treatments-xml/data/AC/C6/01/ACC601C874745647AAF5598883342EE4.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1086.77408" ID-GBIF-Dataset="dd81d919-f6d9-4fda-bbd1-7c45de307fc2" ID-PMC="PMC8863766" ID-Pensoft-Pub="1313-2970-1086-33" ID-Pensoft-UUID="43AFBD7D61105F598ADA74076D751C11" ID-PubMed="35210912" ID-ZooBank="6C32148CA0144AC488EA82A1509E44F8" ModsDocID="1313-2970-1086-33" checkinTime="1644966834654" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Meng, Kaibaryer &amp; Fekete, Judit" docDate="2022" docId="ACC601C874745647AAF5598883342EE4" docLanguage="en" docName="ZooKeys 1086: 33-76" docOrigin="ZooKeys 1086" docPubDate="2022-02-15" docSource="http://dx.doi.org/10.3897/zookeys.1086.77408" docTitle="Laeocathaica carinalis Chen &amp; Zhang 2004" docType="treatment" docVersion="5" id="43AFBD7D61105F598ADA74076D751C11" lastPageNumber="33" masterDocId="43AFBD7D61105F598ADA74076D751C11" masterDocTitle="A review of the genus Laeocathaica Moellendorff, 1899 (Gastropoda, Pulmonata, Camaenidae)" masterLastPageNumber="76" masterPageNumber="33" pageNumber="33" updateTime="1668151486723" updateUser="ExternalLinkService">
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<mods:title>A review of the genus Laeocathaica Moellendorff, 1899 (Gastropoda, Pulmonata, Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022, Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., Budapest 1148, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Meng, Kaibaryer</mods:namePart>
<mods:affiliation>National Zoological Museum of China, Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, H- 8200 Veszprem, Egyetem u. 10, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1086</mods:number>
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<mods:start>33</mods:start>
<mods:end>76</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1086.77408</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1086.77408</mods:identifier>
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<treatment ID-GBIF-Taxon="193377842" LSID="urn:lsid:plazi:treatment:ACC601C874745647AAF5598883342EE4" httpUri="http://treatment.plazi.org/id/ACC601C874745647AAF5598883342EE4" lastPageNumber="33" pageId="0" pageNumber="33">
<subSubSection pageId="0" pageNumber="33" type="nomenclature">
<paragraph pageId="0" pageNumber="33">
<taxonomicName LSID="ACC601C8-7474-5647-AAF5-598883342EE4" authority="Chen &amp; Zhang, 2004" authorityName="Chen &amp; Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica carinalis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="carinalis">Laeocathaica carinalis Chen &amp; Zhang, 2004</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="description">
<paragraph pageId="0" pageNumber="33">
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Laeocathaica carinalis Chen &amp; Zhang, 2004 (IZCAS TM 097578, holotype). Scale bar: 10 mm. Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.1086.77408.figure9" httpUri="https://binary.pensoft.net/fig/646384" pageId="0" pageNumber="33">Figures 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Laeocathaica carinalis Chen &amp; Zhang, 2004 A, B 2016 / 70 b C, D 2016 / 81 E 2016 / 82 F 2016 / 83. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure10" httpUri="https://binary.pensoft.net/fig/646385" pageId="0" pageNumber="33">, 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="reference_group">
<paragraph pageId="0" pageNumber="33">
<taxonomicName authorityName="Chen &amp; Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica carinalis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="carinalis">Laeocathaica carinalis</taxonomicName>
Chen &amp; Zhang, 2004: 341 [Chinese description], fig. 334 (erroneous! Shows a juvenile 
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="genus">Laeocathaica</taxonomicName>
shell belonging to another species); 444 [English description].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="type locality">
<paragraph pageId="0" pageNumber="33">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="33">
&quot;Town of Wenxian County, (
<geoCoordinate degrees="33" direction="north" minutes="0" orientation="latitude" precision="925" value="33.0">33°0'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="6" orientation="longitude" precision="925" value="104.1">104°6'E</geoCoordinate>
), Gansu Province, China&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="type material">
<paragraph pageId="0" pageNumber="33">Type material.</paragraph>
<paragraph pageId="0" pageNumber="33">
The shell we examined (IZCAS TM 097578) is exactly the same as the one figured by 
<bibRefCitation author="Chen, D-N" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="33" refId="B4" refString="Chen, D-N, Zhang, G-Q, 2004. Fauna Sinica Invertebrata Vol. 37 (Mollusca: Gastropoda: Stylommatophora: Bradybaenidae). Science Press, Beijing" title="Fauna Sinica Invertebrata Vol. 37 (Mollusca: Gastropoda: Stylommatophora: Bradybaenidae)." year="2004">Chen and Zhang (2004</bibRefCitation>
: fig. 219) as the holotype of 
<taxonomicName authorityName="Chen &amp; Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Cathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cathaica bizonalis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="bizonalis">
<emphasis italics="true" pageId="0" pageNumber="33">Cathaica bizonalis</emphasis>
</taxonomicName>
Chen &amp; Zhang, 2004. Therefore, we understand this situation as a confusion of specimens and photographs before publication, and consider the figured shell (IZCAS TM 097578, Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Laeocathaica carinalis Chen &amp; Zhang, 2004 (IZCAS TM 097578, holotype). Scale bar: 10 mm. Photographs: Kaibaryer Meng." figureDoi="10.3897/zookeys.1086.77408.figure9" httpUri="https://binary.pensoft.net/fig/646384" pageId="0" pageNumber="33">9</figureCitation>
) as the holotype. See also under 
<taxonomicName authorityName="Chen &amp; Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Cathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cathaica bizonalis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="bizonalis">
<emphasis italics="true" pageId="0" pageNumber="33">Cathaica bizonalis</emphasis>
</taxonomicName>
Chen &amp; Zhang, 2004.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="materials_examined">
<paragraph pageId="0" pageNumber="33">New material.</paragraph>
<paragraph pageId="0" pageNumber="33">
<materialsCitation collectingDate="2016-05-28" collectingDateMax="2016-05-31" collectingDateMin="2016-05-28" collectorName="A. Hunyadi" country="China" county="Longnan Shi" elevation="1800" latitude="32.941185" location="Jianshan Xiang" longLatPrecision="1" longitude="104.67298" municipality="Wenxian" specimenCount="1" stateProvince="Gansu">
<collectingCountry name="China">China</collectingCountry>
• 4 shells; 
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, 
<collectingCounty>Longnan Shi</collectingCounty>
, 
<collectingMunicipality>Wenxian</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:ACC601C874745647AAF5598883342EE4:26EEA9781612E89CC06BB4E603A47721" country="China" county="Longnan Shi" latitude="32.941185" longLatPrecision="1" longitude="104.67298" municipality="Wenxian" name="Jianshan Xiang" stateProvince="Gansu">Jianshan Xiang</location>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</elevation>
</quantity>
west of 
<location LSID="urn:lsid:plazi:treatment:ACC601C874745647AAF5598883342EE4:427C3A59E5E866433A6165B7CD37DBB9" country="China" county="Longnan Shi" latitude="32.941185" longLatPrecision="1" longitude="104.67298" municipality="Wenxian" name="Jianshan" stateProvince="Gansu">Jianshan</location>
towards 
<location LSID="urn:lsid:plazi:treatment:ACC601C874745647AAF5598883342EE4:174C102CC52AE51D4117F75EEE48F1B3" country="China" county="Longnan Shi" latitude="32.941185" longLatPrecision="1" longitude="104.67298" municipality="Wenxian" name="Diaohuya" stateProvince="Gansu">Diaohuya</location>
, right side of road no. 212 (locality code: 2016/70b); 
<geoCoordinate degrees="33" direction="north" minutes="2.922" orientation="latitude" precision="1" value="33.0487">33°2.922'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="50.840" orientation="longitude" precision="1" value="104.847336">104°50.840'E</geoCoordinate>
; 
<collectingDate value="2016-05-29">29 May 2016</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Laeocathaica carinalis Chen &amp; Zhang, 2004 A, B 2016 / 70 b C, D 2016 / 81 E 2016 / 82 F 2016 / 83. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure10" httpUri="https://binary.pensoft.net/fig/646385" pageId="0" pageNumber="33">10A, B</figureCitation>
) • 10 shells; 
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Wenxian, Shifang Xiang, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
northwest from Baiyiba Cun towards Dongyukou Cun, right side of the road (locality code: 2016/82); 
<geoCoordinate degrees="32" direction="north" minutes="59.346" orientation="latitude" precision="1" value="32.9891">32°59.346'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="37.233" orientation="longitude" precision="1" value="104.62055">104°37.233'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.8" unit="m" value="980.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.8" unit="m" value="980.0">980 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Laeocathaica carinalis Chen &amp; Zhang, 2004 A, B 2016 / 70 b C, D 2016 / 81 E 2016 / 82 F 2016 / 83. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure10" httpUri="https://binary.pensoft.net/fig/646385" pageId="0" pageNumber="33">10E</figureCitation>
) • 4 shells; 
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Wenxian, Shifang Xiang, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.0" unit="m" value="800.0">800 m</elevation>
</quantity>
from the northwestern edge of Baiyiba Cun towards Dongyukou Cun, left bank of the river (locality code: 2016/83); 
<geoCoordinate degrees="32" direction="north" minutes="58.985" orientation="latitude" precision="1" value="32.98308">32°58.985'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="37.503" orientation="longitude" precision="1" value="104.62505">104°37.503'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">970 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Laeocathaica carinalis Chen &amp; Zhang, 2004 A, B 2016 / 70 b C, D 2016 / 81 E 2016 / 82 F 2016 / 83. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure10" httpUri="https://binary.pensoft.net/fig/646385" pageId="0" pageNumber="33">10F</figureCitation>
) • 7 shells; 
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Wenxian, Buziba Xiang, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
south of 
<location LSID="urn:lsid:plazi:treatment:ACC601C874745647AAF5598883342EE4:968E29F004FF065F9CA7AF2131785BEB" country="China" county="Longnan Shi" latitude="32.941185" longLatPrecision="1" longitude="104.67298" municipality="Wenxian" name="Taojiaba Cun" stateProvince="Gansu">Taojiaba Cun</location>
towards Dongyukou Cun (locality code: 2016/79); 
<geoCoordinate degrees="33" direction="north" minutes="01.865" orientation="latitude" precision="1" value="33.031082">33°01.865'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="37.329" orientation="longitude" precision="1" value="104.62215">104°37.329'E</geoCoordinate>
; 
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">1150 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HA • 5 shells; 
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Wenxian, Buziba Xiang, southern edge of Liangjiaba (locality code: 2016/81); 
<geoCoordinate degrees="33" direction="north" minutes="00.262" orientation="latitude" precision="1" value="33.004368">33°00.262'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="36.712" orientation="longitude" precision="1" value="104.61187">104°36.712'E</geoCoordinate>
; 
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.005" unit="m" value="1005.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.005" unit="m" value="1005.0">1005 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2016-05-31">31 May 2016</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HA (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Laeocathaica carinalis Chen &amp; Zhang, 2004 A, B 2016 / 70 b C, D 2016 / 81 E 2016 / 82 F 2016 / 83. Scale bars: 10 mm. All photographs: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77408.figure10" httpUri="https://binary.pensoft.net/fig/646385" pageId="0" pageNumber="33">10C, D</figureCitation>
) • 1 shell; 
<collectingRegion country="China" name="Gansu">Gansu</collectingRegion>
, Longnan Shi, Wenxian, Chengguan Zhen, next to a museum (locality code: 2016/64); 
<geoCoordinate degrees="32" direction="north" minutes="56.471" orientation="latitude" precision="1" value="32.941185">32°56.471'N</geoCoordinate>
, 
<geoCoordinate degrees="104" direction="east" minutes="40.379" orientation="longitude" precision="1" value="104.67298">104°40.379'E</geoCoordinate>
; 
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.65" metricValueMax="9.7" metricValueMin="9.6" unit="m" value="965.0" valueMax="970.0" valueMin="960.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.65" metricValueMax="9.7" metricValueMin="9.6" unit="m" value="965.0" valueMax="970.0" valueMin="960.0">960-970 m</quantity>
a.s.l.
</elevation>
; 
<collectingDate value="2016-05-28">28 May 2016</collectingDate>
; 
<collectorName>A. Hunyadi</collectorName>
leg.; HA
</materialsCitation>
.
</paragraph>
<caption ID-Table-UUID="A68A5616E42A25407B14AD4933F57581" httpUri="http://table.plazi.org/id/A68A5616E42A25407B14AD4933F57581" pageId="0" pageNumber="33" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Table 1.</emphasis>
Shell morphological traits of 
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1899 populations.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="33">
<table pageId="0" pageNumber="33">
<tr pageId="0" pageNumber="33">
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Locality no.</th>
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">D (in mm)</th>
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">H (in mm)</th>
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Dorsal side</th>
<th colspan="1" pageId="0" pageNumber="33" rowspan="1">Body whorl</th>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/64</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">21.9-27.1</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">12-14.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded to slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/65</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">25.5-26.2</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">13.3-14.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded to slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/66</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">23.7-24.5</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.2-12.9</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/67</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">19.9-23.3</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8.8-11</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/68</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">24.3-25.4</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.4-10.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">flat to slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/69</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">27.4-30.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">12.2-14.1</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/70a</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">30-31.3</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">13.1-14.9</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/71</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">28.5-29.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">13-13.6</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/72</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">19.8-24.2</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9.1-12.2</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/73</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">21.8</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/74</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">21.4-25.4</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9-10.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">flat to slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly to slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/75</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">25.1-27.4</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9.8-12.5</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed to domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly to slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/76</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">20.8</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">flat</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/77</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">20.8-21.6</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.5-11.5</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/78</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">22.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.6</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/79</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">20.7-21.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10-10.3</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed to domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly to slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/82</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">20.6-23</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9.8-11.6</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed to domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded to slightly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/83</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">27.7-28.3</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">13-13.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">rounded</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/87</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">23-26.9</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">8.1-11.4</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">flat to slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/88</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">26.6-27.7</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.9-11.3</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">flat to slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/89</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">27.9-30.6</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">11.3-12.1</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">flat to slightly domed</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/91</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">27.5-30</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">10.4-11.5</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">flat</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly keeled</td>
</tr>
<tr pageId="0" pageNumber="33">
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">2016/95</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">23.6-27.4</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">9-10.1</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">flat</td>
<td colspan="1" pageId="0" pageNumber="33" rowspan="1">strongly keeled</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/zookeys.1086.77408.figure9" httpUri="https://binary.pensoft.net/fig/646384" pageId="0" pageNumber="33" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 9.</emphasis>
<taxonomicName authorityName="Chen &amp; Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica carinalis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="carinalis">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica carinalis</emphasis>
</taxonomicName>
Chen &amp; Zhang, 2004 (IZCAS 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/TM097578">TM 097578</accessionNumber>
, 
<typeStatus>holotype</typeStatus>
). Scale bar: 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
. Photographs: Kaibaryer Meng.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="description">
<paragraph pageId="0" pageNumber="33">Description.</paragraph>
<paragraph pageId="0" pageNumber="33">Shell sinistral, depressed, strongly keeled, dorsal side with flat, scalariform whorls; ventral side widely conical; dorsal side chocolate brown, ornamented with a white keel on all whorls; ventral side primarily white, below the white keel there is a chocolate brown belt, white part ornamented with greyish radial stripes that sometimes reach the umbilicus, but sometimes thin and stop before umbilicus; umbilicus inside with a chocolate-brown and a white belt; entire shell consists of 7.25-7.75 whorls; protoconch consists of 1.5-1.75 whorls, brownish, seemingly smooth, extremely finely granulose, rather matte, slightly protruding above first whorls of teleoconch; white keels of every whorl slightly elevated from dorsal surface, but dorsal surface flat with usually the last one being scalariform; dorsal side with fine, irregular wrinkles and between the main wrinkles there are very fine radial lines; ventral surface with less prominent wrinkles; umbilicus rather narrow, funnel-shaped, shows all whorls; periumbilical keel absent; aperture oblique to shell axis, semilunar, with pointed incision at the keel; peristome expanded and slightly thickened, but not reflexed; palatal swelling whitish, with a low, blunt basal tooth; parietal callus practically absent, in some old specimens with translucent calcareous layer.</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Measurements</emphasis>
(in mm): D: 18.6-22.9; H: 6.8-9.8 (
<emphasis italics="true" pageId="0" pageNumber="33">n</emphasis>
= 13, newly collected shells from multiple samples).
</paragraph>
<caption doi="10.3897/zookeys.1086.77408.figure10" httpUri="https://binary.pensoft.net/fig/646385" pageId="0" pageNumber="33" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 10.</emphasis>
<taxonomicName authorityName="Chen &amp; Zhang" authorityYear="2004" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica carinalis" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="carinalis">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica carinalis</emphasis>
</taxonomicName>
Chen &amp; Zhang, 2004 
<emphasis bold="true" pageId="0" pageNumber="33">A, B</emphasis>
2016/70b 
<emphasis bold="true" pageId="0" pageNumber="33">C, D</emphasis>
2016/81 
<emphasis bold="true" pageId="0" pageNumber="33">E</emphasis>
2016/82 
<emphasis bold="true" pageId="0" pageNumber="33">F</emphasis>
2016/83. Scale bars: 10 mm. All photographs: B. 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="differential diagnosis">
<paragraph pageId="0" pageNumber="33">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="33">
The most similar species is 
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. pewzowi" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="pewzowi">
<emphasis italics="true" pageId="0" pageNumber="33">L. pewzowi</emphasis>
</taxonomicName>
, which is smaller, paler in colour, has a wider umbilicus, a more domed (and not scalariform) dorsal side, stronger radial sculpture, and a more oblique aperture with a more pointed basal tooth. Furthermore, there is a second broken belt between the main belt and the umbilicus in 
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. pewzowi" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="pewzowi">
<emphasis italics="true" pageId="0" pageNumber="33">L. pewzowi</emphasis>
</taxonomicName>
, which is not present in any specimens of this species. 
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica potanini" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="potanini">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica potanini</emphasis>
</taxonomicName>
has a more scalariform, uniformly light brown shell, and the basal tooth (when present) is situated closer to the columella than in 
<taxonomicName family="Bradybaenidae" kingdom="Animalia" lsidName="L. carinalis" order="Stylommatophora" pageId="0" pageNumber="33" rank="species" species="carinalis">
<emphasis italics="true" pageId="0" pageNumber="33">L. carinalis</emphasis>
</taxonomicName>
. 
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica amdoana" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="species" species="amdoana">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica amdoana</emphasis>
</taxonomicName>
is also similar in colouration, but it is larger, has a blunter keel, a weaker sculpture, and its whorls are never scalariform.
</paragraph>
<caption doi="10.3897/zookeys.1086.77408.figure11" httpUri="https://binary.pensoft.net/fig/646386" pageId="0" pageNumber="33" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Figure 11.</emphasis>
Distribution of 
<taxonomicName authorityName="Moellendorff" authorityYear="1899" class="Gastropoda" family="Bradybaenidae" genus="Laeocathaica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laeocathaica" order="Stylommatophora" pageId="0" pageNumber="33" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="33">Laeocathaica</emphasis>
</taxonomicName>
species in China (detail from Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of the genus Laeocathaica in China A see Figs 8, 11, 17, 24 for details B see Fig. 14 for detail." figureDoi="10.3897/zookeys.1086.77408.figure2" httpUri="https://binary.pensoft.net/fig/646377" pageId="0" pageNumber="33">2A</figureCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="distribution">
<paragraph pageId="0" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="33">
Most precise locality data are from the rocky area along the Baishui River, whereas one sample was collected on the bank of the Yangtang River (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of Laeocathaica species in China (detail from Fig. 2 A)." figureDoi="10.3897/zookeys.1086.77408.figure11" httpUri="https://binary.pensoft.net/fig/646386" pageId="0" pageNumber="33">11</figureCitation>
). The type locality is situated ca. 50 km west in a straight line.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="remarks">
<paragraph pageId="0" pageNumber="33">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="33">
We here provide a redescription, an updated differential diagnosis, and notes on the differences between different populations (Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Shell morphological traits of Laeocathaica carinalis Chen &amp; Zhang, 2004 populations." httpUri="http://table.plazi.org/id/5D856BFDDD1FAE9FD3739F51A2F34FD0" pageId="0" pageNumber="33" tableUuid="5D856BFDDD1FAE9FD3739F51A2F34FD0">2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>