149 lines
18 KiB
XML
149 lines
18 KiB
XML
<document id="63D2A15576A48E92A1B88259286EB860" ID-CLB-Dataset="33935" ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="2D1E41EDDB2BEFB81562A8B314FD74CB" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha zehntneri Carl 1902" docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="62" updateTime="1732923565725" updateUser="ExternalLinkService">
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<mods:title id="85010E0C53444A0D6E38B34FA4B01672">Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="3AD530C7D506FC49B39292BDA714595C">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="AF6A55AACCCE491A12EA31AB9C75A6B5">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="C91DFBF46EC0FCBC161BA371A8F5D817">Panha, Somsak</mods:namePart>
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<mods:typeOfResource id="92ACE8CF30E8CE902AC8E33AA6F07B3F">text</mods:typeOfResource>
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<mods:title id="7499BE8215B7F1B5BF41271C54B0235E">ZooKeys</mods:title>
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<mods:date id="3B4A44E660D0FA16401F4A7D295E577C">2011</mods:date>
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<mods:number id="51202B51D19D89246FA5B50C65B88620">131</mods:number>
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<mods:classification id="E981E5F391F0365C8CFC17E92C9CAF48">journal article</mods:classification>
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<mods:identifier id="2D9F97010A3AD453CDD309CF09814C8F" type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
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<treatment id="2D1E41EDDB2BEFB81562A8B314FD74CB" ID-GBIF-Taxon="152031148" LSID="urn:lsid:plazi:treatment:2D1E41EDDB2BEFB81562A8B314FD74CB" httpUri="http://treatment.plazi.org/id/2D1E41EDDB2BEFB81562A8B314FD74CB" lastPageId="64" lastPageNumber="65" pageId="61" pageNumber="62" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
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<subSubSection id="202B29AC112ACFE41837DAEC8850BFDD" pageId="61" pageNumber="62" type="nomenclature">
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<paragraph id="0066032C7334AC3CEAB43B805879664F" pageId="61" pageNumber="62">
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<taxonomicName id="47D06C3039A8D97ED7ADA2549F37DB78" ID-CoL="74WQ8" LSID="http://species-id.net/wiki/Orthomorpha_zehntneri" authority="Carl, 1902" authorityName="Carl" authorityYear="1902" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha zehntneri" order="Polydesmida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="zehntneri">Orthomorpha zehntneri Carl, 1902</taxonomicName>
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Figs 5354
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</paragraph>
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</subSubSection>
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<subSubSection id="43BC4FD5472AA274149ADCEE2B5EC8F9" pageId="61" pageNumber="62" type="reference_group">
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<paragraph id="58A58F5C50F1770340E9FD5C9AAD3D50" pageId="61" pageNumber="62">
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<taxonomicName id="358A812418D907D1AAA934FA9763E4B9" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha (Zehntneri)" order="Polydesmida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="subGenus" subGenus="Zehntneri">Orthomorpha Zehntneri</taxonomicName>
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<bibRefCitation id="73A597829CB3D751312523C79238FFC1" author="Carl, J" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="563 - 679" title="Exotische Polydesmiden." volume="10" year="1902">Carl 1902</bibRefCitation>
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: 584 (D).
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</paragraph>
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<paragraph id="D99D5DB82701AA1FE735710EFBD5085E" pageId="61" pageNumber="62">
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<taxonomicName id="692D85DD4FAA73FA12455B3FB27D1578" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha (Zehntneri)" order="Polydesmida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="subGenus" subGenus="Zehntneri">Orthomorpha Zehntneri</taxonomicName>
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-
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<bibRefCitation id="60B88DE1BBCFB91627C42D5D61616941" author="Attems, C" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" pageId="114" pageNumber="115" pagination="77 - 142" title="Javanische Myriopoden, gesammelt von Direktor Dr. K. Kraepelin im Jahre 1903." volume="24" year="1907">Attems 1907</bibRefCitation>
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: 83 (M);
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<bibRefCitation id="15DDE8A4D1E320B84E0A26F9A6B28760" pageId="61" pageNumber="62">1914</bibRefCitation>
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: 192 (M, D).
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</paragraph>
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<paragraph id="CDF3D81ED6ED363625B6A3CC155D6FF3" pageId="61" pageNumber="62">
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<taxonomicName id="F981659C2A089539E07E8FDC677EDC65" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha zehntneri" order="Polydesmida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="zehntneri">Orthomorpha zehntneri</taxonomicName>
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-
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<bibRefCitation id="77A72515930CFE62EA6F7534768D9B15" author="Attems, C" journalOrPublisher="- Bd" pageId="114" pageNumber="115" pagination="115 - 192" title="Myriopoden von Java, Sumatra und Bali. Archiv fuer Hydrobiologie, Suppl." volume="8" year="1930 a">Attems 1930a</bibRefCitation>
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: 127 (D);
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<bibRefCitation id="C678B08DD973113AAAA6B03BCE3966D3" pageId="61" pageNumber="62">1937</bibRefCitation>
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: 73 (D);
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<bibRefCitation id="45530890186FA879207D9793EEF5D733" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
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: 265 (M);
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<bibRefCitation id="EE2046C0853215A26226A491954046AD" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1964</bibRefCitation>
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: 361 (M, D);
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<bibRefCitation id="2A8F4F779C71B8F041C4E94C9A9CCB9C" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="71 - 88" title="A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part III)." volume="123" year="1980 a">1968</bibRefCitation>
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: 45 (M);
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<bibRefCitation id="AA416F79EC12B29A9DEE61394539A106" author="Jeekel, CAW" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="323 - 340" title="On the classification of some little-known Paradoxosomatidae from Java and Sumatra (Diplopoda, Polydesmida)." volume="87" year="1980 b">1980b</bibRefCitation>
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: 325 (D);
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<bibRefCitation id="5EEC12C6EE15AA6A22AAE9611B589C40" pageId="61" pageNumber="62">Golovatch 1998</bibRefCitation>
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: 42 (D, M).
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</paragraph>
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</subSubSection>
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<subSubSection id="0373016EA9ADEAA87FE2ED278F5E8C60" pageId="61" pageNumber="62" type="syntypes">
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<paragraph id="D2E7CF8C0F8D4B585A3F14A7199AA087" pageId="61" pageNumber="62">Syntypes.</paragraph>
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<paragraph id="548D6D44466EAACB7B8681A494C731FE" pageId="61" pageNumber="62">2 ♂, 1 ♀ (MHNG), Indonesia, Java, no date, leg. L. Zehntner.</paragraph>
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</subSubSection>
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<subSubSection id="61CB03E35CC2F08B0764680AF2358F29" lastPageId="64" lastPageNumber="65" pageId="61" pageNumber="62" type="redescription">
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<paragraph id="CB0D8D07149FD791B9A90F089AB5A447" pageId="61" pageNumber="62">Redescription.</paragraph>
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<paragraph id="40A0D0E8BEE0FCD672FD87BC112B9F0F" pageId="61" pageNumber="62">
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Length 19-22 (♂) or 24 mm (♀), width of midbody pro- and metazona 1.4-1.5 and 2.1-2.4 mm (♂), 2.0 and 2.6 mm (♀), respectively (vs 2.8 mm in width, as given in the original description (
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<bibRefCitation id="647FB8A65A73EF34990AA672ABDDEA19" author="Carl, J" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="563 - 679" title="Exotische Polydesmiden." volume="10" year="1902">Carl 1902</bibRefCitation>
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)). Coloration of alcohol material after long-term preservation dark castaneous brown with contrastingly light red-brown to pallid paraterga, venter, and basal podomeres; antenomere 7 blackish, tip of antennae pallid (Fig. 53).
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</paragraph>
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<paragraph id="2A1521B86CE23DA00F130B7879741D24" lastPageId="64" lastPageNumber="65" pageId="61" pageNumber="62">
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Clypeolabral region densely setose, vertex sparsely so, epicranial suture distinct. Antennae moderately long, clavate (antennomere 6 broadest), extending behind body segment 3 (♂) or 2 (♀) dorsally (Fig. 53A & B). Head in width <collum <segment 2 <3 and 4 <5-16; thereafter body gently and gradually tapering. Collum semi-lunar, with three transverse rows of setae, 4+4 anterior, 2+2 intermediate and 3+3 post
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<pageBreakToken id="E8EE9F6A490EC0B2CC074885A6AFBF26" pageId="62" pageNumber="63" start="start">erior</pageBreakToken>
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; caudal corner of paraterga pointed, slightly declined ventrally and continuing
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<normalizedToken id="4ED6444B2BBACD1216C49E5D1D70BE63" originalValue="collum’s">collum's</normalizedToken>
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convexity (Fig. 53B); paraterga subrectangular, slightly surpassing rear tergal margin (Fig. 53A & J). Tegument smooth and shining, prozona very finely shagreened, metaterga leathery, faintly rugulose, below paraterga microgranular, faintly rugulose. Metaterga 2-19 with an anterior transverse row of 2+2, mostly abraded setae; caudal row barely traceable as 3+3 insertion points. Tergal setae simple, rather long, about 1/3 metatergal length. Axial line faint, starting from collum. Paraterga very well developed (Fig. 53A-G & J-L), especially well so in ♂, all lying below dorsum (at about 1/4 midbody height), subhorizontal, in lateral view modestly enlarged on pore-bearing
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<pageBreakToken id="C766D76AB9199C8653026FB6F9CBF737" pageId="63" pageNumber="64" start="start">segments</pageBreakToken>
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, thinner on poreless ones; shoulders always present, mostly regularly rounded and narrowly bordered, fused to callus; caudal corner of all paraterga pointed, beak-like, extending increasingly beyond rear tergal margin, best developed and slightly curved mesad on segments 15-19. Calluses delimited by a sulcus both dorsally and ventrally, with three small, but evident lateral denticles on callus 2, with one or two similar, often setigerous incisions in front of following poreless segments, and with one incision on pore-bearing ones. Posterior edge of paraterga evidently concave, especially strongly so on segments 16-19. Ozopores evident, lateral, lying in an ovoid groove at about 1/4 in front of caudal corner. Transverse sulcus complete on metaterga 5-17, incomplete on segments 4 and 18, narrow, shallow, reaching bases of paraterga, finely beaded at bottom, better developed in ♂ (Fig. 53A, C, F & J-L). Stricture between pro- and metazona narrow and rather shallow, evidently beaded at bottom down to base of paraterga (Fig. 53A-G & J-L). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2 and 3, thereafter like increasingly poorly developed, flat ridges with small caudal teeth until segment 8, further on caudally only like an
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<pageBreakToken id="62456673408EB7EA9345991F38A0894E" pageId="64" pageNumber="65" start="start">increasingly</pageBreakToken>
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small, sharp, caudal tooth until segment 16 (♂, ♀) (Fig. 53B, D & E). Epiproct (Fig. 53E-G & L) conical, inflated dorsoventrally, with two evident apical papillae directed ventrocaudally, subtruncate at tip; pre-apical papillae evident, lying close to tip. Hypoproct (Fig. 53G) subtrapeziform, slightly rounded at caudal margin, setiferous knobs at caudal edge large and well-separated.
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</paragraph>
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<paragraph id="D64C7A056BB5A5EEF4A22BEA8E74BBD1" pageId="64" pageNumber="65">Sterna sparsely setose, without modifications; cross-impressions shallow; with a paramedian pair of very evident, anteroventrally directed lobe deeply, but incompletely split into two between ♂ coxae 4 (Fig. 53H & I). A paramedian pair of small tubercles in front of gonopod aperture. Legs long and slender, midbody ones ca 1.3-1.5 (♂) or 0.9-1.1 times (♀) as long as body height, prefemora without modifications, ♂ tarsal brushes present until legs of segment 16.</paragraph>
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<paragraph id="90C08C32E11C3EB3955AF814467802F9" pageId="64" pageNumber="65">
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Gonopods (Fig. 54) simple. Coxa long and slender, with numerous strong setae distodorsally. Prefemur densely setose, almost 3 times shorter than femorite +
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<normalizedToken id="0CE40DB014CFEE7DDF2C16CE932614ED" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part. Femorite slender, slightly curved and rather evidently enlarged distally, with a
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<normalizedToken id="D6A9DE9E4A55031C46332D62A98850D2" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part demarcated by an oblique lateral sulcus; solenophore with lamina lateralis much smaller than lamina medialis, the former slightly helicoid, forming a tridentate and mesally curved tip; both terminal and subterminal denticles/spikes subequal, acute and narrow, middle denticle nearly as long or shorter; solenomere long and flagelliform.
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</paragraph>
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<caption id="8BE1EE802A5564A6FE726DB8700A8F0D" pageId="64" pageNumber="65">
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<paragraph id="DC5510DF68EBB2F7D3787EEB41991A6E" pageId="64" pageNumber="65">
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Figure 53.
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<taxonomicName id="B48EB1B56C88127B42C1F9344F499478" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha zehntneri" order="Polydesmida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="zehntneri">Orthomorpha zehntneri</taxonomicName>
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Carl, 1902, ♂ syntype (
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<normalizedToken id="95E2D7D9C43765D265D3607B413D42C7" originalValue="A–I">A-I</normalizedToken>
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), ♀ syntype (
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<normalizedToken id="B83F48515752F72562E16A60EB85948F" originalValue="J–L">J-L</normalizedToken>
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). A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D, K segments 10 and 11, dorsal, lateral and dorsal views, respectively
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<normalizedToken id="5ACE61E32D6A29A1E92308CBFC16EF36" originalValue="E–G">E-G</normalizedToken>
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, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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<caption id="431B63F9DF4D71CD3B156391EA478634" pageId="64" pageNumber="65">
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<paragraph id="10F318647D611DD5ABD6AE173DD34DE5" pageId="64" pageNumber="65">
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Figure 54.
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<taxonomicName id="C4B496BA8CB577935497401CFEF98E63" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha zehntneri" order="Polydesmida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="zehntneri">Orthomorpha zehntneri</taxonomicName>
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Carl, 1902, ♂ syntype.
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<normalizedToken id="7A409144F62F5C8C846D9A37569747CD" originalValue="A–C">A-C</normalizedToken>
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right gonopod, lateral, mesal and subcaudal views, respectively.Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="6FD3142209F8044FE92131FEC4016F4C" pageId="64" pageNumber="65" type="remarks">
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<paragraph id="CABD53953FC415098EFCBF7D58998B06" pageId="64" pageNumber="65">Remarks.</paragraph>
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<paragraph id="04FAFD5DBCFA8253F95B3FA0A3A7A79E" pageId="64" pageNumber="65">
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The above redescription is meant to augment the fairly complete one by
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<bibRefCitation id="0E311AAFDF553969A8872F3EF3D89770" author="Jeekel, CAW" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="323 - 340" title="On the classification of some little-known Paradoxosomatidae from Java and Sumatra (Diplopoda, Polydesmida)." volume="87" year="1980 b">Jeekel (1980b)</bibRefCitation>
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which was also based on syntypes.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |