248 lines
28 KiB
XML
248 lines
28 KiB
XML
<document id="4107F1679131225D5B94B17F27040D1A" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki & Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB705FFBC69F0FBD741275FD1" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Adelothyreus totus Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="4" lastPageNumber="10" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="9" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="5DF3654915C01418ABA28CFFE49FE4CF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2ACCFA63E90DF15B5A83C87A9643FC66">
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<mods:title id="1388AFE148B5D37A007B8E1097C76339">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
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</mods:titleInfo>
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<mods:name id="E9A957E1DD15432DB307681EB82A6A6B" type="personal">
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<mods:role id="4D00DB8A071FF1F5E74978C6683D06C0">
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<mods:roleTerm id="344F8AD1DCC534DA4415DBFE33BD9C84">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7B025C2E033C12A0FC4ADB70D152A48E">Otto, Robert L.</mods:namePart>
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<mods:affiliation id="DC6300C5B1DF0602EEFC573ABE6C7A8B">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:name id="A10EF8C9FEA84DCF2B47421B5619B217" type="personal">
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<mods:role id="6791C32EF39746089DE5B29B542DC551">
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<mods:roleTerm id="DD60DAEBD26DCE4E9EC5D62AC06DCCEF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1A0735A5A96334CCD4D064F4F923710">Muona, Jyrki</mods:namePart>
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<mods:affiliation id="0769CC7BDB3669E52A035B6862521F2F">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
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</mods:name>
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<mods:name id="F183872C599CAC47A4C15B8AECC1FF0A" type="personal">
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<mods:roleTerm id="AA715B806BCECF6AF7E05B28B84D8522">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FC9FC4D69EDE3B14FF97311D1ED3DB5C">Córdoba-Alfaro, Jim</mods:namePart>
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<mods:affiliation id="5201F147C9A3A6E6C89B1E3DAAF5A29F">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="D5DE1766010B12B9E090330A4853B3D6">text</mods:typeOfResource>
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<mods:titleInfo id="A3749FD10C3AA9F345E2E15CEAB2C1AB">
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<mods:title id="5F019B08D0B6B77C70D2BD4BAE2D6FAE">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="6D82369ADD3AA82AE2E47C9EAF6105E2">
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<mods:date id="9CAB45322ACE52D48DFFD0F90F4BB251">2023</mods:date>
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<mods:detail id="FCD4A9370CA668EB999A10A23A90D899" type="pubDate">
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<mods:number id="C8C1D255FA986142518D7658BA5BA77A">2023-05-26</mods:number>
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</mods:detail>
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<mods:detail id="92F5073CB41A1FBF7C0C8BBC194AC17D" type="volume">
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<mods:number id="C5E1A11CB38023A6496A72744E5B81F4">2023</mods:number>
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</mods:detail>
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<mods:detail id="00C1C5AC5F82DF5310605B71B9FF94CC" type="issue">
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<mods:number id="9F023BB7674C9D6BD2063F0063A32CE7">991</mods:number>
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<mods:start id="9CAC0BE2A589BF10DD79D48C7DE1A423">1</mods:start>
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<mods:classification id="573161A54994848203E2BEC0B2F92976">journal article</mods:classification>
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<mods:identifier id="1A9FCFA4E76368612B71E84E17024A49" type="DOI">10.5281/zenodo.10831085</mods:identifier>
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<mods:identifier id="0EF3E1C0284DA455B613DF7807C6C41A" type="ISSN">1942-1354</mods:identifier>
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<treatment id="954287CDB705FFBC69F0FBD741275FD1" ID-DOI="http://doi.org/10.5281/zenodo.8109072" ID-Zenodo-Dep="8109072" LSID="urn:lsid:plazi:treatment:954287CDB705FFBC69F0FBD741275FD1" httpUri="http://treatment.plazi.org/id/954287CDB705FFBC69F0FBD741275FD1" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
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<subSubSection id="55F16550B705FFBD69F0FBD7476D5E53" box="[192,1028,1025,1054]" pageId="10" pageNumber="9" type="nomenclature">
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<paragraph id="1D5436DBB705FFBD69F0FBD7476D5E53" blockId="10.[192,1028,1025,1088]" box="[192,1028,1025,1054]" pageId="10" pageNumber="9">
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<heading id="461C81B7B705FFBD69F0FBD7476D5E53" box="[192,1028,1025,1054]" fontSize="12" level="2" pageId="10" pageNumber="9" reason="2">
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<emphasis id="2F9FEAC9B705FFBD69F0FBD7476D5E53" bold="true" box="[192,1028,1025,1054]" pageId="10" pageNumber="9">
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<taxonomicName id="DAEB4D58B705FFBD69F0FBD740085E53" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,865,1025,1054]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="totus" status="sp. nov.">
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<emphasis id="2F9FEAC9B705FFBD69F0FBD742C35E53" bold="true" box="[192,426,1025,1054]" italics="true" pageId="10" pageNumber="9">Adelothyreus totus</emphasis>
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Otto, Muona and Córdoba-Alfaro
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</taxonomicName>
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,
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<taxonomicNameLabel id="34AC57B2B705FFBD6A40FBD4476D5E53" box="[880,1028,1025,1054]" pageId="10" pageNumber="9" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="55F16550B705FFBD69F0FBF342375E0D" box="[192,350,1061,1088]" pageId="10" pageNumber="9" type="description">
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||
<paragraph id="1D5436DBB705FFBD69F0FBF342375E0D" blockId="10.[192,1028,1025,1088]" box="[192,350,1061,1088]" pageId="10" pageNumber="9">
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||
<figureCitation id="85D02A5EB705FFBD69F0FBF342375E0D" box="[192,350,1061,1088]" captionStart="Figures 17–22" captionStartId="11.[192,278,1085,1111]" captionTargetBox="[449,1187,218,1056]" captionTargetId="figure-12@11.[449,1187,218,1056]" captionTargetPageId="11" captionText="Figures 17–22. Costa Rican Eucnemidae. 17) Adelothyreus totus sp. nov., dorsal habitus. 18) Adelothyreus totus sp. nov., pronotal lateral habitus. 19) Adelothyreus totus sp. nov., ventral habitus. 20) Quirsfeldia stethonoides sp. nov., dorsal habitus. 21) Quirsfeldia stethonoides sp. nov., pronotal lateral habitus. 22) Quirsfeldia stethonoides sp. nov., ventral habitus. (Scale: 17–22 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831093" httpUri="https://zenodo.org/record/10831093/files/figure.png" pageId="10" pageNumber="9">Figures 17–19</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="55F16550B705FFBD69F0FB99428C5EC6" pageId="10" pageNumber="9" type="diagnosis">
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<paragraph id="1D5436DBB705FFBD69F0FB99428C5EC6" blockId="10.[192,1440,1103,1164]" pageId="10" pageNumber="9">
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<emphasis id="2F9FEAC9B705FFBD69F0FB9942555E24" bold="true" box="[192,316,1103,1129]" pageId="10" pageNumber="9">Diagnosis.</emphasis>
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A complete lateral pronotal ridge will distinguish the new species from all known
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<taxonomicName id="DAEB4D58B705FFBD6DECFB9943835EC6" pageId="10" pageNumber="9">
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<emphasis id="2F9FEAC9B705FFBD6DECFB9946035E24" box="[1244,1386,1103,1129]" italics="true" pageId="10" pageNumber="9">Adelothyreus</emphasis>
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species
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</taxonomicName>
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present in
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<collectingCountry id="65FC764BB705FFBD6855FBA442895EC6" box="[357,480,1137,1164]" name="Costa Rica" pageId="10" pageNumber="9">Costa Rica</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="55F16550B705FFBD69F0FB4D46C85EB4" pageId="10" pageNumber="9" type="materials_examined">
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<paragraph id="1D5436DBB705FFBD69F0FB4D46C85EB4" blockId="10.[192,1441,1179,1274]" pageId="10" pageNumber="9">
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<emphasis id="2F9FEAC9B705FFBD69F0FB4D42D85EF8" bold="true" box="[192,433,1179,1205]" pageId="10" pageNumber="9">Type material. Male</emphasis>
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<materialsCitation id="AD833C86B705FFBD688CFB4D46C85EB4" collectingDate="1988-04" collectionCode="R, INBC" collectorName="H. A. Hespenheide & L. Otto" country="Costa Rica" latitude="10.433333" location="Pto. Viejo" longLatPrecision="1298" longitude="-84.01667" pageId="10" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
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<emphasis id="2F9FEAC9B705FFBD688CFB4D414A5EF8" bold="true" box="[444,547,1179,1205]" pageId="10" pageNumber="9">
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<typeStatus id="C2508879B705FFBD688CFB4D414A5EF8" box="[444,547,1179,1205]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
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</emphasis>
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: “
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<collectingCountry id="65FC764BB705FFBD6B0DFB4D418C5EF8" box="[573,741,1179,1205]" name="Costa Rica" pageId="10" pageNumber="9">COSTA RICA</collectingCountry>
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:
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<collectingRegion id="DF2FF839B705FFBD6BC3FB4D40255EF8" box="[755,844,1179,1205]" country="Costa Rica" name="Heredia" pageId="10" pageNumber="9">Heredia</collectingRegion>
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pr.; La Selva Bio. Sta.;
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<quantity id="DA139B3EB705FFBD6D51FB4D47F65EF8" box="[1121,1183,1179,1205]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="9" unit="km" value="3.0">3 km</quantity>
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S
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<location id="18346000B705FFBD6DF2FB4A465E5EF8" LSID="urn:lsid:plazi:treatment:954287CDB705FFBC69F0FBD741275FD1:18346000B705FFBD6DF2FB4A465E5EF8" box="[1218,1335,1179,1206]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" name="Pto. Viejo" pageId="10" pageNumber="9" stateProvince="Heredia">Pto. Viejo</location>
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;
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<geoCoordinate id="78DF501CB705FFBD6C76FB4D46C95EFB" box="[1350,1440,1179,1206]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="10" pageNumber="9" precision="925" value="10.433333">10°26′N</geoCoordinate>
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<geoCoordinate id="78DF501CB705FFBD69F0FB6B42485E9A" box="[192,289,1213,1239]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="10" pageNumber="9" precision="925" value="-84.01667">84°01′W</geoCoordinate>
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” / “I.
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<date id="6955101BB705FFBD686BFB6B42DE5E9A" box="[347,439,1213,1239]" pageId="10" pageNumber="9" value="1988-04">
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<collectingDate id="7911E9F3B705FFBD686BFB6B42DE5E9A" box="[347,439,1213,1239]" pageId="10" pageNumber="9" value="1988-04">IV.1988</collectingDate>
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</date>
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;
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<collectorName id="B01E530DB705FFBD68F2FB6B41FB5E9A" box="[450,658,1213,1239]" pageId="10" pageNumber="9">H.A. Hespenheide</collectorName>
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||
” (“I”, “IV” and “8” handwritten) / “
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<emphasis id="2F9FEAC9B705FFBD6D14FB6B47A95E9A" bold="true" box="[1060,1216,1213,1239]" pageId="10" pageNumber="9">
|
||
<typeStatus id="C2508879B705FFBD6D14FB6B47D25E9A" box="[1060,1211,1213,1239]" pageId="10" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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;
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<taxonomicName id="DAEB4D58B705FFBD6DFEFB6B46355E9A" authorityName="Chevrolat" authorityYear="1868" box="[1230,1372,1213,1239]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="2F9FEAC9B705FFBD6DFEFB6B46355E9A" box="[1230,1372,1213,1239]" italics="true" pageId="10" pageNumber="9">Adelothyreus</emphasis>
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</taxonomicName>
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;
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<taxonomicName id="DAEB4D58B705FFBD6C5AFB6846C95E9A" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[1386,1440,1214,1239]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="totus">
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<emphasis id="2F9FEAC9B705FFBD6C5AFB6846C95E9A" box="[1386,1440,1214,1239]" italics="true" pageId="10" pageNumber="9">totus</emphasis>
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</taxonomicName>
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<specimenCount id="0BEDFD52B705FFBD69F0FB3543BC5EB4" box="[192,213,1251,1273]" pageId="10" pageNumber="9" type="male">♂</specimenCount>
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; Otto, Muona & Córdoba-Alfaro; Det.
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<collectionCode id="7BFAAE1EB705FFBD6BB8FB3641F55EB7" box="[648,668,1248,1274]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="9">R</collectionCode>
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||
.
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<collectorName id="B01E530DB705FFBD6B90FB36419B5EB4" box="[672,754,1247,1274]" pageId="10" pageNumber="9">L. Otto</collectorName>
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; 2021” (red printed label).
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||
<typeStatus id="C2508879B705FFBD6D2CFB0947ED5EB4" box="[1052,1156,1247,1273]" pageId="10" pageNumber="9" type="holotype">Holotype</typeStatus>
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to be deposited in
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<collectionCode id="7BFAAE1EB705FFBD6C68FB0946F45EB4" box="[1368,1437,1247,1273]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="10" pageNumber="9">INBC</collectionCode>
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||
.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="55F16550B705FFBD69F0FADF40205DCB" pageId="10" pageNumber="9" type="description">
|
||
<paragraph id="1D5436DBB705FFBD69F0FADF40205DCB" blockId="10.[191,1442,1289,1927]" pageId="10" pageNumber="9">
|
||
<emphasis id="2F9FEAC9B705FFBD69F0FADF41695F6E" bold="true" box="[192,512,1289,1315]" pageId="10" pageNumber="9">
|
||
Description. Male
|
||
<typeStatus id="C2508879B705FFBD68A9FADF41695F6E" box="[409,512,1289,1315]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: Length, 3.0 mm. Width,
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||
<quantity id="DA139B3EB705FFBD6A13FADF40E05F69" box="[803,905,1289,1316]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="9" unit="mm" value="0.75">0.75 mm</quantity>
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. Body elongate; uniformly dark black; antennae dark brownish black; legs dark brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (
|
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<figureCitation id="85D02A5EB705FFBD69F8FA9B425C5F2A" box="[200,309,1357,1383]" captionStart="Figures 17–22" captionStartId="11.[192,278,1085,1111]" captionTargetBox="[449,1187,218,1056]" captionTargetId="figure-12@11.[449,1187,218,1056]" captionTargetPageId="11" captionText="Figures 17–22. Costa Rican Eucnemidae. 17) Adelothyreus totus sp. nov., dorsal habitus. 18) Adelothyreus totus sp. nov., pronotal lateral habitus. 19) Adelothyreus totus sp. nov., ventral habitus. 20) Quirsfeldia stethonoides sp. nov., dorsal habitus. 21) Quirsfeldia stethonoides sp. nov., pronotal lateral habitus. 22) Quirsfeldia stethonoides sp. nov., ventral habitus. (Scale: 17–22 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831093" httpUri="https://zenodo.org/record/10831093/files/figure.png" pageId="10" pageNumber="9">Figure 17</figureCitation>
|
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).
|
||
<emphasis id="2F9FEAC9B705FFBD6876FA9B42ED5F2A" bold="true" box="[326,388,1357,1383]" pageId="10" pageNumber="9">Head</emphasis>
|
||
: Surface densely punctate, somewhat dullish, subspherical; frons convex, without median fovea or carina; apical margin of frontoclypeal region rounded, more than 2.0 times wider than base; mandibles stout, bidentate, densely punctate.
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||
<emphasis id="2F9FEAC9B705FFBD68CEFA47410A5FE6" bold="true" box="[510,611,1425,1451]" pageId="10" pageNumber="9">Antenna</emphasis>
|
||
: Pectinate from flagellomeres III–VIII, attaining about 3/4 the length of the body; flagellomere I longer than II; flagellomere II strongly, asymmetrically serrate; flagellomere III longer than wide, shorter than IV; flagellomeres IV–VIII each sub-equal, longer than wide; flagellomere IX longer than VIII; rami on flagellomeres III–VIII arising at apical end, elongate and about 2.0 times longer than length of segment.
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<emphasis id="2F9FEAC9B705FFBD69F0F9CF42515C7E" bold="true" box="[192,312,1561,1587]" pageId="10" pageNumber="9">Pronotum</emphasis>
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: Surface somewhat dullish, closely punctate; longer than wide, with poorly developed hind angles; lateral sides parallel-sided, apically arcuate; disc flattened with delicate median groove; anterior lateral pronotal ridge completely connected to the posterior lateral pronotal ridge forming a complete lateral ridge (
|
||
<figureCitation id="85D02A5EB705FFBD6C14F98A46FA5C3A" box="[1316,1427,1628,1655]" captionStart="Figures 17–22" captionStartId="11.[192,278,1085,1111]" captionTargetBox="[449,1187,218,1056]" captionTargetId="figure-12@11.[449,1187,218,1056]" captionTargetPageId="11" captionText="Figures 17–22. Costa Rican Eucnemidae. 17) Adelothyreus totus sp. nov., dorsal habitus. 18) Adelothyreus totus sp. nov., pronotal lateral habitus. 19) Adelothyreus totus sp. nov., ventral habitus. 20) Quirsfeldia stethonoides sp. nov., dorsal habitus. 21) Quirsfeldia stethonoides sp. nov., pronotal lateral habitus. 22) Quirsfeldia stethonoides sp. nov., ventral habitus. (Scale: 17–22 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831093" httpUri="https://zenodo.org/record/10831093/files/figure.png" pageId="10" pageNumber="9">Figure 18</figureCitation>
|
||
); base sinuous, strongly impressed above scutellar shield.
|
||
<emphasis id="2F9FEAC9B705FFBD6A0BF9A840875CD5" bold="true" box="[827,1006,1662,1688]" pageId="10" pageNumber="9">Scutellar shield</emphasis>
|
||
: Short, transverse, shallowly punctate, setose and distally truncated.
|
||
<emphasis id="2F9FEAC9B705FFBD6B24F97741355CF6" bold="true" box="[532,604,1697,1723]" pageId="10" pageNumber="9">Elytra</emphasis>
|
||
: Shallow striate indicated at humeri, indistinct elsewhere; interstices flattened; surfaces shiny, closely punctate to rugose at basal 1/2, shallowly punctate at apical 1/2.
|
||
<emphasis id="2F9FEAC9B705FFBD6D84F91447805C91" bold="true" box="[1204,1257,1730,1756]" pageId="10" pageNumber="9">Legs</emphasis>
|
||
: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I–III simple; metatarsomere IV very short, excavate; metatarsomere V short with simple claws.
|
||
<emphasis id="2F9FEAC9B705FFBD69F1F8FE42665D0F" bold="true" box="[193,271,1832,1858]" pageId="10" pageNumber="9">Venter</emphasis>
|
||
(
|
||
<figureCitation id="85D02A5EB705FFBD682CF8FE42E55D0F" box="[284,396,1832,1858]" captionStart="Figures 17–22" captionStartId="11.[192,278,1085,1111]" captionTargetBox="[449,1187,218,1056]" captionTargetId="figure-12@11.[449,1187,218,1056]" captionTargetPageId="11" captionText="Figures 17–22. Costa Rican Eucnemidae. 17) Adelothyreus totus sp. nov., dorsal habitus. 18) Adelothyreus totus sp. nov., pronotal lateral habitus. 19) Adelothyreus totus sp. nov., ventral habitus. 20) Quirsfeldia stethonoides sp. nov., dorsal habitus. 21) Quirsfeldia stethonoides sp. nov., pronotal lateral habitus. 22) Quirsfeldia stethonoides sp. nov., ventral habitus. (Scale: 17–22 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831093" httpUri="https://zenodo.org/record/10831093/files/figure.png" pageId="10" pageNumber="9">Figure 19</figureCitation>
|
||
): Closely punctate, with short, recumbent yellowish setae; hypomeron with apically widened notosternal antennal grooves; antennal grooves apically punctate, basally smooth; metepisterna caudally wide; elytral epipleura punctate; metacoxal plates parallel-sided.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="49946653B704FFBC69F0FBEB41A95EF0" ID-DOI="http://doi.org/10.5281/zenodo.10831093" ID-Zenodo-Dep="10831093" httpUri="https://zenodo.org/record/10831093/files/figure.png" pageId="11" pageNumber="10" startId="11.[192,278,1085,1111]" targetBox="[449,1187,218,1056]" targetPageId="11" targetType="figure">
|
||
<paragraph id="1D5436DBB704FFBC69F0FBEB41A95EF0" blockId="11.[192,1440,1085,1213]" pageId="11" pageNumber="10">
|
||
<emphasis id="2F9FEAC9B704FFBC69F0FBEB42035E15" bold="true" box="[192,362,1085,1112]" pageId="11" pageNumber="10">Figures 17–22.</emphasis>
|
||
Costa Rican
|
||
<taxonomicName id="DAEB4D58B704FFBC6B33FBE841E65E15" authorityName="Eschscholtz" authorityYear="1829" box="[515,655,1086,1112]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="family">Eucnemidae</taxonomicName>
|
||
.
|
||
<emphasis id="2F9FEAC9B704FFBC6BAAFBEB41D65E1A" bold="true" box="[666,703,1085,1111]" pageId="11" pageNumber="10">17)</emphasis>
|
||
<taxonomicName id="DAEB4D58B704FFBC6BF6FBE840F95E15" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[710,912,1086,1112]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="totus" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B704FFBC6BF6FBE840F95E15" box="[710,912,1086,1112]" italics="true" pageId="11" pageNumber="10">Adelothyreus totus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B704FFBC6AA8FBE840995E15" bold="true" box="[920,1008,1086,1112]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="34AC57B2B704FFBC6AA8FBE840995E15" box="[920,1008,1086,1112]" pageId="11" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal habitus.
|
||
<emphasis id="2F9FEAC9B704FFBC6D9AFBEB47A65E1A" bold="true" box="[1194,1231,1085,1111]" pageId="11" pageNumber="10">18)</emphasis>
|
||
<taxonomicName id="DAEB4D58B704FFBC6DE6FBE846C95E15" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[1238,1440,1086,1112]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="totus" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B704FFBC6DE6FBE846C95E15" box="[1238,1440,1086,1112]" italics="true" pageId="11" pageNumber="10">Adelothyreus totus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B704FFBC69F0FB89427E5E34" bold="true" box="[192,279,1119,1145]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="34AC57B2B704FFBC69F0FB89427E5E34" box="[192,279,1119,1145]" pageId="11" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, pronotal lateral habitus.
|
||
<emphasis id="2F9FEAC9B704FFBC6B04FB8941305E34" bold="true" box="[564,601,1119,1145]" pageId="11" pageNumber="10">19)</emphasis>
|
||
<taxonomicName id="DAEB4D58B704FFBC6B50FB8940425E34" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[608,811,1119,1145]" class="Insecta" family="Eucnemidae" genus="Adelothyreus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="totus" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B704FFBC6B50FB8940425E34" box="[608,811,1119,1145]" italics="true" pageId="11" pageNumber="10">Adelothyreus totus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B704FFBC6A01FB8940E05E34" bold="true" box="[817,905,1119,1145]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="34AC57B2B704FFBC6A01FB8940E05E34" box="[817,905,1119,1145]" pageId="11" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ventral habitus.
|
||
<emphasis id="2F9FEAC9B704FFBC6D78FB8947075E34" bold="true" box="[1096,1134,1119,1145]" pageId="11" pageNumber="10">20)</emphasis>
|
||
<taxonomicName id="DAEB4D58B704FFBC6D44FB8946105E34" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[1140,1401,1119,1145]" class="Insecta" family="Eucnemidae" genus="Quirsfeldia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="stethonoides" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B704FFBC6D44FB8946105E34" box="[1140,1401,1119,1145]" italics="true" pageId="11" pageNumber="10">Quirsfeldia stethonoides</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B704FFBC6CB0FB8943985ED6" bold="true" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="34AC57B2B704FFBC6CB0FB8943985ED6" pageId="11" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, dorsal habitus.
|
||
<emphasis id="2F9FEAC9B704FFBC6894FB5742A05ED6" bold="true" box="[420,457,1153,1179]" pageId="11" pageNumber="10">21)</emphasis>
|
||
<taxonomicName id="DAEB4D58B704FFBC68E0FB5741BC5ED6" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[464,725,1153,1179]" class="Insecta" family="Eucnemidae" genus="Quirsfeldia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="stethonoides" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B704FFBC68E0FB5741BC5ED6" box="[464,725,1153,1179]" italics="true" pageId="11" pageNumber="10">Quirsfeldia stethonoides</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B704FFBC6BE9FB5740595ED6" bold="true" box="[729,816,1153,1179]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="34AC57B2B704FFBC6BE9FB5740595ED6" box="[729,816,1153,1179]" pageId="11" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, pronotal lateral habitus.
|
||
<emphasis id="2F9FEAC9B704FFBC6D79FB5747065ED6" bold="true" box="[1097,1135,1153,1179]" pageId="11" pageNumber="10">22)</emphasis>
|
||
<taxonomicName id="DAEB4D58B704FFBC6D46FB5746105ED6" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[1142,1401,1153,1179]" class="Insecta" family="Eucnemidae" genus="Quirsfeldia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="stethonoides" status="sp. nov.">
|
||
<emphasis id="2F9FEAC9B704FFBC6D46FB5746105ED6" box="[1142,1401,1153,1179]" italics="true" pageId="11" pageNumber="10">Quirsfeldia stethonoides</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2F9FEAC9B704FFBC6CB0FB5743985EF0" bold="true" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="34AC57B2B704FFBC6CB0FB5743985EF0" pageId="11" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ventral habitus. (Scale: 17–22 = 1.0 mm)
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="55F16550B704FFBC69F0FADB47FE5F65" box="[192,1175,1293,1320]" pageId="11" pageNumber="10" type="distribution">
|
||
<paragraph id="1D5436DBB704FFBC69F0FADB47FE5F65" blockId="11.[192,1175,1293,1320]" box="[192,1175,1293,1320]" pageId="11" pageNumber="10">
|
||
<emphasis id="2F9FEAC9B704FFBC69F0FADB42325F6A" bold="true" box="[192,347,1293,1319]" pageId="11" pageNumber="10">Distribution.</emphasis>
|
||
This eucnemid species is only known from its
|
||
<typeStatus id="C2508879B704FFBC6A53FAD840FA5F65" box="[867,915,1294,1320]" pageId="11" pageNumber="10">type</typeStatus>
|
||
location in
|
||
<collectingCountry id="65FC764BB704FFBC6D28FAD847FB5F65" box="[1048,1170,1294,1320]" name="Costa Rica" pageId="11" pageNumber="10">Costa Rica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55F16550B704FFBC69F0FAE1466C5F1C" box="[192,1285,1335,1362]" pageId="11" pageNumber="10" type="biology_ecology">
|
||
<paragraph id="1D5436DBB704FFBC69F0FAE1466C5F1C" blockId="11.[192,1285,1335,1362]" box="[192,1285,1335,1362]" pageId="11" pageNumber="10">
|
||
<emphasis id="2F9FEAC9B704FFBC69F0FAE142495F1C" bold="true" box="[192,288,1335,1361]" pageId="11" pageNumber="10">Biology.</emphasis>
|
||
No biological information is known for the new species. Larvae and pupae are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="55F16550B704FFBC69F0FAB741275FD1" pageId="11" pageNumber="10" type="etymology">
|
||
<paragraph id="1D5436DBB704FFBC69F0FAB741275FD1" blockId="11.[192,1440,1377,1437]" pageId="11" pageNumber="10">
|
||
<emphasis id="2F9FEAC9B704FFBC69F0FAB7422D5F36" bold="true" box="[192,324,1377,1403]" pageId="11" pageNumber="10">Etymology.</emphasis>
|
||
The specific epithet is derived from ‘totus’, a Latin adjectival word for complete due to the presence of a complete lateral pronotal ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |