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<mods:title>Phylogenetic and multivariate analyses of Gekko smithii Gray, 1842 recover a new species from Peninsular Malaysia and support the resurrection of G. albomaculatus (Giebel, 1861) from Sumatra</mods:title>
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<mods:namePart>Grismer, L. Lee</mods:namePart>
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<mods:namePart>Quah, Evan S. H.</mods:namePart>
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<mods:affiliation>Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400, Kota Kinabalu, Sabah, Malaysia &amp; School of Biological Sciences, Universiti Sains Malaysia, 11800 Minden, Penang, Malaysia</mods:affiliation>
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<mods:namePart>Anuar, Shahrul</mods:namePart>
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<taxonomicName LSID="BD8129DA-B813-5A5D-A30F-7B6887ECE0E6" authority="Gray, 1842" authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">Gekko smithii Gray, 1842</taxonomicName>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution and color pattern variation in Gekko smithii and G. hulk sp. nov. on the Thai-Malay Peninsula. A. Gekko smithii (LSUHC 15052) from Perlis State Park, Perlis, Peninsular Malaysia. B. Gekko smithii (LSUDPC 12884) from Puncak Janing, Kedah, Peninsualr Malaysia. C. Gekko smithii (LSUHC 9626) from Sedim, Kedah, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12869) from Penang Hill (type locality), Penang, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12886) from Lata Kedondong, Selangor, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12887) from Kota Damansara, Selangor, Peninsular Malaysia. G. Gekko smithii (LSUDPC 11174) from Khaopu Khaoya, Nakhon Si Thamarat Province, Thailand. H. Gekko smithii (LSUDPC 11173) from Bacho Falls, Narathiwat Province, Thailand. I. Gekko smithii (LSUDPC 12878) from the Weang District, Narathiwat Province, Thailand. J. Gekko hulk sp. nov. (LSUDPC 12880) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 8696) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUDPC 7991) from Gunung Tebu, Terengganu, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12897) from Merapoh, Pahang, Peninsular Malaysia. N. Gekko hulk sp. nov. (LSUDPC 12897) from Pulau Tioman, Pahang, Peninsular Malaysia. Photographs by L. Lee Grismer (A, K, L, N), Evan S. H. Quah (B, C, D), Kurt H. P. Guek (E), Steven Wong (F), Henrick Bringsoe (G, H); Parinya Pawangkhanant (I), Ian Dugdale (J), and Nick Baker (M). Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature." figureDoi="10.3897/vz.72.e77702.figure7" httpUri="https://binary.pensoft.net/fig/641483" pageId="0" pageNumber="47">Figures 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">, 8</figureCitation>
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<taxonomicName authority="Brongniart, 1800" authorityName="Stoliczka" authorityYear="1870" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko Smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="Smithii">Gecko Smithii</taxonomicName>
: 
<bibRefCitation author="Gray, JE" journalOrPublisher="Zoological Miscellany" pageId="0" pageNumber="47" pagination="57 - 59" refId="B37" refString="Gray, JE, 1842. Description of some new species of reptiles, chiefly from the British Museum collection. Zoological Miscellany 2: 57 - 59" title="Description of some new species of reptiles, chiefly from the British Museum collection." volume="2" year="1842">Gray 1842</bibRefCitation>
:57; 
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B124" refString="Stoliczka, F, 1870. Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39(2): 134-228, pls. IX-XII." title="Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39 (2): 134 - 228, pls. IX-XII." year="1870">Stoliczka 1870</bibRefCitation>
:161
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<taxonomicName authorityName="Cantor" authorityYear="1847" class="Insecta" family="Gekkonidae" genus="Platydactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platydactylus Stentor" order="Squamata" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="Stentor">Platydactylus Stentor</taxonomicName>
: 
<bibRefCitation author="Cantor, TE" journalOrPublisher="Zoologische Mededeelingen uitgegeven door het Rijksmuseum van Natuurlijke Historie" pageId="0" pageNumber="47" refId="B17" refString="Cantor, TE, 1847. Catalogue of retiles inhabiting the Malayan Peninsula and islands, collected or observed by Theodore Cantor, Esq., M.D. Bengal Medical Services. Journal of the Asiatic Society of Bengal 16: 607-656, 897-952, 1026-1078." title="Catalogue of retiles inhabiting the Malayan Peninsula and islands, collected or observed by Theodore Cantor, Esq., M. D. Bengal Medical Services. Journal of the Asiatic Society of Bengal 16: 607 - 656, 897 - 952, 1026 - 1078." year="1847">Cantor 1847</bibRefCitation>
:624
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1856:449; 
<bibRefCitation author="Anderson, J" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="149 - 211" refId="B2" refString="Anderson, J, 1871. On some Indian reptiles. Proceedings of the Zoological Society of London 1871: 149 - 211" title="On some Indian reptiles." volume="1871" year="1871">Anderson 1871</bibRefCitation>
:150; 
<bibRefCitation author="Mueller, F" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="0" pageNumber="47" pagination="120 - 165" refId="B96" refString="Mueller, F, 1882. Erster Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museums. Verhandlungen der Naturforschenden Gesellschaft in Basel 7: 120 - 165" title="Erster Nachtrag zum Katalog der herpetologischen Sammlung des Basler Museums." volume="7" year="1882">
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:124
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: 
<bibRefCitation author="Guenther, A" journalOrPublisher="Sauria" pageId="0" pageNumber="47" refId="B57" refString="Guenther, A, 1864. The Reptiles of British India. The Royal Society of London, xxvii + 452 pp., 26 pls." title="The Reptiles of British India. The Royal Society of London, xxvii + 452 pp., 26 pls." year="1864">
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1864
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:103; 
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1889.tb06764.x" author="Boulenger, GA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="143 - 145" refId="B13" refString="Boulenger, GA, 1889. On some specimens of lizards in the Zoological Musuem of Halle (Saale). Proceedings of the Zoological Society of London 57: 143 - 145, DOI: https://doi.org/10.1111/j.1469-7998.1889.tb06764.x" title="On some specimens of lizards in the Zoological Musuem of Halle (Saale)." url="https://doi.org/10.1111/j.1469-7998.1889.tb06764.x" volume="57" year="1889">Boulenger 1889</bibRefCitation>
:143, 1890:103, 1912:51 (in part); 
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
1895:832 (in part); 
<bibRefCitation author="Flower, SS" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="47" pagination="856 - 914" refId="B32" refString="Flower, SS, 1896. Notes on a second collection of batrachians made in the Malay Peninsula 1895-96, with a list of the species recorded from that region. Proceedings of the Zoological Society of London 1896: 856 - 914" title="Notes on a second collection of batrachians made in the Malay Peninsula 1895 - 96, with a list of the species recorded from that region." volume="1896" year="1896">Flower 1896</bibRefCitation>
:867 (in part), 1899:634 (in part); Annandale 1906:92; 
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:57 (in part)
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authority="Brongniart, 1800" authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko Stentor" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="Stentor">Gecko Stentor</taxonomicName>
: 
<bibRefCitation author="Theobald, W" journalOrPublisher="The University of Kansas Science Bulletin" pageId="0" pageNumber="47" refId="B126" refString="Theobald, W, 1876. Descriptive Catalog of the Reptiles of British India. Thacker, Spink and Co., London." title="Descriptive Catalog of the Reptiles of British India. Thacker, Spink and Co., London." year="1876">Theobald 1876</bibRefCitation>
:72
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko stentor" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="stentor">Gekko stentor</taxonomicName>
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<bibRefCitation author="Brongersma, LD" journalOrPublisher="Zoologische Mededeelingen uitgegeven door het Rijksmuseum van Natuurlijke Historie" pageId="0" pageNumber="47" pagination="161 - 251" refId="B16" refString="Brongersma, LD, 1934. Contributions to the Indo-Australian herpetology. Zoologische Mededeelingen uitgegeven door het Rijksmuseum van Natuurlijke Historie 17: 161 - 251" title="Contributions to the Indo-Australian herpetology." volume="17" year="1934">Brongersma 1934</bibRefCitation>
:165
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithi" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithi">Gekko smithi</taxonomicName>
: 
<bibRefCitation author="Smith, MA" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B123" refString="Smith, MA, 1935. The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, Taylor &amp; Francis Ltd." title="The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, Taylor &amp; Francis Ltd." year="1935">Smith 1935</bibRefCitation>
:113 (in part); 
<bibRefCitation author="Taylor, EH" journalOrPublisher="The University of Kansas Science Bulletin" pageId="0" pageNumber="47" pagination="687 - 1077" refId="B125" refString="Taylor, EH, 1963. The Lizards of Thailand. The University of Kansas Science Bulletin 44: 687 - 1077" title="The Lizards of Thailand." volume="44" year="1963">Taylor 1963</bibRefCitation>
:803 (in part); 
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:477 (in part)
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authority="Brongniart, 1800" authorityName="Stoliczka" authorityYear="1870" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">Gecko smithii</taxonomicName>
: 
<bibRefCitation author="Mertens, R" journalOrPublisher="Oecologia" pageId="0" pageNumber="47" refId="B92" refString="Mertens, R, 1946.: Die Warn- und Drohreaktionen der Reptilien. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 471: 1108." title=": Die Warn- und Drohreaktionen der Reptilien. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 471: 1108." year="1946">Mertens 1946</bibRefCitation>
:16
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<bibRefCitation author="Biswas, S" journalOrPublisher="Records of the Zoological Survey of India" pageId="0" pageNumber="47" pagination="107 - 124" refId="B7" refString="Biswas, S, Sanyal, DP, 1977. Notes on the Reptilia collection from the Great Nicobar Island during the Great Nicobar Expedition in 1966. Records of the Zoological Survey of India 72: 107 - 124" title="Notes on the Reptilia collection from the Great Nicobar Island during the Great Nicobar Expedition in 1966." volume="72" year="1977">Biswas and Sanyal 1977</bibRefCitation>
:111 (in part)
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<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">Gekko smithii</taxonomicName>
: 
<bibRefCitation author="Wermuth, H" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="47" refId="B133" refString="Wermuth, H, 1965. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Berlin, Das Tierreich, 80, Walterv de Gruyter, XXII + 246 pp." title="Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Berlin, Das Tierreich, 80, Walterv de Gruyter, XXII + 246 pp." year="1965">Wermuth 1965</bibRefCitation>
:40 (in part); 
<bibRefCitation author="Kluge, AG" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="47" pagination="1 - 59" refId="B69" refString="Kluge, AG, 1967.: Higher taxonomic categories of gekkonid lizards and their evolution. Bulletin of the American Museum of Natural History 135: 1 - 59" title=": Higher taxonomic categories of gekkonid lizards and their evolution." volume="135" year="1967">Kluge 1967</bibRefCitation>
:135 (in part); 
<bibRefCitation author="Grossmann, W" journalOrPublisher="Sauria" pageId="0" pageNumber="47" pagination="9 - 18" refId="B56" refString="Grossmann, W, Ulber, T, 1990. Ein neuer Gecko aus Zentral-Thailand: Gekko siamensis sp. nov. (Reptilia: Sauria: Gekkoni-dae). Sauria 12: 9 - 18" title="Ein neuer Gecko aus Zentral-Thailand: Gekko siamensis sp. nov. (Reptilia: Sauria: Gekkoni-dae)." volume="12" year="1990">Grossmann and Ulber 1990</bibRefCitation>
:9; 
<bibRefCitation author="Kluge, AG" journalOrPublisher="Smithsonian Herpetological Information Service" pageId="0" pageNumber="47" pagination="1 - 35" refId="B70" refString="Kluge, AG, 1991. Checklist of gekkonoid lizards. Smithsonian Herpetological Information Service 85: 1 - 35" title="Checklist of gekkonoid lizards." volume="85" year="1991">Kluge 1991</bibRefCitation>
:10 (in part); 
<bibRefCitation author="Ota, H" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="47" pagination="147 - 151" refId="B103" refString="Ota, H, Hikida, T, Matsui, M, 1991. Re-evaluation of the status of Gecko verreauxi Tytler, 1864, from the Andaman Islands, India. Journal of Herpetology 25: 147 - 151" title="Re-evaluation of the status of Gecko verreauxi Tytler, 1864, from the Andaman Islands, India." volume="25" year="1991">Ota et al. 1991</bibRefCitation>
:147; 
<bibRefCitation author="Manthey, U" journalOrPublisher="Natur und Tier-Verlag, Muenster" pageId="0" pageNumber="47" refId="B87" refString="Manthey, U, Grossmann, W, 1997. Amphibien und Reptilien Suedostasiens. Natur und Tier-Verlag, Muenster" title="Amphibien und Reptilien Suedostasiens." year="1997">Manthey and Grossmann 1997</bibRefCitation>
:234 (in part); 
<bibRefCitation author="Grismer, LL" journalOrPublisher="Edition Chimaira, Frankfurt am Main" pageId="0" pageNumber="47" refId="B40" refString="Grismer, LL, 2011a. Field Guide to the Amphibians and Reptiles of the Seribuat Archipelago (Peninsular Malaysia). Edition Chimaira, Frankfurt am Main" title="Field Guide to the Amphibians and Reptiles of the Seribuat Archipelago (Peninsular Malaysia)." year="2011 a">Grismer 2011a</bibRefCitation>
:127 (in part), 2011b:469 (in part)
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">Gekko albofasciolatus</taxonomicName>
: 
<bibRefCitation author="Ota, H" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="47" pagination="147 - 151" refId="B103" refString="Ota, H, Hikida, T, Matsui, M, 1991. Re-evaluation of the status of Gecko verreauxi Tytler, 1864, from the Andaman Islands, India. Journal of Herpetology 25: 147 - 151" title="Re-evaluation of the status of Gecko verreauxi Tytler, 1864, from the Andaman Islands, India." volume="25" year="1991">Ota et al. 1991</bibRefCitation>
:150 (in part)
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko (Gekko) smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii" subGenus="Gekko">Gekko (Gekko) smithii</taxonomicName>
: 
<bibRefCitation author="Wood, PL" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="47" refId="B135" refString="Wood, PL, Guo, X, Travers, SL, Su, YC, Olson, KV, Bauer, AM, Grismer, LL, Siler, CD, Moyle, RG, Andersen, MJ, Brown, RF, 2020a. . Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Molecular Phylogenetics and Evolution 146: 106731." title=". Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Molecular Phylogenetics and Evolution 146: 106731." year="2020 a">Wood et al. 2020a</bibRefCitation>
:7 (in part)
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">Gekko cf. smithii</taxonomicName>
: 
<bibRefCitation author="Chandramouli, SR" journalOrPublisher="Amphibian &amp; Reptile Conservation" pageId="0" pageNumber="47" pagination="108 - 125" refId="B22" refString="Chandramouli, SR, Gokulakrishnan, G, Sivaperuman, C, Grismer, LL, 2021. A new species of the genus Gekko Laurenti, 1768 (Squamata: Gekkonidae) from the Nicobar Archipelago, with an overview of congeners from the Andaman and Nicobar Islands. Amphibian &amp; Reptile Conservation 15: 108 - 125" title="A new species of the genus Gekko Laurenti, 1768 (Squamata: Gekkonidae) from the Nicobar Archipelago, with an overview of congeners from the Andaman and Nicobar Islands." volume="15" year="2021">Chandramouli et al. 2021</bibRefCitation>
:108 (in part)
</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="genus">Gekko</taxonomicName>
Non-technical books, field guides, and pockets guides are not listed
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="diagnosis">
<paragraph pageId="0" pageNumber="47">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="0" pageNumber="47">sensu stricto</emphasis>
herein after referred to as 
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
</taxonomicName>
unless noted otherwise can be separated from all other species of 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko</emphasis>
</taxonomicName>
in the 
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
</taxonomicName>
species complex by having the combination of a maximum SVL of 191.1 mm, 11-17 supralabials, 9-14 infralabials, 3-6 internarial scales, 19-26 frontal scales, 4-9 chin scales, 94-137 midbody scales, 17-23 paravertebral tubercles, 8-11 longitudinal rows of tubercles, 23-35 ventral scales, 15-20 1st toe subdigital lamellae, 19-24 4th toe lamellae; 13-15 precloacal pores in males (absent in females); enlarged subcaudal scales; thin, white nuchal band at base of occiput composed of closely spaced spots; thin dark nuchal band contacting the eyes; large white ocelli surrounding dorsal tubercles or bordering them posteriorly in six or seven transverse rows; and a thick dark reticulum to diffuse banded dorsal pattern (Tables 
<tableCitation captionStart="Table 9" captionStartId="T9" captionText="Table 9. Summary statistics of the meristic data. sd = 1 standard deviation. N = sample size. Character abbreviations are in the Materials and methods." httpUri="http://table.plazi.org/id/A165CC91E2126A36916C509BD46E86AC" pageId="0" pageNumber="47" tableUuid="A165CC91E2126A36916C509BD46E86AC">9</tableCitation>
, 
<tableCitation captionStart="Table 10" captionStartId="T10" captionText="Table 10. Diagnostic character states (shaded cells) separating Gekko hulk sp. nov. from the other species of G. (Gekko). Data on G. gecko, G. nutaphandi, G. reevesi, G. siamensis, G. stoliczkai, and G. verreauxi come from Grossmann and Ulber (1990), Ota and Nabhitabhata (1991), Bauer et al. (2008), Rosler et al. (2011), and Chandramouil et al. (2021)." httpUri="http://table.plazi.org/id/2FCA3F971C5DFDE7C899FA2A5FBD60B3" pageId="0" pageNumber="47" tableUuid="2FCA3F971C5DFDE7C899FA2A5FBD60B3">10</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="2FCA3F971C5DFDE7C899FA2A5FBD60B3" httpUri="http://table.plazi.org/id/2FCA3F971C5DFDE7C899FA2A5FBD60B3" pageId="0" pageNumber="47" start="Table 10" startId="T10">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Table 10.</emphasis>
Diagnostic character states (shaded cells) separating 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko hulk" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov.</emphasis>
from the other species of 
<taxonomicName authority="" genus="G." lsidName="G. (Gekko)" pageId="0" pageNumber="47" rank="subGenus" subGenus="Gekko">G. (Gekko)</taxonomicName>
. Data on 
<taxonomicName genus="G." lsidName="G. gecko" pageId="0" pageNumber="47" rank="species" species="gecko">
<emphasis italics="true" pageId="0" pageNumber="47">G. gecko</emphasis>
</taxonomicName>
, 
<taxonomicName genus="G." lsidName="G. nutaphandi" pageId="0" pageNumber="47" rank="species" species="nutaphandi">
<emphasis italics="true" pageId="0" pageNumber="47">G. nutaphandi</emphasis>
</taxonomicName>
, 
<taxonomicName genus="G." lsidName="G. reevesi" pageId="0" pageNumber="47" rank="species" species="reevesi">
<emphasis italics="true" pageId="0" pageNumber="47">G. reevesi</emphasis>
</taxonomicName>
, 
<taxonomicName genus="G." lsidName="G. siamensis" pageId="0" pageNumber="47" rank="species" species="siamensis">
<emphasis italics="true" pageId="0" pageNumber="47">G. siamensis</emphasis>
</taxonomicName>
, 
<taxonomicName genus="G." lsidName="G. stoliczkai" pageId="0" pageNumber="47" rank="species" species="stoliczkai">
<emphasis italics="true" pageId="0" pageNumber="47">G. stoliczkai</emphasis>
</taxonomicName>
, and 
<taxonomicName genus="G." lsidName="G. verreauxi" pageId="0" pageNumber="47" rank="species" species="verreauxi">
<emphasis italics="true" pageId="0" pageNumber="47">G. verreauxi</emphasis>
</taxonomicName>
come from 
<bibRefCitation author="Grossmann, W" journalOrPublisher="Sauria" pageId="0" pageNumber="47" pagination="9 - 18" refId="B56" refString="Grossmann, W, Ulber, T, 1990. Ein neuer Gecko aus Zentral-Thailand: Gekko siamensis sp. nov. (Reptilia: Sauria: Gekkoni-dae). Sauria 12: 9 - 18" title="Ein neuer Gecko aus Zentral-Thailand: Gekko siamensis sp. nov. (Reptilia: Sauria: Gekkoni-dae)." volume="12" year="1990">Grossmann and Ulber (1990)</bibRefCitation>
, 
<bibRefCitation author="Ota, H" journalOrPublisher="Copeia" pageId="0" pageNumber="47" pagination="503 - 509" refId="B104" refString="Ota, H, Nabhitabhata, J, 1991. A new species of Gekko (Gekkonidae: Squamata) from Thailand. Copeia 1991: 503 - 509" title="A new species of Gekko (Gekkonidae: Squamata) from Thailand." volume="1991" year="1991">Ota and Nabhitabhata (1991)</bibRefCitation>
, 
<bibRefCitation author="Bauer, AM" journalOrPublisher="Zootaxa" pageId="0" pageNumber="47" pagination="32 - 42" refId="B6" refString="Bauer, AM, Sumontha, M, Pauwels, OSG, 2008. A new red-eyed Gekko (Reptilia: Gekkonidae) from Kanchanaburi Province, Thailand. Zootaxa 1750: 32 - 42" title="A new red-eyed Gekko (Reptilia: Gekkonidae) from Kanchanaburi Province, Thailand." volume="1750" year="2008">Bauer et al. (2008)</bibRefCitation>
, Rosler et al. (2011), and Chandramouil et al. (2021).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="47">
<table pageId="0" pageNumber="47">
<tr pageId="0" pageNumber="47">
<td colspan="1" isEmpty="true" pageId="0" pageNumber="47" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName baseAuthorityName="Guenther" baseAuthorityYear="1867" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko albofasciolatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="albofasciolatus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gekko albofasciolatus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko alblomaculatus" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="alblomaculatus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gekko alblomaculatus</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko gecko" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="gecko">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gekko gecko</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="47">
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko hulk" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko hulk</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName authorityName="Bauer, Sumontha &amp; Pauwels" authorityYear="2008" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko nutaphandi" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="nutaphandi">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gekko nutaphandi</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko reevesi" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="reevesi">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gekko reevesi</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName authorityName="Grossmann &amp; Ulber" authorityYear="1990" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko siamensis" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="siamensis">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gekko siamensis</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko stoliczkai" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="stoliczkai">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gekko stoliczkai</emphasis>
</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">
<taxonomicName authority="Brongniart, 1800" authorityName="Brongniart" authorityYear="1800" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko verreauxi" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="verreauxi">
<emphasis bold="true" italics="true" pageId="0" pageNumber="47">Gecko verreauxi</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">max SVL</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">165.1</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">157.4</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">185</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">161.3</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">117</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">173</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">150</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">191</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">128.4</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">155</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">precloacal pores (males)</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">13-15</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">11-17</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">12-16</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">6-13</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">17-22</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">13-20</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">10-13</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">13-15</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">13-15</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">11-13</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">rostral scale contacting nares</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">occiput scales smaller than snout scales</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">longitudinal rows of tubercles</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9-12</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9-11</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9-18</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">9-11</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">14</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">12-18</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">16-19</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">8-11</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">10-12</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">10-14</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">subcaudals enlarged</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">iris color</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">green</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">green</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">gold/copper/ olive/brown</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">turquoise/green</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">red</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">gold/copper/ olive/brown</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">green</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">turquoise/green</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">green</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">golden</td>
</tr>
<tr pageId="0" pageNumber="47">
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">solid, thin, white nuchal band</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">yes</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
<td colspan="1" pageId="0" pageNumber="47" rowspan="1">no</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="distribution">
<paragraph pageId="0" pageNumber="47">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authority="Brongniart, 1800" authorityName="Stoliczka" authorityYear="1870" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gecko smithii</emphasis>
</taxonomicName>
ranges from southern Thailand south of the Isthmus of Kra from at least Khao Phanom Bencha National Park in Krabi and Khao Nan National Park in Nakhon Si Thammarat Province to the northern border of the Banjaran Titiwangsa in southeastern Thailand and northwestern Peninsular Malaysia. Its range continues southward along the west side of the Banjaran Titiwangsa to at least the state of Selangor but very likely farther as well (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the species of the Gekko smithii complex from throughout their respective Sundaic land masses. Circles denote the localities of specimens that were both examined first hand and are represented in the phylogeny (Fig. 2). Stars denote type localities. Squares denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature. Localities from non-peer reviewed literature or unverifiable online data were not included." figureDoi="10.3897/vz.72.e77702.figure1" httpUri="https://binary.pensoft.net/fig/641477" pageId="0" pageNumber="47">1</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution and color pattern variation in Gekko smithii and G. hulk sp. nov. on the Thai-Malay Peninsula. A. Gekko smithii (LSUHC 15052) from Perlis State Park, Perlis, Peninsular Malaysia. B. Gekko smithii (LSUDPC 12884) from Puncak Janing, Kedah, Peninsualr Malaysia. C. Gekko smithii (LSUHC 9626) from Sedim, Kedah, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12869) from Penang Hill (type locality), Penang, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12886) from Lata Kedondong, Selangor, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12887) from Kota Damansara, Selangor, Peninsular Malaysia. G. Gekko smithii (LSUDPC 11174) from Khaopu Khaoya, Nakhon Si Thamarat Province, Thailand. H. Gekko smithii (LSUDPC 11173) from Bacho Falls, Narathiwat Province, Thailand. I. Gekko smithii (LSUDPC 12878) from the Weang District, Narathiwat Province, Thailand. J. Gekko hulk sp. nov. (LSUDPC 12880) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 8696) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUDPC 7991) from Gunung Tebu, Terengganu, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12897) from Merapoh, Pahang, Peninsular Malaysia. N. Gekko hulk sp. nov. (LSUDPC 12897) from Pulau Tioman, Pahang, Peninsular Malaysia. Photographs by L. Lee Grismer (A, K, L, N), Evan S. H. Quah (B, C, D), Kurt H. P. Guek (E), Steven Wong (F), Henrick Bringsoe (G, H); Parinya Pawangkhanant (I), Ian Dugdale (J), and Nick Baker (M). Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature." figureDoi="10.3897/vz.72.e77702.figure7" httpUri="https://binary.pensoft.net/fig/641483" pageId="0" pageNumber="47">7</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="47">
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B124" refString="Stoliczka, F, 1870. Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39(2): 134-228, pls. IX-XII." title="Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39 (2): 134 - 228, pls. IX-XII." year="1870">Stoliczka (1870)</bibRefCitation>
reports &quot; 
<taxonomicName authority="Brongniart, 1800" authorityName="Stoliczka" authorityYear="1870" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko Smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="Smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gecko Smithii</emphasis>
</taxonomicName>
&quot; from Java based on a juvenile specimen (SVL 86.3 mm) specimen sent to him but not collected by him. However, his description of the color pattern is well within the range of variation of both 
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
</taxonomicName>
and 
<taxonomicName genus="G." lsidName="G. gecko" pageId="0" pageNumber="47" rank="species" species="gecko">
<emphasis italics="true" pageId="0" pageNumber="47">G. gecko</emphasis>
</taxonomicName>
. 
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B124" refString="Stoliczka, F, 1870. Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39(2): 134-228, pls. IX-XII." title="Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39 (2): 134 - 228, pls. IX-XII." year="1870">Stoliczka (1870)</bibRefCitation>
stated his specimen had 12 longitudinal rows of &quot;small flattened sub-equal granules [=longitudinal rows of dorsal tubercles], slightly varying in size on the posterior part of the body and especially at the sides&quot;. The dorsal tubercles of species in the 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
complex are distinctly raised and sub-conical in adults but less so in juveniles. Furthermore, of the 93 specimens of the 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
complex examined here, only one specimen from Borneo had 12 rows dorsal tubercles. The others ranged from 8-11. Awal Riyanto (pers com. in lit. 2021) says he nor any of his colleagues working in Java have ever seen 
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
</taxonomicName>
. Photographic material and specimens we have examined from Java cataloged as 
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
</taxonomicName>
in the Zoological Museum Amsterdam, now officially part of Naturalis Biodiversity Center in Leiden, are 
<taxonomicName genus="G." lsidName="G. gecko" pageId="0" pageNumber="47" rank="species" species="gecko">
<emphasis italics="true" pageId="0" pageNumber="47">G. gecko</emphasis>
</taxonomicName>
. We believe reports of 
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
</taxonomicName>
from Java stem from the possible misidentification of this species by 
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B124" refString="Stoliczka, F, 1870. Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39(2): 134-228, pls. IX-XII." title="Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39 (2): 134 - 228, pls. IX-XII." year="1870">Stoliczka (1870)</bibRefCitation>
as no new naturally occurring populations have been reported or observed to our knowledge. The less likely scenario exists, however, that Stoliczka described a juvenile 
<taxonomicName genus="G." lsidName="G. albomaculatus" pageId="0" pageNumber="47" rank="species" species="albomaculatus">
<emphasis italics="true" pageId="0" pageNumber="47">G. albomaculatus</emphasis>
</taxonomicName>
from Bangka island, Indonesia as he notes &quot;I have also added a complete description of the rare 
<taxonomicName authority="Brongniart, 1800" authorityName="Stoliczka" authorityYear="1870" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gecko smithii</emphasis>
</taxonomicName>
, Gray, a specimen of which was sent to me from Java, and that of what appears to be a full grown specimen of 
<taxonomicName authorityName="Gunther" authorityYear="1858" class="Reptilia" family="Colubridae" genus="Tetragonosoma" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tetragonosoma" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Tetragonosoma</emphasis>
</taxonomicName>
[
<taxonomicName class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" lsidName="Lycodon" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Lycodon</emphasis>
</taxonomicName>
] 
<emphasis italics="true" pageId="0" pageNumber="47">effrene</emphasis>
, CANT., from the island Banca.&quot; The ambiguity in this sentence could mean these two species were sent to him from Java but were collected on Bangka Island.
</paragraph>
<paragraph pageId="0" pageNumber="47">
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B124" refString="Stoliczka, F, 1870. Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39(2): 134-228, pls. IX-XII." title="Observations on some Indian and Malayan Amphibia and Reptilia. Journal of the Asiatic Society of Bengal 39 (2): 134 - 228, pls. IX-XII." year="1870">Stoliczka (1870)</bibRefCitation>
reported 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(as 
<taxonomicName authority="Brongniart, 1800" authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko stentor" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="stentor">
<emphasis italics="true" pageId="0" pageNumber="47">Gecko stentor</emphasis>
</taxonomicName>
) from the coastal cities of Chittagong, Bangladesh and Akyab (= Sittwe = Sittway, Rakhine [Arakan] State), Myanmar which was followed by 
<bibRefCitation author="Theobald, W" journalOrPublisher="The University of Kansas Science Bulletin" pageId="0" pageNumber="47" refId="B126" refString="Theobald, W, 1876. Descriptive Catalog of the Reptiles of British India. Thacker, Spink and Co., London." title="Descriptive Catalog of the Reptiles of British India. Thacker, Spink and Co., London." year="1876">Theobald (1876)</bibRefCitation>
, Annandale (1906), 
<bibRefCitation author="Boulenger, GA" journalOrPublisher="London, Taylor &amp; Francis" pageId="0" pageNumber="47" refId="B15" refString="Boulenger, GA, 1912. A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. Reptilia and Batrachia. London, Taylor &amp; Francis" title="A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. Reptilia and Batrachia." year="1912">Boulenger (1912)</bibRefCitation>
, and 
<bibRefCitation author="Taylor, EH" journalOrPublisher="The University of Kansas Science Bulletin" pageId="0" pageNumber="47" pagination="687 - 1077" refId="B125" refString="Taylor, EH, 1963. The Lizards of Thailand. The University of Kansas Science Bulletin 44: 687 - 1077" title="The Lizards of Thailand." volume="44" year="1963">Taylor (1963)</bibRefCitation>
. 
<bibRefCitation author="Smith, MA" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="47" refId="B123" refString="Smith, MA, 1935. The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, Taylor &amp; Francis Ltd." title="The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, Taylor &amp; Francis Ltd." year="1935">Smith (1935)</bibRefCitation>
stated &quot;I do not know of any specimens to prove that this 
<taxonomicName authority="Brongniart, 1800" authorityName="Brongniart" authorityYear="1800" class="Reptilia" family="Gekkonidae" genus="Gecko" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gecko" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="genus">Gecko</taxonomicName>
inhabits Burma, as has been stated.&quot; 
<bibRefCitation author="Ota, H" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="47" pagination="147 - 151" refId="B103" refString="Ota, H, Hikida, T, Matsui, M, 1991. Re-evaluation of the status of Gecko verreauxi Tytler, 1864, from the Andaman Islands, India. Journal of Herpetology 25: 147 - 151" title="Re-evaluation of the status of Gecko verreauxi Tytler, 1864, from the Andaman Islands, India." volume="25" year="1991">Ota et al. (1991)</bibRefCitation>
examined a specimen (MCZ 3120) putatively from &quot;Burma: Rangoon&quot; (=Myanmar, Yangon). MCZ has no other data on this specimen and the only species of 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko</emphasis>
</taxonomicName>
we have seen in Yangon and throughout all of Myanmar over the last five years of field work has been 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko gecko" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="gecko">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko gecko</emphasis>
</taxonomicName>
. Furthermore, the herpetological surveys in Myanmar by the California Academy of Sciences (www.calacademy.org/research/herpetology/myanmar) which resulted in the collection of over 14,000 specimens, found no 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
. We do not consider this species as part of the Burmese herpetofauna. However, the type material of 
<taxonomicName authorityName="Mertens" authorityYear="1955" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko gecko subsp. azhari" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="subSpecies" species="gecko" subSpecies="azhari">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko gecko azhari</emphasis>
</taxonomicName>
Mertens, 1955 from Barkal, Chittagong Hill Tracts, Bangladesh does bear some resemblance to 
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
</taxonomicName>
in head shape, gular scales, and dorsal tubercle shape (
<bibRefCitation author="Roesler, H" journalOrPublisher="Gekkota" pageId="0" pageNumber="47" pagination="33 - 46" refId="B116" refString="Roesler, H, 2001. Studien am Tokeh: 1. Gekko gecko azhari Mertens, 1825 (Sauria: Gekkonidae). Gekkota 3: 33 - 46" title="Studien am Tokeh: 1. Gekko gecko azhari Mertens, 1825 (Sauria: Gekkonidae)." volume="3" year="2001">
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
2001
</bibRefCitation>
). 
<bibRefCitation author="Mahony, s" journalOrPublisher="Hamadryad" pageId="0" pageNumber="47" pagination="80 - 94" refId="B85" refString="Mahony, s, Hasan, MK, Kabir, MM, Ahmed, M, Hossain, MK, 2009. A catalogue of amphibinas and reptiles in the collection of Jahangirnagar University, Dhaka, Bangladesh. Hamadryad 34: 80 - 94" title="A catalogue of amphibinas and reptiles in the collection of Jahangirnagar University, Dhaka, Bangladesh." volume="34" year="2009">Mahony et al. (2009)</bibRefCitation>
reported on 23 additional specimens from throughout the Chittagong Division that they considered 
<taxonomicName genus="G." lsidName="G. g. subsp. azhari" pageId="0" pageNumber="47" rank="subSpecies" species="g." subSpecies="azhari">
<emphasis italics="true" pageId="0" pageNumber="47">G. g. azhari</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="variation">
<paragraph pageId="0" pageNumber="47">Variation in color pattern.</paragraph>
<paragraph pageId="0" pageNumber="47">
The dorsal body tubercles of 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
from northernmost Peninsular Malaysia and Thailand north of the Kangar-Pattani Line (generally running from Gunung Jerai, Kedah, Peninsular Malaysia to Songkhla, Songkhla, Thailand) are surrounded by a large white ocellus whereas in southern populations, the ocelli are smaller and may only border the tubercles posteriorly (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution and color pattern variation in Gekko smithii and G. hulk sp. nov. on the Thai-Malay Peninsula. A. Gekko smithii (LSUHC 15052) from Perlis State Park, Perlis, Peninsular Malaysia. B. Gekko smithii (LSUDPC 12884) from Puncak Janing, Kedah, Peninsualr Malaysia. C. Gekko smithii (LSUHC 9626) from Sedim, Kedah, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12869) from Penang Hill (type locality), Penang, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12886) from Lata Kedondong, Selangor, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12887) from Kota Damansara, Selangor, Peninsular Malaysia. G. Gekko smithii (LSUDPC 11174) from Khaopu Khaoya, Nakhon Si Thamarat Province, Thailand. H. Gekko smithii (LSUDPC 11173) from Bacho Falls, Narathiwat Province, Thailand. I. Gekko smithii (LSUDPC 12878) from the Weang District, Narathiwat Province, Thailand. J. Gekko hulk sp. nov. (LSUDPC 12880) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 8696) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUDPC 7991) from Gunung Tebu, Terengganu, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12897) from Merapoh, Pahang, Peninsular Malaysia. N. Gekko hulk sp. nov. (LSUDPC 12897) from Pulau Tioman, Pahang, Peninsular Malaysia. Photographs by L. Lee Grismer (A, K, L, N), Evan S. H. Quah (B, C, D), Kurt H. P. Guek (E), Steven Wong (F), Henrick Bringsoe (G, H); Parinya Pawangkhanant (I), Ian Dugdale (J), and Nick Baker (M). Unannotated circles denote localities of specimens or photographs examined here or verified from other publications as well as vouchered samples in the literature." figureDoi="10.3897/vz.72.e77702.figure7" httpUri="https://binary.pensoft.net/fig/641483" pageId="0" pageNumber="47">7</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">8</figureCitation>
). Although the latter creates a general resemblance to 
<taxonomicName genus="G." lsidName="G. hulk" pageId="0" pageNumber="47" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">G. hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov.</emphasis>
, the small white spots in 
<taxonomicName genus="G." lsidName="G. hulk" pageId="0" pageNumber="47" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">G. hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov.</emphasis>
, when not confined to the tubercles, tend to border them anteriorly. Although not as adept at substrate matching as 
<taxonomicName genus="G." lsidName="G. gecko" pageId="0" pageNumber="47" rank="species" species="gecko">
<emphasis italics="true" pageId="0" pageNumber="47">G. gecko</emphasis>
</taxonomicName>
(see figs 374-378 in 
<bibRefCitation author="Grismer, LL" journalOrPublisher="Edition Chimaira, Frankfurt am Main" pageId="0" pageNumber="47" refId="B40" refString="Grismer, LL, 2011a. Field Guide to the Amphibians and Reptiles of the Seribuat Archipelago (Peninsular Malaysia). Edition Chimaira, Frankfurt am Main" title="Field Guide to the Amphibians and Reptiles of the Seribuat Archipelago (Peninsular Malaysia)." year="2011 a">Grismer 2011a</bibRefCitation>
), considerable variation has been observed wherein some individuals of 
<taxonomicName genus="G." lsidName="G. smithii" pageId="0" pageNumber="47" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">G. smithii</emphasis>
</taxonomicName>
from Penang and Selangor states have very faded dark dorsal markings (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">8F</figureCitation>
). Although rare, other geckos from Selangor state may resemble east peninsular lineage geckos in lacking a dark nuchal loop and having a darkly speckled dorsum (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">8C</figureCitation>
). Iris color can also range from turquoise to lime green in the same population (Figs 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">8A</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">8D</figureCitation>
) and along with ground color (which can vary from yellow to dark gray (Figs 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">8B</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic variation in Gekko smithii (A-G) and G. hulk sp. nov. (H-M). A. Gekko smithii (LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. Gekko smithii (LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. Gekko smithii (LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. Gekko smithii (LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. Gekko smithii (LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. Gekko smithii (LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. Gekko smithii (THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. Gekko hulk sp. nov. (LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. Gekko hulk sp. nov. (LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. Gekko hulk sp. nov. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. Gekko hulk sp. nov. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. Gekko hulk sp. nov. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. Gekko hulk sp. nov. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M)." figureDoi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47">8E</figureCitation>
)), is not a diagnostic character (contra 
<bibRefCitation author="Grossmann, W" journalOrPublisher="Natur und Tier Verlag GmbH, Muenster" pageId="0" pageNumber="47" refId="B53" refString="Grossmann, W, 2006. Gruenaugengeckos Gekko smithii &amp; Gekko siamensis. Natur und Tier Verlag GmbH, Muenster" title="Gruenaugengeckos Gekko smithii &amp; Gekko siamensis." year="2006">Grossmann 2006</bibRefCitation>
). See Tables 
<tableCitation captionStart="Table 9" captionStartId="T9" captionText="Table 9. Summary statistics of the meristic data. sd = 1 standard deviation. N = sample size. Character abbreviations are in the Materials and methods." httpUri="http://table.plazi.org/id/A165CC91E2126A36916C509BD46E86AC" pageId="0" pageNumber="47" tableUuid="A165CC91E2126A36916C509BD46E86AC">9</tableCitation>
and 
<tableCitation captionStart="Table 10" captionStartId="T10" captionText="Table 10. Diagnostic character states (shaded cells) separating Gekko hulk sp. nov. from the other species of G. (Gekko). Data on G. gecko, G. nutaphandi, G. reevesi, G. siamensis, G. stoliczkai, and G. verreauxi come from Grossmann and Ulber (1990), Ota and Nabhitabhata (1991), Bauer et al. (2008), Rosler et al. (2011), and Chandramouil et al. (2021)." httpUri="http://table.plazi.org/id/2FCA3F971C5DFDE7C899FA2A5FBD60B3" pageId="0" pageNumber="47" tableUuid="2FCA3F971C5DFDE7C899FA2A5FBD60B3">10</tableCitation>
for variation in morphology.
</paragraph>
<caption doi="10.3897/vz.72.e77702.figure8" httpUri="https://binary.pensoft.net/fig/641484" pageId="0" pageNumber="47" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="47">
<emphasis bold="true" pageId="0" pageNumber="47">Figure 8.</emphasis>
Geographic variation in 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(A-G) and 
<taxonomicName genus="G." lsidName="G. hulk" pageId="0" pageNumber="47" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">G. hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov</emphasis>
. (H-M). A. 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(LSUDPC 12899) from Ulu Yam, Selangor, Peninsular Malaysia. B. 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(LSUHC 4959) from Pulau Pangkor, Perak, Peninsular Malaysia. C. 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(LSUDPC 12898) from Ulu Yam, Selangor, Peninsular Malaysia. D. 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(LSUDPC 12895) from Kota Damansara, Selangor, Peninsular Malaysia. E. 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(LSUDPC 12903) from Bukit Panchor, Penang, Peninsular Malaysia. F. 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(LSUDPC 12902) from Ulu Yam, Selangor, Peninsular Malaysia. G. 
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
(THNHM 01844) from Phu Pha Phet, Satun, Province, Thailand. H. 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko hulk" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov.</emphasis>
(LSUDPC 12912) from Lata Kekabu, Setiu, Terengganu, Peninsular Malaysia. I. 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko hulk" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov.</emphasis>
(LSUDPC 8696) from Endau-Rompin, Johor, Peninsular Malaysia. J. 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko hulk" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov</emphasis>
. (LSUDPC 12953) from the Hala-Bala Wildlife Sanctuary, Narathiwat Province, Thailand. K. 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko hulk" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov</emphasis>
. (LSUHC 5152) from the Tekek-Juara Trail, Pulau Tioman, Pahang, Peninsular Malaysia. L. 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko hulk" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov</emphasis>
. (LSUHC 5873) from Kota Tinggi, Peninsular Malaysia. M. 
<taxonomicName authorityName="Grismer &amp; del Pinto &amp; Quah &amp; Anuar &amp; Cota &amp; McGuire &amp; Iskandar &amp; Wood Jr &amp; Grismer" authorityYear="2022" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko hulk" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="hulk">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko hulk</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="47">sp. nov</emphasis>
. (LSUDPC 12955) from Hutan Lipur Sekayu, Terengganu, Peninsular Malaysia. Photographs by Kurt H. P. Guek (A, C, D, F), L. Lee Grismer (B, K, L), Evan S. H. Quah (E, H), Michael Cota (G, J), M. A. Muin (I) and Syed A. Rizal (M).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="47" type="natural history">
<paragraph pageId="0" pageNumber="47">Natural history (adapted from Grismer 2011a,b).</paragraph>
<paragraph pageId="0" pageNumber="47">
<taxonomicName authorityName="Gray" authorityYear="1842" class="Reptilia" family="Gekkonidae" genus="Gekko" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gekko smithii" order="Squamata" pageId="0" pageNumber="47" phylum="Chordata" rank="species" species="smithii">
<emphasis italics="true" pageId="0" pageNumber="47">Gekko smithii</emphasis>
</taxonomicName>
is an arboreal nocturnal species that is well-established in all types of primary and secondary forests. It is particularly well-suited for inhabiting occupied and abandoned human dwellings along forest edges where it is most easily observed. In forested habitats, geckos occur on the trunks of large trees and boulders from where males are commonly heard calling during the day and early in the evening.
</paragraph>
</subSubSection>
</treatment>
</document>