243 lines
19 KiB
XML
243 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e7863" ID-PMC="PMC4867709" ID-Pensoft-Pub="1314-2828--7863" ID-PubMed="27226748" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828--e7863" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="A new species of Phosocephala Townsend, 1908 (Diptera: Tachinidae) from Area de Conservación Guanacaste in northwestern Costa Rica" checkinTime="1585357430397" checkinUser="pensoft" docAuthor="Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Janzen, Daniel H, Hallwachs, Winnie & Dapkey, Tanya" docDate="2016" docId="F458FB5FC76FC7A96370B82AA9500021" docLanguage="en" docName="BiodivDatJour 4: e7863" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e7863" docTitle="Phosocephala metallica Townsend 1908" docType="treatment" docVersion="3" lastPageNumber="7863" masterDocId="ED4BFF9883042C43FFAEFF954127FF89" masterDocTitle="A new species of Phosocephala Townsend, 1908 (Diptera: Tachinidae) from Area de Conservacion Guanacaste in northwestern Costa Rica" masterLastPageNumber="7863" masterPageNumber="7863" pageNumber="7863" updateTime="1668123564579" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>A new species of Phosocephala Townsend, 1908 (Diptera: Tachinidae) from Area de Conservacion Guanacaste in northwestern Costa Rica</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Fleming, AJ</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Wood, D. Monty</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Smith, M. Alex</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Janzen, Daniel H</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Hallwachs, Winnie</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Dapkey, Tanya</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Biodiversity Data Journal</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2016</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>4</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>7863</mods:start>
|
|
<mods:end>7863</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/BDJ.4.e7863</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.4.e7863</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1314-2828--7863</mods:identifier>
|
|
</mods:mods>
|
|
<treatment LSID="urn:lsid:plazi:treatment:F458FB5FC76FC7A96370B82AA9500021" httpUri="http://treatment.plazi.org/id/F458FB5FC76FC7A96370B82AA9500021" lastPageNumber="7863" pageId="0" pageNumber="7863">
|
|
<subSubSection pageId="0" pageNumber="7863" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="7863">
|
|
<taxonomicName authority="Townsend, 1908" authorityName="Townsend" authorityYear="1908" class="Insecta" family="Tachinidae" genus="Phosocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phosocephala metallica" order="Diptera" pageId="0" pageNumber="7863" phylum="Arthropoda" rank="species" species="metallica">Phosocephala metallica Townsend, 1908</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="7863" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="7863">Materials</paragraph>
|
|
<paragraph pageId="0" pageNumber="7863">
|
|
<materialsCitation collectingDate="N / A" collectingMethod="hand collected during field expedition of Schild and Burgdorf" collectionCode="USNM, Insects" collectorName="D. M. Wood" country="Costa Rica" location="Tucurrique" pageId="0" pageNumber="7863" specimenCode="No. 10902" specimenCount="1" typeStatus="Holotype">
|
|
Type status:
|
|
<typeStatus pageId="0" pageNumber="7863">Holotype</typeStatus>
|
|
. Occurrence: occurrenceDetails: http://n2t.net/ark:/65665/39956764d-8fca-4c6c-a18b-a3946f68ea3c; catalogNumber:
|
|
<specimenCode pageId="0" pageNumber="7863">No.10902</specimenCode>
|
|
; recordedBy:
|
|
<collectorName pageId="0" pageNumber="7863">D.M. Wood</collectorName>
|
|
; individualID: No.10902; individualCount:
|
|
<specimenCount pageId="0" pageNumber="7863">1</specimenCount>
|
|
; sex:
|
|
<specimenType pageId="0" pageNumber="7863">F</specimenType>
|
|
; lifeStage:
|
|
<specimenType pageId="0" pageNumber="7863">adult</specimenType>
|
|
; preparations: pinned; Taxon: scientificName: Phosocephalametallica; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Phosocephala; specificEpithet: metallica; scientificNameAuthorship: Townsend, 1908; Location: continent: Central America; country:
|
|
<collectingCountry pageId="0" pageNumber="7863">Costa Rica</collectingCountry>
|
|
; countryCode: CR; stateProvince: Cartago; county: Turrialba; locality:
|
|
<location LSID="urn:lsid:plazi:treatment:F458FB5FC76FC7A96370B82AA9500021:1ABDD83E9E207ECFFBDD35803D746F0A" country="Costa Rica" name="Tucurrique" pageId="0" pageNumber="7863">Tucurrique</location>
|
|
; Event: samplingProtocol:
|
|
<collectingMethod pageId="0" pageNumber="7863">hand collected during field expedition of Schild and Burgdorf</collectingMethod>
|
|
; verbatimEventDate:
|
|
<collectingDate pageId="0" pageNumber="7863">N/A</collectingDate>
|
|
; Record Level: language: en; institutionCode:
|
|
<collectionCode pageId="0" pageNumber="7863">USNM</collectionCode>
|
|
; collectionCode:
|
|
<collectionCode pageId="0" pageNumber="7863">Insects</collectionCode>
|
|
; basisOfRecord: Pinned Specimen
|
|
</materialsCitation>
|
|
<materialsCitation collectingDate="2006-02-15" collectingMethod="Reared from a Nolidae moth larva, Iscadiapurpurascens" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Roster Moraga" country="Costa Rica" latitude="11.027" location="Area de Conservacion Guanacaste" longitude="-85.495" pageId="0" pageNumber="7863" specimenCode="DHJPAR 0006933" specimenCount="1" typeStatus="Other material">
|
|
Type status:
|
|
<typeStatus pageId="0" pageNumber="7863">Other material</typeStatus>
|
|
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
|
|
<specimenCode pageId="0" pageNumber="7863">DHJPAR0006933</specimenCode>
|
|
; recordedBy:
|
|
<collectorName pageId="0" pageNumber="7863">D.H. Janzen & W. Hallwachs, Roster Moraga</collectorName>
|
|
; individualID: DHJPAR0006933; individualCount:
|
|
<specimenCount pageId="0" pageNumber="7863">1</specimenCount>
|
|
; sex:
|
|
<specimenType pageId="0" pageNumber="7863">M</specimenType>
|
|
; lifeStage:
|
|
<specimenType pageId="0" pageNumber="7863">adult</specimenType>
|
|
; preparations: pinned; otherCatalogNumbers: 06-SRNP-20280,BOLD:AAF5995,ASTAV175-06; Taxon: scientificName: Phosocephalametallica; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Phosocephala; specificEpithet: metallica; scientificNameAuthorship: Townsend, 1908; Location: continent: Central America; country:
|
|
<collectingCountry pageId="0" pageNumber="7863">Costa Rica</collectingCountry>
|
|
; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality:
|
|
<location LSID="urn:lsid:plazi:treatment:F458FB5FC76FC7A96370B82AA9500021:099495CF2EA51BC05F6787EBF2BDB74C" country="Costa Rica" latitude="11.027" longitude="-85.495" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="7863">Area de Conservacion Guanacaste</location>
|
|
; verbatimLocality: Quebrada Trigal; verbatimElevation:
|
|
<elevation pageId="0" pageNumber="7863">290</elevation>
|
|
; verbatimLatitude: 11.027; verbatimLongitude: -85.495; verbatimCoordinateSystem: Decimal; decimalLatitude:
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7863" value="11.027">11.027</geoCoordinate>
|
|
; decimalLongitude:
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7863" value="-85.495">-85.495</geoCoordinate>
|
|
; Identification: identifiedBy:
|
|
<determinerName pageId="0" pageNumber="7863">AJ Fleming</determinerName>
|
|
; dateIdentified: 2105; Event: samplingProtocol:
|
|
<collectingMethod pageId="0" pageNumber="7863">Reared from a Nolidae moth larva, Iscadiapurpurascens</collectingMethod>
|
|
; verbatimEventDate:
|
|
<collectingDate pageId="0" pageNumber="7863" value="2006-02-15">15-Feb-2006</collectingDate>
|
|
; Record Level: language: en; institutionCode:
|
|
<collectionCode pageId="0" pageNumber="7863">CNC</collectionCode>
|
|
; collectionCode:
|
|
<collectionCode pageId="0" pageNumber="7863">Insects</collectionCode>
|
|
; basisOfRecord: Pinned Specimen
|
|
</materialsCitation>
|
|
<materialsCitation collectingDate="2003-03-18" collectingMethod="Reared from a Nolidae moth larva, Iscadiapurpurascens" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Roster Moraga" country="Costa Rica" latitude="11.027" location="Area de Conservacion Guanacaste" longitude="-85.495" pageId="0" pageNumber="7863" specimenCode="DHJPAR 0018275" specimenCount="1" typeStatus="Other material">
|
|
Type status:
|
|
<typeStatus pageId="0" pageNumber="7863">Other material</typeStatus>
|
|
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
|
|
<specimenCode pageId="0" pageNumber="7863">DHJPAR0018275</specimenCode>
|
|
; recordedBy:
|
|
<collectorName pageId="0" pageNumber="7863">D.H. Janzen & W. Hallwachs, Roster Moraga</collectorName>
|
|
; individualID: DHJPAR0018275; individualCount:
|
|
<specimenCount pageId="0" pageNumber="7863">1</specimenCount>
|
|
; sex:
|
|
<specimenType pageId="0" pageNumber="7863">M</specimenType>
|
|
; lifeStage:
|
|
<specimenType pageId="0" pageNumber="7863">adult</specimenType>
|
|
; preparations: pinned; otherCatalogNumbers: 03-SRNP-1672,BOLD:AAF5995,ASTAR985-07; Taxon: scientificName: Phosocephalametallica; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Phosocephala; specificEpithet: metallica; scientificNameAuthorship: Townsend, 1908; Location: continent: Central America; country:
|
|
<collectingCountry pageId="0" pageNumber="7863">Costa Rica</collectingCountry>
|
|
; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality:
|
|
<location LSID="urn:lsid:plazi:treatment:F458FB5FC76FC7A96370B82AA9500021:159BCBF7B4112B9B0519D5E8E9C2E113" country="Costa Rica" latitude="11.027" longitude="-85.495" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="7863">Area de Conservacion Guanacaste</location>
|
|
; verbatimLocality: Quebrada Trigal; verbatimElevation:
|
|
<elevation pageId="0" pageNumber="7863">290</elevation>
|
|
; verbatimLatitude: 11.027; verbatimLongitude: -85.495; verbatimCoordinateSystem: Decimal; decimalLatitude:
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7863" value="11.027">11.027</geoCoordinate>
|
|
; decimalLongitude:
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7863" value="-85.495">-85.495</geoCoordinate>
|
|
; Identification: identifiedBy:
|
|
<determinerName pageId="0" pageNumber="7863">AJ Fleming</determinerName>
|
|
; dateIdentified: 2105; Event: samplingProtocol:
|
|
<collectingMethod pageId="0" pageNumber="7863">Reared from a Nolidae moth larva, Iscadiapurpurascens</collectingMethod>
|
|
; verbatimEventDate:
|
|
<collectingDate pageId="0" pageNumber="7863" value="2003-03-18">18-Mar-2003</collectingDate>
|
|
; Record Level: language: en; institutionCode:
|
|
<collectionCode pageId="0" pageNumber="7863">CNC</collectionCode>
|
|
; collectionCode:
|
|
<collectionCode pageId="0" pageNumber="7863">Insects</collectionCode>
|
|
; basisOfRecord: Pinned Specimen
|
|
</materialsCitation>
|
|
<materialsCitation collectingDate="2003-09-10" collectingMethod="Reared from a Nolidae moth larva, Iscadiapurpurascens" collectionCode="CNC, Insects" collectorName="D. H. Janzen & W. Hallwachs, Manuel Pereira" country="Costa Rica" latitude="11.027" location="Area de Conservacion Guanacaste" longitude="-85.495" pageId="0" pageNumber="7863" specimenCode="DHJPAR 0018276" specimenCount="1" typeStatus="Other material">
|
|
Type status:
|
|
<typeStatus pageId="0" pageNumber="7863">Other material</typeStatus>
|
|
. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber:
|
|
<specimenCode pageId="0" pageNumber="7863">DHJPAR0018276</specimenCode>
|
|
; recordedBy:
|
|
<collectorName pageId="0" pageNumber="7863">D.H. Janzen & W. Hallwachs, Manuel Pereira</collectorName>
|
|
; individualID: DHJPAR0018276; individualCount:
|
|
<specimenCount pageId="0" pageNumber="7863">1</specimenCount>
|
|
; sex:
|
|
<specimenType pageId="0" pageNumber="7863">F</specimenType>
|
|
; lifeStage:
|
|
<specimenType pageId="0" pageNumber="7863">adult</specimenType>
|
|
; preparations: pinned; otherCatalogNumbers: 03-SRNP-18792,BOLD:AAF5995,ASTAR986-07; Taxon: scientificName: Phosocephalametallica; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Phosocephala; specificEpithet: metallica; scientificNameAuthorship: Townsend, 1908; Location: continent: Central America; country:
|
|
<collectingCountry pageId="0" pageNumber="7863">Costa Rica</collectingCountry>
|
|
; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality:
|
|
<location LSID="urn:lsid:plazi:treatment:F458FB5FC76FC7A96370B82AA9500021:C7B9AE0F385425D534023F6A4CE9D5C4" country="Costa Rica" latitude="11.027" longitude="-85.495" name="Area de Conservacion Guanacaste" pageId="0" pageNumber="7863">Area de Conservacion Guanacaste</location>
|
|
; verbatimLocality: Quebrada Trigal; verbatimElevation:
|
|
<elevation pageId="0" pageNumber="7863">290</elevation>
|
|
; verbatimLatitude: 11.027; verbatimLongitude: -85.495; verbatimCoordinateSystem: Decimal; decimalLatitude:
|
|
<geoCoordinate orientation="latitude" pageId="0" pageNumber="7863" value="11.027">11.027</geoCoordinate>
|
|
; decimalLongitude:
|
|
<geoCoordinate orientation="longitude" pageId="0" pageNumber="7863" value="-85.495">-85.495</geoCoordinate>
|
|
; Identification: identifiedBy:
|
|
<determinerName pageId="0" pageNumber="7863">AJ Fleming</determinerName>
|
|
; dateIdentified: 2105; Event: samplingProtocol:
|
|
<collectingMethod pageId="0" pageNumber="7863">Reared from a Nolidae moth larva, Iscadiapurpurascens</collectingMethod>
|
|
; verbatimEventDate:
|
|
<collectingDate pageId="0" pageNumber="7863" value="2003-09-10">10-Sep-2003</collectingDate>
|
|
; Record Level: language: en; institutionCode:
|
|
<collectionCode pageId="0" pageNumber="7863">CNC</collectionCode>
|
|
; collectionCode:
|
|
<collectionCode pageId="0" pageNumber="7863">Insects</collectionCode>
|
|
; basisOfRecord: Pinned Specimen
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="7863" type="description">
|
|
<paragraph pageId="0" pageNumber="7863">Description</paragraph>
|
|
<paragraph pageId="0" pageNumber="7863">Described from 2 males and 2 females. Length: 8-9mm.</paragraph>
|
|
<paragraph pageId="0" pageNumber="7863">Head: (Fig. 2b, e) dark reddish-orange to dark yellow; ocellar bristles strongly lateraloclinate; fronto-orbital plate densely haired, with short dark hairs interspersed with yellow hairs; parafacial densely populated with short blond hairs, not obvious unless viewed under varying angles of light; facial carina flat to absent; antennal grooves almost indistinguishable; first flagellomere brilliant orange, with hints of dark brown; gena 0.3x height of eye in males, 0.36x height of eye in females; gena densely haired with short yellow-orange hairs, and with a small tuft of black hairs at base of eye, only apparent in the female. Thorax: (Fig. 2a, c, d, f) scutum gold-pollinose presuturally, with 4 faintly visible thoracic; scutum, postsuturally, with gold pollinosity only on the anterior corners, otherwise glabrous black, including scutellum; thorax, laterally, yellow in ground color, with long yellow hairs laterally, and a light gray pollinosity giving it a silver sheen; katepisternum with 3 bristles. Abdomen: (Fig. 2a, c, d, f) ground color deep shiny reddish-brown dorsally, changing to a lighter reddish-orange ventrally; T1+2 with silver pollinosity covering posterior 1/3 of underside of T1+2 extending into anterior 1/6 of T3; anterior margins of T3 and T4 with a narrow band of thin silver pollinosity, reaching the lateral and ventral sides of the tergites. Legs: bright yellow, with a dense covering of short black hairs. Wings: (Fig. 2a, d) smoky, translucent, bearing 4-5 short setulae at the base of R4+5. Male terminalia: (Fig. 3) sternite 5 with deeply excavated median cleft in a rounded V-shape; apical lobes squared at apex, densely covered in long thick bristles; cercus in dorsal view fused medially, apically pointed with a small bifurcation at tip, densely hirsute basally with numerous long bristles; surstylus divided into two fused processes, both with a strong inward curve; ventral process apically rounded, dorsal process apically hooked.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="7863" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="7863">Diagnosis</paragraph>
|
|
<paragraph pageId="0" pageNumber="7863">
|
|
<taxonomicName class="Insecta" family="Tachinidae" genus="Phosocephala" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phosocephala metallica" order="Diptera" pageId="0" pageNumber="7863" phylum="Arthropoda" rank="species" species="metallica">Phosocephala metallica</taxonomicName>
|
|
differs from
|
|
<taxonomicName lsidName="P. alexanderi" pageId="0" pageNumber="7863" rank="species" species="alexanderi">P. alexanderi</taxonomicName>
|
|
by its larger size, the dark orange head color, the yellow ground color of the thorax laterally, encompassing the katepisternum, anepisternum, katepimeron, and meron, and the conspicuous yellow hairs laterally on the thorax.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="7863" type="distribution">
|
|
<paragraph pageId="0" pageNumber="7863">Distribution</paragraph>
|
|
<paragraph pageId="0" pageNumber="7863">Holotype: Costa Rica, Tucurrique (USNM). Other material: Costa Rica, ACG, Guanacaste, Quebrada Trigal, rain forest, 290m (CNC).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="7863" type="ecology">
|
|
<paragraph pageId="0" pageNumber="7863">Ecology</paragraph>
|
|
<paragraph pageId="0" pageNumber="7863">
|
|
This species was reared from three out of a total of 88 caterpillars of
|
|
<taxonomicName class="Insecta" family="Nolidae" genus="Iscadia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iscadia purpurascens" order="Lepidoptera" pageId="0" pageNumber="7863" phylum="Arthropoda" rank="species" species="purpurascens">Iscadia purpurascens</taxonomicName>
|
|
(Schaus, 1910) (
|
|
<taxonomicName lsidName="" order="Lepidoptera" pageId="0" pageNumber="7863" rank="order">Lepidoptera</taxonomicName>
|
|
:
|
|
<taxonomicName family="Nolidae" lsidName="" pageId="0" pageNumber="7863" rank="family">Nolidae</taxonomicName>
|
|
) bred during the project. The host species feeds on leaves of
|
|
<taxonomicName class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Garcinia intermedia" order="Malpighiales" pageId="0" pageNumber="7863" phylum="Tracheophyta" rank="species" species="intermedia">Garcinia intermedia</taxonomicName>
|
|
(
|
|
<taxonomicName family="Clusiaceae" lsidName="" pageId="0" pageNumber="7863" rank="family">Clusiaceae</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |