5E3E" box="[278,420,1113,1139]" italics="true" pageId="8" pageNumber="7">Adelothyreus</emphasis>
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species
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</taxonomicName>
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present in
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<collectingCountry id="65FC764BB707FFBF6B45FB8F41995E3E" box="[629,752,1113,1139]" name="Costa Rica" pageId="8" pageNumber="7">Costa Rica</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="55F16550B707FFBF69F2FB55462B5EAD" pageId="8" pageNumber="7" type="materials_examined">
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<paragraph id="1D5436DBB707FFBF69F2FB55462B5EAD" blockId="8.[193,1443,1154,1248]" pageId="8" pageNumber="7">
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<emphasis id="2F9FEAC9B707FFBF69F2FB5542C45ED0" bold="true" box="[194,429,1154,1181]" pageId="8" pageNumber="7">Type material. Male</emphasis>
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<materialsCitation id="AD833C86B707FFBF6885FB55462B5EAD" collectingDate="1993-05-02" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Est. Bio. La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
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<emphasis id="2F9FEAC9B707FFBF6885FB5541755ED0" bold="true" box="[437,540,1155,1181]" pageId="8" pageNumber="7">
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<typeStatus id="C2508879B707FFBF6885FB5541755ED0" box="[437,540,1155,1181]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
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</emphasis>
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: “
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<collectingCountry id="65FC764BB707FFBF6B04FB5541B15ED0" box="[564,728,1155,1181]" name="Costa Rica" pageId="8" pageNumber="7">COSTA RICA</collectingCountry>
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:
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<collectingRegion id="DF2FF839B707FFBF6BD3FB5440565ED1" box="[739,831,1154,1180]" country="Costa Rica" name="Heredia" pageId="8" pageNumber="7">Heredia</collectingRegion>
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;
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||
<location id="18346000B707FFBF6A7AFB54476E5ED1" LSID="urn:lsid:plazi:treatment:954287CDB707FFBE69F2FC3C460A5F0D:18346000B707FFBF6A7AFB54476E5ED1" box="[842,1031,1154,1181]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Est. Bio. La Selva" pageId="8" pageNumber="7" stateProvince="Heredia">Est.Bio. La Selva</location>
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, 50–;
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<quantity id="DA139B3EB707FFBF6D7BFB5447FB5ED0" box="[1099,1170,1154,1181]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="7" unit="m" value="150.0">
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||
<elevation id="96C6D1E8B707FFBF6D7BFB5447FB5ED0" box="[1099,1170,1154,1181]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="7" unit="m" value="150.0">150 m</elevation>
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</quantity>
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||
<geoCoordinate id="78DF501CB707FFBF6DAAFB54479D5ED0" box="[1178,1268,1154,1181]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="8" pageNumber="7" precision="925" value="10.433333">10°26′N</geoCoordinate>
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<geoCoordinate id="78DF501CB707FFBF6DCCFB5546365ED0" box="[1276,1375,1155,1181]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="8" pageNumber="7" precision="925" value="-84.01667">84°01′W</geoCoordinate>
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; Proy.
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<collectionCode id="7BFAAE1EB707FFBF69F2FB7242615EF3" box="[194,264,1188,1214]" pageId="8" pageNumber="7">ALAS</collectionCode>
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, INBio-OET” / “M/06/085;
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<date id="6955101BB707FFBF6B08FB7241BC5EF3" box="[568,725,1188,1215]" pageId="8" pageNumber="7" value="1993-05-02">
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<collectingDate id="7911E9F3B707FFBF6B08FB7241BC5EF3" box="[568,725,1188,1215]" pageId="8" pageNumber="7" value="1993-05-02">02 Mayo 1993</collectingDate>
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</date>
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; Bosque primario” / “
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<emphasis id="2F9FEAC9B707FFBF6AF1FB7247345EF3" bold="true" box="[961,1117,1188,1214]" pageId="8" pageNumber="7">
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<typeStatus id="C2508879B707FFBF6AF1FB7247315EF3" box="[961,1112,1188,1214]" pageId="8" pageNumber="7" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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;
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<taxonomicName id="DAEB4D58B707FFBF6D58FB72479F5EF3" authorityName="Chevrolat" authorityYear="1868" box="[1128,1270,1188,1214]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="2F9FEAC9B707FFBF6D58FB72479F5EF3" box="[1128,1270,1188,1214]" italics="true" pageId="8" pageNumber="7">Adelothyreus</emphasis>
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</taxonomicName>
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;
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<taxonomicName id="DAEB4D58B707FFBF6C32FB72462B5EF3" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[1282,1346,1188,1214]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="brevis">
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<emphasis id="2F9FEAC9B707FFBF6C32FB72462B5EF3" box="[1282,1346,1188,1214]" italics="true" pageId="8" pageNumber="7">brevis</emphasis>
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</taxonomicName>
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<specimenCount id="0BEDFD52B707FFBF6C78FB7E46345EF3" box="[1352,1373,1192,1214]" pageId="8" pageNumber="7" type="male">♂</specimenCount>
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; Otto, Muona & Córdoba-Alfaro; Det.
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<collectionCode id="7BFAAE1EB707FFBF6B18FB1041555EAD" box="[552,572,1222,1248]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="8" pageNumber="7">R</collectionCode>
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.
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<collectorName id="B01E530DB707FFBF6B70FB1041FB5EAD" box="[576,658,1222,1248]" pageId="8" pageNumber="7">L. Otto</collectorName>
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; 2019” (red printed label).
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<typeStatus id="C2508879B707FFBF6A8DFB10474C5EAD" box="[957,1061,1222,1248]" pageId="8" pageNumber="7" type="holotype">Holotype</typeStatus>
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to be deposited in
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<collectionCode id="7BFAAE1EB707FFBF6DC9FB1046575EAD" box="[1273,1342,1222,1248]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="8" pageNumber="7">INBC</collectionCode>
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||
.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="55F16550B707FFBF69F2FB3942CE5DC5" pageId="8" pageNumber="7" type="description">
|
||
<paragraph id="1D5436DBB707FFBF69F2FB3942CE5DC5" blockId="8.[192,1443,1263,1928]" pageId="8" pageNumber="7">
|
||
<emphasis id="2F9FEAC9B707FFBF69F2FB39416B5F44" bold="true" box="[194,514,1263,1289]" pageId="8" pageNumber="7">
|
||
Description. Male
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<typeStatus id="C2508879B707FFBF68ABFB39416B5F44" box="[411,514,1263,1289]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
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||
</emphasis>
|
||
: Length,
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||
<quantity id="DA139B3EB707FFBF6B5CFB3941AC5F47" box="[620,709,1263,1290]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="7" unit="mm" value="2.5">2.5 mm</quantity>
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||
. Width,
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||
<quantity id="DA139B3EB707FFBF6A14FB3940E25F47" box="[804,907,1263,1290]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="7" unit="mm" value="0.5">0.50 mm</quantity>
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||
. Body elongate; uniformly dark black; antennae dark brownish black; legs dark brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (
|
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<figureCitation id="85D02A5EB707FFBF69F9FAE4425E5F00" box="[201,311,1330,1357]" captionStart="Figures 10–16" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 10–16. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 10–16 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="8" pageNumber="7">Figure 10</figureCitation>
|
||
).
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||
<emphasis id="2F9FEAC9B707FFBF6878FAE542EF5F00" bold="true" box="[328,390,1331,1357]" pageId="8" pageNumber="7">Head</emphasis>
|
||
: Surface densely punctate, somewhat dullish, subspherical; frons convex, without median fovea or carina; apical margin of frontoclypeal region rounded, more than 2.0 times wider than base; mandibles stout, bidentate, densely punctate.
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<emphasis id="2F9FEAC9B707FFBF68CFFAA3410D5FC2" bold="true" box="[511,612,1397,1423]" pageId="8" pageNumber="7">Antenna</emphasis>
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: Pectinate from flagellomeres III–VIII, attaining about 2/3 the length of the body; flagellomere I longer than II; flagellomere II strongly, asymmetrically serrate; flagellomeres III–IV longer than wide, sub-equal, each shorter than V; flagellomeres V–VIII each sub-equal, longer than wide; flagellomere IX longer than VIII; rami on flagellomeres III–VIII arising at apical end, short and about as long as the length of the segment.
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<emphasis id="2F9FEAC9B707FFBF686CFA2A42BB5C5B" bold="true" box="[348,466,1532,1558]" pageId="8" pageNumber="7">Pronotum</emphasis>
|
||
: Surface somewhat dullish, closely punctate; longer than wide, with poorly developed hind angles; lateral sides parallel-sided, laterally sinuous above pronotal hind angle, apically arcuate; disc flattened without median groove; anterior lateral pronotal ridge extremely short, directed posteroventrally; posterior lateral pronotal ridge elongate, extending at least 3/4the length of pronotum (
|
||
<figureCitation id="85D02A5EB707FFBF6AC6F9B7470D5C36" box="[1014,1124,1633,1659]" captionStart="Figures 10–16" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 10–16. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 10–16 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="8" pageNumber="7">Figure 11</figureCitation>
|
||
); base sinuous, with circular strong impressions above scutellar shield.
|
||
<emphasis id="2F9FEAC9B707FFBF6BABF95440275CD1" bold="true" box="[667,846,1666,1692]" pageId="8" pageNumber="7">Scutellar shield</emphasis>
|
||
: Quadrate, sub-triangular, shallowly punctate, setose and distally rounded.
|
||
<emphasis id="2F9FEAC9B707FFBF6888F97242965CF3" bold="true" box="[440,511,1700,1726]" pageId="8" pageNumber="7">Elytra</emphasis>
|
||
: Striate indistinct; interstices flattened; surfaces shiny, closely punctate to rugose at basal 1/2, shallowly and closely punctate at apical 1/2.
|
||
<emphasis id="2F9FEAC9B707FFBF6A34F91040505CAD" bold="true" box="[772,825,1734,1760]" pageId="8" pageNumber="7">Legs</emphasis>
|
||
: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I–III simple; metatarsomere IV very short, excavate; metatarsomere V short with simple claws.
|
||
<emphasis id="2F9FEAC9B707FFBF6D33F8DF47385D6E" bold="true" box="[1027,1105,1801,1827]" pageId="8" pageNumber="7">Venter</emphasis>
|
||
(
|
||
<figureCitation id="85D02A5EB707FFBF6D6FF8DF47A75D6E" box="[1119,1230,1801,1827]" captionStart="Figures 10–16" captionStartId="9.[193,280,1019,1045]" captionTargetBox="[449,1184,213,989]" captionTargetId="figure-360@9.[449,1184,213,989]" captionTargetPageId="9" captionText="Figures 10–16. Costa Rican Adelothyreus species. 10) Adelothyreus brevis sp. nov., dorsal habitus. 11) Adelothyreus brevis sp. nov., pronotal lateral habitus. 12) Adelothyreus brevis sp. nov., ventral habitus. 13) Adelothyreus costaricensis sp. nov., male holotype, dorsal habitus. 14) Adelothyreus costaricensis sp. nov., male holotype, pronotal lateral habitus. 15) Adelothyreus costaricensis sp. nov., male holotype, ventral habitus. 16) Adelothyreus costaricensis sp. nov. female allotype, dorsal habitus. (Scale: 10–16 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="8" pageNumber="7">Figure 12</figureCitation>
|
||
): Closely punctate, with short, recumbent yellowish setae; hypomeron with apically widened notosternal antennal grooves; antennal grooves smooth, scarcely punctate apically; metepisterna caudally wide; elytral epipleura punctate; metacoxal plates parallel-sided.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="49946653B706FFBE69F1FC2D47735ED1" ID-DOI="http://doi.org/10.5281/zenodo.10831091" ID-Zenodo-Dep="10831091" httpUri="https://zenodo.org/record/10831091/files/figure.png" pageId="9" pageNumber="8" startId="9.[193,280,1019,1045]" targetBox="[449,1184,213,989]" targetPageId="9" targetType="figure">
|
||
<paragraph id="1D5436DBB706FFBE69F1FC2D47735ED1" blockId="9.[192,1440,1019,1180]" pageId="9" pageNumber="8">
|
||
<emphasis id="2F9FEAC9B706FFBE69F1FC2D42075E58" bold="true" box="[193,366,1019,1045]" pageId="9" pageNumber="8">Figures 10–16.</emphasis>
|
||
Costa Rican
|
||
<taxonomicName id="DAEB4D58B706FFBE6B3AFC2D419E5E58" box="[522,759,1019,1045]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="2F9FEAC9B706FFBE6B3AFC2D41F15E58" box="[522,664,1019,1045]" italics="true" pageId="9" pageNumber="8">Adelothyreus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B706FFBE6A30FC2D404C5E58" bold="true" box="[768,805,1019,1045]" pageId="9" pageNumber="8">10)</emphasis>
|
||
<taxonomicName id="DAEB4D58B706FFBE6A1EFC2D476F5E58" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[814,1030,1019,1045]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B706FFBE6A1EFC2D476F5E58" box="[814,1030,1019,1045]" italics="true" pageId="9" pageNumber="8">Adelothyreus brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B706FFBE6D20FC2A47035E5B" bold="true" box="[1040,1130,1020,1046]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="34AC57B2B706FFBE6D20FC2A47035E5B" box="[1040,1130,1020,1046]" pageId="9" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal habitus.
|
||
<emphasis id="2F9FEAC9B706FFBE6C19FC2D46265E58" bold="true" box="[1321,1359,1019,1045]" pageId="9" pageNumber="8">11)</emphasis>
|
||
<taxonomicName id="DAEB4D58B706FFBE6C68FC2D423F5E7A" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B706FFBE6C68FC2D423F5E7A" italics="true" pageId="9" pageNumber="8">Adelothyreus brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B706FFBE686CFBCB42DA5E7A" bold="true" box="[348,435,1053,1079]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="34AC57B2B706FFBE686CFBCB42DA5E7A" box="[348,435,1053,1079]" pageId="9" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, pronotal lateral habitus.
|
||
<emphasis id="2F9FEAC9B706FFBE6BFCFBCB41985E7A" bold="true" box="[716,753,1053,1079]" pageId="9" pageNumber="8">12)</emphasis>
|
||
<taxonomicName id="DAEB4D58B706FFBE6BC8FBCB40A55E7A" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[760,972,1053,1079]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="brevis" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B706FFBE6BC8FBCB40A55E7A" box="[760,972,1053,1079]" italics="true" pageId="9" pageNumber="8">Adelothyreus brevis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B706FFBE6AE2FBCB47405E7A" bold="true" box="[978,1065,1053,1079]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="34AC57B2B706FFBE6AE2FBCB47405E7A" box="[978,1065,1053,1079]" pageId="9" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ventral habitus.
|
||
<emphasis id="2F9FEAC9B706FFBE6DD5FBCB46625E7A" bold="true" box="[1253,1291,1053,1079]" pageId="9" pageNumber="8">13)</emphasis>
|
||
<taxonomicName id="DAEB4D58B706FFBE6C22FBCB422E5E15" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B706FFBE6C22FBCB422E5E15" italics="true" pageId="9" pageNumber="8">Adelothyreus costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B706FFBE687EFBE942CF5E14" bold="true" box="[334,422,1087,1113]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="34AC57B2B706FFBE687EFBE942CF5E14" box="[334,422,1087,1113]" pageId="9" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, dorsal habitus.
|
||
<emphasis id="2F9FEAC9B706FFBE6A3DFBE8405B5E15" bold="true" box="[781,818,1086,1112]" pageId="9" pageNumber="8">14)</emphasis>
|
||
<taxonomicName id="DAEB4D58B706FFBE6A09FBE9473F5E15" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[825,1110,1087,1113]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B706FFBE6A09FBE9473F5E15" box="[825,1110,1087,1113]" italics="true" pageId="9" pageNumber="8">Adelothyreus costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B706FFBE6D6DFBE947DC5E14" bold="true" box="[1117,1205,1087,1113]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="34AC57B2B706FFBE6D6DFBE947DC5E14" box="[1117,1205,1087,1113]" pageId="9" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, pronotal lateral habitus.
|
||
<emphasis id="2F9FEAC9B706FFBE6884FBB642B05E37" bold="true" box="[436,473,1120,1146]" pageId="9" pageNumber="8">15)</emphasis>
|
||
<taxonomicName id="DAEB4D58B706FFBE68D3FBB6406B5E37" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[483,770,1120,1146]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B706FFBE68D3FBB6406B5E37" box="[483,770,1120,1146]" italics="true" pageId="9" pageNumber="8">Adelothyreus costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B706FFBE6A3CFBB6400E5E37" bold="true" box="[780,871,1120,1146]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="34AC57B2B706FFBE6A3CFBB6400E5E37" box="[780,871,1120,1146]" pageId="9" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, ventral habitus.
|
||
<emphasis id="2F9FEAC9B706FFBE6DD2FBB646615E37" bold="true" box="[1250,1288,1120,1146]" pageId="9" pageNumber="8">16)</emphasis>
|
||
<taxonomicName id="DAEB4D58B706FFBE6C22FBB6422E5ED1" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B706FFBE6C22FBB6422E5ED1" italics="true" pageId="9" pageNumber="8">Adelothyreus costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B706FFBE687EFB5442CC5ED1" bold="true" box="[334,421,1154,1180]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="34AC57B2B706FFBE687EFB5442CC5ED1" box="[334,421,1154,1180]" pageId="9" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female allotype, dorsal habitus. (Scale: 10–16 = 1.0 mm)
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="55F16550B706FFBE69F0FB0747E65EA6" box="[192,1167,1233,1259]" pageId="9" pageNumber="8" type="distribution">
|
||
<paragraph id="1D5436DBB706FFBE69F0FB0747E65EA6" blockId="9.[192,1167,1233,1259]" box="[192,1167,1233,1259]" pageId="9" pageNumber="8">
|
||
<emphasis id="2F9FEAC9B706FFBE69F0FB0742325EA6" bold="true" box="[192,347,1233,1259]" pageId="9" pageNumber="8">Distribution.</emphasis>
|
||
This eucnemid species is known only from its
|
||
<typeStatus id="C2508879B706FFBE6A53FB0740FA5EA6" box="[867,915,1233,1259]" pageId="9" pageNumber="8">type</typeStatus>
|
||
location in
|
||
<collectingCountry id="65FC764BB706FFBE6D28FB0747E65EA6" box="[1048,1167,1233,1259]" name="Costa Rica" pageId="9" pageNumber="8">Costa Rica</collectingCountry>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55F16550B706FFBE69F0FB2D47F05F5B" box="[192,1177,1275,1302]" pageId="9" pageNumber="8" type="biology_ecology">
|
||
<paragraph id="1D5436DBB706FFBE69F0FB2D47F05F5B" blockId="9.[192,1177,1275,1302]" box="[192,1177,1275,1302]" pageId="9" pageNumber="8">
|
||
<emphasis id="2F9FEAC9B706FFBE69F0FB2D42495F58" bold="true" box="[192,288,1275,1301]" pageId="9" pageNumber="8">Biology.</emphasis>
|
||
The
|
||
<typeStatus id="C2508879B706FFBE6866FB2A42D15F5B" box="[342,440,1276,1302]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
was taken from a primary forest. Larvae and pupae are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55F16550B706FFBE69F0FAF1460A5F0D" box="[192,1379,1318,1345]" pageId="9" pageNumber="8" type="etymology">
|
||
<paragraph id="1D5436DBB706FFBE69F0FAF1460A5F0D" blockId="9.[192,1379,1318,1345]" box="[192,1379,1318,1345]" pageId="9" pageNumber="8">
|
||
<emphasis id="2F9FEAC9B706FFBE69F0FAF1422D5F0C" bold="true" box="[192,324,1319,1345]" pageId="9" pageNumber="8">Etymology.</emphasis>
|
||
The specific epithet is derived from the presence of a very short anterior pronotal lateral ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |