treatments-xml/data/A9/6C/87/A96C87A66A76E97CFF0CB0F6FC331378.xml

<document id="7835035ABC91B86B3B41741BA7D982D7" ID-CLB-Dataset="89525" ID-DOI="10.5281/zenodo.5416213" ID-GBIF-Dataset="2d1c446a-b436-4f55-a074-3b4b5b8771c3" ID-ISSN="0253-116X" ID-Zenodo-Dep="5416213" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1630358880360" checkinUser="marcus" docAuthor="Assing, Volker" docDate="2016" docId="A96C87A66A76E97CFF0CB0F6FC331378" docLanguage="en" docName="LinzerbiolBeitr.2016.48.1.229-234.pdf" docOrigin="Linzer biologische Beiträge 48 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Tetrasticta laotica Assing 2016, nov.sp." docType="treatment" docVersion="9" lastPageNumber="233" masterDocId="5555FFDE6A77E978FFBEB130FFCF1144" masterDocTitle="A new species of Tetrasticta from Laos (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="234" masterPageNumber="229" pageNumber="230" updateTime="1699247009521" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6D52F9DCBC874806C6D3608C3FEB2689">A new species of Tetrasticta from Laos (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="C4305AA269E248ACC67603456BE4933C">Assing, Volker</mods:namePart>
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<paragraph id="217A36B06A76E979FF0CB0F6FDB61098" blockId="1.[178,633,453,476]" box="[178,633,453,476]" pageId="1" pageNumber="230">
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<emphasis id="13B1EAA26A76E979FF0CB0F6FE0E109F" bold="true" box="[178,449,453,475]" pageId="1" pageNumber="230">
<taxonomicName id="E6C54D336A76E979FF0CB0F6FEA2109F" ID-CoL="55TK5" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[178,365,453,475]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="230" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
<emphasis id="13B1EAA26A76E979FF0CB0F6FEA2109F" bold="true" box="[178,365,453,475]" italics="true" pageId="1" pageNumber="230">Tetrasticta laotica</emphasis>
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<taxonomicNameLabel id="088257D96A76E979FECAB0F6FE0E109F" box="[372,449,454,475]" pageId="1" pageNumber="230" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B9FE2A356A76E979FE70B0F6FDEB109F" box="[462,548,454,476]" captionStart="Figs 1-4" captionStartId="2.[178,218,916,938]" captionTargetBox="[178,1052,211,898]" captionTargetId="figure-1@2.[178,1057,211,906]" captionTargetPageId="2" captionText="Figs 1-4: Tetrasticta laotica nov.sp.: forebody (1); antenna (2); median lobe of aedeagus (3); spermatheca (4). Scale bars: 1-2: 0.5 mm; 3: 0.2 mm; 4: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5416217" httpUri="https://zenodo.org/record/5416217/files/figure.png" pageId="1" pageNumber="230">Figs 1-4</figureCitation>
, 
<figureCitation id="B9FE2A356A76E979FD91B0F6FDA01098" box="[559,623,454,476]" captionStart="Map 1" captionStartId="3.[178,223,1064,1085]" captionTargetBox="[178,1054,211,1054]" captionTargetId="figure-1@3.[178,1054,211,1054]" captionTargetPageId="3" captionText="Map 1: Distribution of Tetrasticta laotica nov.sp." figureDoi="http://doi.org/10.5281/zenodo.5416219" httpUri="https://zenodo.org/record/5416219/files/figure.png" pageId="1" pageNumber="230">Map 1</figureCitation>
)
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<subSubSection id="69DF653B6A76E979FF0CB0DBFEC3131F" pageId="1" pageNumber="230" type="materials_examined">
<paragraph id="217A36B06A76E979FF0CB0DBFEC3131F" blockId="1.[178,1086,491,603]" pageId="1" pageNumber="230">
Type material: 
<materialsCitation id="91AD3CED6A76E979FE0CB0DBFE66136A" ID-GBIF-Occurrence="3829957301" collectingDate="2003-05-28" collectingDateMax="2003-06-20" collectingDateMin="2003-05-28" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos" determinerName="V. Assing" elevation="1500" latitude="21.6833" location="Phongsaly" longitude="102.1000" municipality="Phongsaly" pageId="1" pageNumber="230" specimenCount="1" specimenCount-male="1" stateProvince="Phongsaly prov." typeStatus="holotype">
<typeStatus id="FE7E88126A76E979FE0CB0DBFDC81345" box="[434,519,491,513]" pageId="1" pageNumber="230" type="holotype">Holotype</typeStatus>
<specimenCount id="37C3FD396A76E979FDAAB0DBFDE81344" box="[532,551,491,512]" pageId="1" pageNumber="230" type="male">♂</specimenCount>
: &quot; 
<collectingCountry id="59D276206A76E979FDFFB0DBFDA71344" box="[577,616,491,512]" pageId="1" pageNumber="230">Lao</collectingCountry>
, 
<collectingRegion id="E301F8526A76E979FDC6B0DBFCD91344" box="[632,790,491,513]" country="Laos" name="Phongsali" pageId="1" pageNumber="230">Phongsaly prov.</collectingRegion>
, 
<geoCoordinate id="44F150776A76E979FC9BB0DBFC4C1344" box="[805,899,491,512]" degrees="21" direction="north" minutes="41.5" orientation="latitude" pageId="1" pageNumber="230" value="21.6833">21°41-2'N</geoCoordinate>
<geoCoordinate id="44F150776A76E979FC2EB0DBFC371345" box="[912,1016,491,513]" degrees="102" direction="east" minutes="06" orientation="longitude" pageId="1" pageNumber="230" value="102.1">102°06-8'E</geoCoordinate>
, 
<date id="557B10706A76E979FBB6B0DBFEDA1353" pageId="1" pageNumber="230" value="2003-05-28" valueMax="2003-06-20" valueMin="2003-05-28">
<collectingDate id="453FE9986A76E979FBB6B0DBFEDA1353" pageId="1" pageNumber="230" value="2003-05-28" valueMax="2003-06-20" valueMin="2003-05-28">28.v.- 20.vi.2003</collectingDate>
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, 
<collectingMunicipality id="C11EACCA6A76E979FEA1B331FEB11353" box="[287,382,513,535]" pageId="1" pageNumber="230">Phongsaly</collectingMunicipality>
env., ~ 
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</quantity>
, 
<collectorName id="8C3053666A76E979FDABB331FDBA1352" box="[533,629,513,535]" pageId="1" pageNumber="230">Vít Kubáň</collectorName>
leg. / Holotypus ♂ 
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<taxonomicName id="E6C54D336A76E979FC91B332FC181352" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[815,983,513,535]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="230" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
<emphasis id="13B1EAA26A76E979FC91B332FC181352" box="[815,983,513,535]" italics="true" pageId="1" pageNumber="230">Tetrasticta laotica</emphasis>
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<taxonomicNameLabel id="088257D96A76E979FC60B332FBDE1353" box="[990,1041,514,535]" pageId="1" pageNumber="230" rank="species">sp. n.</taxonomicNameLabel>
</originalDetermination>
, det. 
<determinerName id="D4A6DA746A76E979FF0CB328FEC1136A" box="[178,270,536,558]" pageId="1" pageNumber="230">V. Assing</determinerName>
2015&quot; (
<collectionCode id="47D4AE756A76E979FEE4B328FE6C136A" box="[346,419,536,558]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="1" pageNumber="230">NHMB</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="91AD3CED6A76E979FE0DB328FC75136A" ID-GBIF-Occurrence="3829957302" box="[435,954,536,562]" collectingDate="2003-05-28" collectingDateMax="2003-06-20" collectingDateMin="2003-05-28" collectionCode="NHMB, cAss" collectorName="Vit Kuban" country="Laos" determinerName="V. Assing" elevation="1500" latitude="21.6833" location="Phongsaly" municipality="Phongsaly" pageId="1" pageNumber="230" specimenCount="2" specimenCount-female="2" stateProvince="Phongsaly prov." typeStatus="paratype">
<typeStatus id="FE7E88126A76E979FE0DB328FDC01369" box="[435,527,536,557]" pageId="1" pageNumber="230" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="37C3FD396A76E979FDA7B328FD851376" box="[537,586,536,562]" pageId="1" pageNumber="230" type="female">2♀♀</specimenCount>
: same data as holotype (
<collectionCode id="47D4AE756A76E979FC8FB328FCB71369" box="[817,888,536,557]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="1" pageNumber="230">NHMB</collectionCode>
, 
<collectionCode id="47D4AE756A76E979FC3DB328FC7C136A" box="[899,947,536,558]" pageId="1" pageNumber="230">cAss</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="91AD3CED6A76E979FC7AB328FEC8131F" ID-GBIF-Occurrence="3829957303" collectingDate="2002-05-01" collectingDateMax="2002-05-09" collectingDateMin="2002-05-01" collectionCode="NHMB" collectorName="Vit Kuban" country="Laos-N" elevation="1100" latitude="20.75" location="Oudom Xai" longLatPrecision="1266" longitude="102.15" pageId="1" pageNumber="230" specimenCount="1" specimenCount-female="1" stateProvince="Oudomxai">
<specimenCount id="37C3FD396A76E979FC7AB328FC2D1376" box="[964,994,536,562]" pageId="1" pageNumber="230" type="female">1♀</specimenCount>
: &quot; 
<collectingCountry id="59D276206A76E979FC49B328FBF21369" box="[1015,1085,536,557]" pageId="1" pageNumber="230">Laos-N</collectingCountry>
(
<collectingRegion id="E301F8526A76E979FF04B31FFED01301" box="[186,287,559,581]" country="Laos" name="Oudomxai" pageId="1" pageNumber="230">Oudomxai</collectingRegion>
), 
<date id="557B10706A76E979FE91B31FFE5C1300" box="[303,403,559,580]" pageId="1" pageNumber="230" value="2002-05-01" valueMax="2002-05-09" valueMin="2002-05-01">
<collectingDate id="453FE9986A76E979FE91B31FFE5C1300" box="[303,403,559,580]" pageId="1" pageNumber="230" value="2002-05-01" valueMax="2002-05-09" valueMin="2002-05-01">1-9.v.2002</collectingDate>
</date>
, ~ 
<quantity id="E63D9B556A76E979FE14B31FFE3C1300" box="[426,499,559,580]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="230" unit="m" value="1100.0">
<elevation id="AAE8D1836A76E979FE14B31FFE3C1300" box="[426,499,559,580]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="230" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 
<geoCoordinate id="44F150776A76E979FE43B31EFD871300" box="[509,584,558,580]" degrees="20" direction="north" minutes="45" orientation="latitude" pageId="1" pageNumber="230" precision="925" value="20.75">20°45'N</geoCoordinate>
<geoCoordinate id="44F150776A76E979FDF1B31FFD6B1300" box="[591,676,559,580]" degrees="102" direction="east" minutes="09" orientation="longitude" pageId="1" pageNumber="230" precision="925" value="102.15">102°09'E</geoCoordinate>
, 
<location id="241A606B6A76E979FD10B31FFCD51301" LSID="urn:lsid:plazi:treatment:A96C87A66A76E97CFF0CB0F6FC331378:241A606B6A76E979FD10B31FFCD51301" box="[686,794,559,581]" country="Laos-N" latitude="20.75" longLatPrecision="1266" longitude="102.15" name="Oudom Xai" pageId="1" pageNumber="230" stateProvince="Oudomxai">Oudom Xai</location>
(
<quantity id="E63D9B556A76E979FC96B31FFCAC1301" box="[808,867,559,581]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="230" unit="km" value="17.0">
<locationDeviation id="30C82F926A76E979FC96B31FFCAC1301" box="[808,867,559,581]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="230" unit="km" value="17.0">17 km</locationDeviation>
</quantity>
<location id="241A606B6A76E979FCD4B31FFC551300" LSID="urn:lsid:plazi:treatment:A96C87A66A76E97CFF0CB0F6FC331378:241A606B6A76E979FCD4B31FFC551300" box="[874,922,559,580]" country="Laos-N" latitude="20.75" longLatPrecision="1266" longitude="102.15" name="NEE" pageId="1" pageNumber="230" stateProvince="Oudomxai">NEE</location>
), 
<collectorName id="8C3053666A76E979FC15B31FFBC41300" box="[939,1035,559,581]" pageId="1" pageNumber="230">Vít Kubáň</collectorName>
leg.&quot; (
<collectionCode id="47D4AE756A76E979FF06B375FECE131F" box="[184,257,581,603]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="1" pageNumber="230">NHMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="69DF653B6A76E979FF0CB359FC67133B" box="[178,936,617,639]" pageId="1" pageNumber="230" type="etymology">
<paragraph id="217A36B06A76E979FF0CB359FC67133B" blockId="1.[178,936,617,639]" box="[178,936,617,639]" pageId="1" pageNumber="230">
Etymology: The specific epithet is an adjective derived from 
<collectingCountry id="59D276206A76E979FCD1B35AFC6C133B" box="[879,931,618,639]" name="Laos" pageId="1" pageNumber="230">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="69DF653B6A76E97AFF0CB3BFFEAB1545" lastPageId="2" lastPageNumber="231" pageId="1" pageNumber="230" type="description">
<paragraph id="217A36B06A76E979FF0CB3BFFDB813B9" blockId="1.[178,1086,655,765]" pageId="1" pageNumber="230">
Description: Body length 4.2-5.0 mm; length of forebody 
<quantity id="E63D9B556A76E979FC08B3BFFBF613E0" box="[950,1081,655,676]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="1" pageNumber="230" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9-2.2 mm</quantity>
. Coloration: forebody blackish; abdomen pale-reddish; legs dark-yellowish to palebrown; antennae blackish-brown to black with the basal 2-3 antennomeres yellowish-red to reddish-brown; maxillary palpi yellowish.
</paragraph>
<paragraph id="217A36B06A76E979FF0CB23BFE7812AD" blockId="1.[178,1085,779,1001]" pageId="1" pageNumber="230">
Head (
<figureCitation id="B9FE2A356A76E979FF49B23BFEFB1266" box="[247,308,779,802]" captionStart="Figs 1-4" captionStartId="2.[178,218,916,938]" captionTargetBox="[178,1052,211,898]" captionTargetId="figure-1@2.[178,1057,211,906]" captionTargetPageId="2" captionText="Figs 1-4: Tetrasticta laotica nov.sp.: forebody (1); antenna (2); median lobe of aedeagus (3); spermatheca (4). Scale bars: 1-2: 0.5 mm; 3: 0.2 mm; 4: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5416217" httpUri="https://zenodo.org/record/5416217/files/figure.png" pageId="1" pageNumber="230">Fig. 1</figureCitation>
) without sexual dimorphism, 1.20-1.25 times as broad as long; punctation moderately dense and rather fine; interstices without distinct microsculpture. Eyes very large and bulging, approximately four times as long as distance from posterior margin of eye to posterior margin of head in dorsal view. Antenna (
<figureCitation id="B9FE2A356A76E979FD44B251FCF81233" box="[762,823,865,887]" captionStart="Figs 1-4" captionStartId="2.[178,218,916,938]" captionTargetBox="[178,1052,211,898]" captionTargetId="figure-1@2.[178,1057,211,906]" captionTargetPageId="2" captionText="Figs 1-4: Tetrasticta laotica nov.sp.: forebody (1); antenna (2); median lobe of aedeagus (3); spermatheca (4). Scale bars: 1-2: 0.5 mm; 3: 0.2 mm; 4: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5416217" httpUri="https://zenodo.org/record/5416217/files/figure.png" pageId="1" pageNumber="230">Fig. 2</figureCitation>
) 
<quantity id="E63D9B556A76E979FCF8B251FC731232" box="[838,956,865,887]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="1" pageNumber="230" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5-1.6 mm</quantity>
long; antennomere IV strongly transverse, asymmetrically disc-shaped, and approximately three times as broad as long; antennomere V much longer than IV; antennomeres V-X of subequal width and approximately 1.5 times as broad as long; XI barely as long as the combined length of IX and X.
</paragraph>
<paragraph id="217A36B06A76E979FF0CB2C8FEEC15D8" blockId="1.[178,1085,1016,1180]" pageId="1" pageNumber="230">
Pronotum (
<figureCitation id="B9FE2A356A76E979FE96B2C8FEA8154A" box="[296,359,1016,1038]" captionStart="Figs 1-4" captionStartId="2.[178,218,916,938]" captionTargetBox="[178,1052,211,898]" captionTargetId="figure-1@2.[178,1057,211,906]" captionTargetPageId="2" captionText="Figs 1-4: Tetrasticta laotica nov.sp.: forebody (1); antenna (2); median lobe of aedeagus (3); spermatheca (4). Scale bars: 1-2: 0.5 mm; 3: 0.2 mm; 4: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5416217" httpUri="https://zenodo.org/record/5416217/files/figure.png" pageId="1" pageNumber="230">Fig. 1</figureCitation>
) strongly transverse, approximately 1.5 times as broad as long and 1.20-1.24 times as broad as head; lateral and posterior margins forming a semi-circle, posterior angles obsolete; disc with four coarse punctures forming a square in the middle, with a coarse lateral puncture on either side, and with additional finer punctation similar to that of head; interstices without microsculpture; lateral margins with five long and erect setae.
</paragraph>
<paragraph id="217A36B06A76E979FF0CB59BFCB115BD" blockId="1.[178,1086,1195,1274]" pageId="1" pageNumber="230">
Elytra (
<figureCitation id="B9FE2A356A76E979FEBEB59BFEF31585" box="[256,316,1195,1217]" captionStart="Figs 1-4" captionStartId="2.[178,218,916,938]" captionTargetBox="[178,1052,211,898]" captionTargetId="figure-1@2.[178,1057,211,906]" captionTargetPageId="2" captionText="Figs 1-4: Tetrasticta laotica nov.sp.: forebody (1); antenna (2); median lobe of aedeagus (3); spermatheca (4). Scale bars: 1-2: 0.5 mm; 3: 0.2 mm; 4: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5416217" httpUri="https://zenodo.org/record/5416217/files/figure.png" pageId="1" pageNumber="230">Fig. 1</figureCitation>
) 0.7-0.8 times as long as pronotum, distinctly gaping posteriorly; punctation fine and dense; interstices without distinct microsculpture. Hind wings present. Metatarsomere I slightly longer than the combined length of II and III.
</paragraph>
<paragraph id="217A36B06A76E979FF0CB439FD891430" blockId="1.[178,1085,1289,1396]" pageId="1" pageNumber="230">Abdomen slightly narrower than elytra; tergites III-IV with, tergite V without shallow and narrow anterior impressions; punctation moderately fine and moderately dense; interstices without microsculpture; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII convex.</paragraph>
<paragraph id="217A36B06A76E979FF0CB4B1FECE1496" blockId="1.[178,1085,1409,1490]" pageId="1" pageNumber="230">
♂: median lobe of aedeagus (
<figureCitation id="B9FE2A356A76E979FE58B4B3FDEA14DD" box="[486,549,1411,1433]" captionStart="Figs 1-4" captionStartId="2.[178,218,916,938]" captionTargetBox="[178,1052,211,898]" captionTargetId="figure-1@2.[178,1057,211,906]" captionTargetPageId="2" captionText="Figs 1-4: Tetrasticta laotica nov.sp.: forebody (1); antenna (2); median lobe of aedeagus (3); spermatheca (4). Scale bars: 1-2: 0.5 mm; 3: 0.2 mm; 4: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5416217" httpUri="https://zenodo.org/record/5416217/files/figure.png" pageId="1" pageNumber="230">Fig. 3</figureCitation>
) 
<quantity id="E63D9B556A76E979FD8BB4B3FD5F14DC" box="[565,656,1411,1433]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="230" unit="mm" value="0.63">0.63 mm</quantity>
long; ventral process weakly sinuate and apically acute; internal sac with long flagellum forming approximately seven coils basally.
</paragraph>
<caption id="75BA66386A75E97AFF0CB2A4FD311284" ID-DOI="http://doi.org/10.5281/zenodo.5416217" ID-Zenodo-Dep="5416217" httpUri="https://zenodo.org/record/5416217/files/figure.png" pageId="2" pageNumber="231" startId="2.[178,218,916,938]" targetBox="[178,1052,211,898]" targetPageId="2">
<paragraph id="217A36B06A75E97AFF0CB2A4FD311284" blockId="2.[178,1085,916,961]" pageId="2" pageNumber="231">
<emphasis id="13B1EAA26A75E97AFF0CB2A4FECC12ED" bold="true" box="[178,259,916,938]" pageId="2" pageNumber="231">Figs 1-4</emphasis>
: 
<taxonomicName id="E6C54D336A75E97AFEAAB2A5FE0F12ED" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[276,448,916,938]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
<emphasis id="13B1EAA26A75E97AFEAAB2A5FE0F12ED" box="[276,448,916,938]" italics="true" pageId="2" pageNumber="231">Tetrasticta laotica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088257D96A75E97AFE75B2A5FDDF12EE" box="[459,528,917,938]" pageId="2" pageNumber="231" rank="species">nov.sp.</taxonomicNameLabel>
: forebody (
<emphasis id="13B1EAA26A75E97AFD3FB2A4FD4312ED" bold="true" box="[641,652,916,937]" pageId="2" pageNumber="231">1</emphasis>
); antenna (
<emphasis id="13B1EAA26A75E97AFD40B2A4FCC612ED" bold="true" box="[766,777,916,937]" pageId="2" pageNumber="231">2</emphasis>
); median lobe of aedeagus (
<emphasis id="13B1EAA26A75E97AFB9AB2A4FBE012ED" bold="true" box="[1060,1071,916,937]" pageId="2" pageNumber="231">3</emphasis>
); spermatheca (
<emphasis id="13B1EAA26A75E97AFE8FB29BFEF31284" bold="true" box="[305,316,939,960]" pageId="2" pageNumber="231">4</emphasis>
). Scale bars: 1-2: 0.5 mm; 3: 0.2 mm; 4: 0.1 mm.
</paragraph>
</caption>
<paragraph id="217A36B06A75E97AFF0CB2E2FEAB1545" blockId="2.[178,1085,974,1025]" pageId="2" pageNumber="231">
<specimenCount id="37C3FD396A75E97AFF0CB2E2FF0812AD" box="[178,199,978,1001]" pageId="2" pageNumber="231" type="female">♀</specimenCount>
: spermatheca (
<figureCitation id="B9FE2A356A75E97AFEDCB2FEFE5012A1" box="[354,415,974,997]" captionStart="Figs 1-4" captionStartId="2.[178,218,916,938]" captionTargetBox="[178,1052,211,898]" captionTargetId="figure-1@2.[178,1057,211,906]" captionTargetPageId="2" captionText="Figs 1-4: Tetrasticta laotica nov.sp.: forebody (1); antenna (2); median lobe of aedeagus (3); spermatheca (4). Scale bars: 1-2: 0.5 mm; 3: 0.2 mm; 4: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5416217" httpUri="https://zenodo.org/record/5416217/files/figure.png" pageId="2" pageNumber="231">Fig. 4</figureCitation>
) 
<quantity id="E63D9B556A75E97AFE13B2FFFE3512A0" box="[429,506,975,996]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="231" unit="mm" value="0.2">0.2 mm</quantity>
long, with oval distal portion and with short and straight proximal portion.
</paragraph>
</subSubSection>
<subSubSection id="69DF653B6A75E97CFF0CB520FE761026" lastPageId="4" lastPageNumber="233" pageId="2" pageNumber="231" type="diagnosis">
<paragraph id="217A36B06A75E97AFF0CB520FE94151A" blockId="2.[178,1085,1040,1118]" pageId="2" pageNumber="231">
Comparative notes: The new species differs from other geographically close 
<taxonomicName id="E6C54D336A75E97AFF51B51DFE921506" authority="KRAATZ, 1857" authorityName="KRAATZ" authorityYear="1857" box="[239,349,1069,1090]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="13B1EAA26A75E97AFF51B51DFE921506" box="[239,349,1069,1090]" italics="true" pageId="2" pageNumber="231">Tetrasticta</emphasis>
</taxonomicName>
species by the shape of the median lobe of the aedeagus and additionally as follows:
</paragraph>
<paragraph id="217A36B06A75E97AFF0CB55DFEDD159D" blockId="2.[178,1086,1133,1241]" pageId="2" pageNumber="231">
from the similarly coloured 
<taxonomicName id="E6C54D336A75E97AFE67B55EFD3115C6" authority="PACE, 2000" authorityName="PACE" authorityYear="2000" box="[473,766,1133,1155]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="13B1EAA26A75E97AFE67B55EFDB215C6" box="[473,637,1133,1155]" italics="true" pageId="2" pageNumber="231">T. thailandensis</emphasis>
<bibRefCitation id="45544B416A75E97AFD38B55EFD3115C6" author="PACE R" box="[646,766,1133,1155]" pageId="2" pageNumber="231" pagination="39 - 86" refId="ref1913" refString="PACE R. (2000): Aleocharinae della Thailandia (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 17 (1): 39 - 86." type="journal article" year="2000">PACE, 2000</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="59D276206A75E97AFCB0B55DFCA215C7" box="[782,877,1133,1155]" name="Thailand" pageId="2" pageNumber="231">Thailand</collectingCountry>
; male unknown) by the distinct punctation of the head and pronotum (indistinct or absent in 
<taxonomicName id="E6C54D336A75E97AFC30B5BAFBE115DB" authorityName="PACE" authorityYear="2000" box="[910,1070,1162,1183]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="13B1EAA26A75E97AFC30B5BAFBE115DB" box="[910,1070,1162,1183]" italics="true" pageId="2" pageNumber="231">T. thailandensis</emphasis>
</taxonomicName>
), as well as the shape of the spermatheca (proximal portion of the capsule distinctly dilated apically);
</paragraph>
<paragraph id="217A36B06A75E97AFF0CB5D7FC13142B" blockId="2.[178,1086,1255,1391]" pageId="2" pageNumber="231">
from 
<taxonomicName id="E6C54D336A75E97AFF54B5D8FD8015B9" authority="(BERNHAUER, 1903)" baseAuthorityName="BERNHAUER" baseAuthorityYear="1903" box="[234,591,1255,1277]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="13B1EAA26A75E97AFF54B5D8FEB515B8" box="[234,378,1255,1277]" italics="true" pageId="2" pageNumber="231">T. brevipennis</emphasis>
(BERNHAUER, 1903)
</taxonomicName>
(widespread in the Oriental and East Palaearctic regions) by the absence of distinct dimorphisms of the head and segment VIII, the absence of microsculpture on the head and pronotum, the presence of four coarse macropunctures in the middle of the pronotum, the completely reddish abdomen (
<taxonomicName id="E6C54D336A75E97AFB96B40EFEE7142A" baseAuthorityName="BERNHAUER" baseAuthorityYear="1903" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="13B1EAA26A75E97AFB96B40EFEE7142A" italics="true" pageId="2" pageNumber="231">T. brevipennis</emphasis>
</taxonomicName>
: at least segments V-VIII blackish), and numerous other characters;
</paragraph>
<paragraph id="217A36B06A75E97AFF0CB44EFE0C1489" blockId="2.[178,1085,1406,1485]" pageId="2" pageNumber="231">
from 
<taxonomicName id="E6C54D336A75E97AFF52B44FFDB214D7" authority="MARUYAMA &amp; SUGAYA, 2002" authorityName="MARUYAMA &amp; SUGAYA" authorityYear="2002" box="[236,637,1406,1428]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="laeta">
<emphasis id="13B1EAA26A75E97AFF52B44FFEF314D7" box="[236,316,1406,1428]" italics="true" pageId="2" pageNumber="231">T. laeta</emphasis>
<bibRefCitation id="45544B416A75E97AFEFAB44EFDB214D7" author="MARUYAMA M" box="[324,637,1406,1427]" firstAuthor="MARUYAMA" pageId="2" pageNumber="231" pagination="17 - 21" refId="ref1870" refString="MARUYAMA M. &amp; H. SUGAYA (2002): A new species of Tetrasticta (Coleoptera, Staphylinidae, Aleocharinae) from Japan and Taiwan. - Japanese Journal of Systematic Entomology 8 (1): 17 - 21." type="journal article" year="2002">MARUYAMA &amp; SUGAYA, 2002</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="59D276206A75E97AFD32B44EFD0614D7" box="[652,713,1406,1427]" name="Japan" pageId="2" pageNumber="231">Japan</collectingCountry>
, Taiwan) by uniformly black elytra and a reddish abdomen (
<taxonomicName id="E6C54D336A75E97AFE7BB4ABFDD514EB" authorityName="MARUYAMA &amp; SUGAYA" authorityYear="2002" box="[453,538,1434,1456]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="laeta">
<emphasis id="13B1EAA26A75E97AFE7BB4ABFDD514EB" box="[453,538,1434,1456]" italics="true" pageId="2" pageNumber="231">T. laeta</emphasis>
</taxonomicName>
: elytra yellow with black postero-lateral portions; abdomen blackish-brown);
</paragraph>
<paragraph id="217A36B06A75E97BFF0CB4ECFD4415CA" blockId="2.[178,1085,1500,1550]" lastBlockId="3.[178,1085,1116,1166]" lastPageId="3" lastPageNumber="232" pageId="2" pageNumber="231">
from 
<taxonomicName id="E6C54D336A75E97AFF52B4ECFD4214B6" authority="(CAMERON, 1943) (Malaysia)" baseAuthorityName="CAMERON" baseAuthorityYear="1943" box="[236,653,1500,1522]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="13B1EAA26A75E97AFF52B4ECFE9D14B5" box="[236,338,1500,1521]" italics="true" pageId="2" pageNumber="231">T. bicolor</emphasis>
(CAMERON, 1943) (
<collectingCountry id="59D276206A75E97AFD98B4ECFD4914B6" box="[550,646,1500,1522]" name="Malaysia" pageId="2" pageNumber="231">Malaysia</collectingCountry>
)
</taxonomicName>
by uniformly blackish elytra (
<taxonomicName id="E6C54D336A75E97AFC71B4ECFBFA14B5" baseAuthorityName="CAMERON" baseAuthorityYear="1943" box="[975,1077,1500,1521]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="13B1EAA26A75E97AFC71B4ECFBFA14B5" box="[975,1077,1500,1521]" italics="true" pageId="2" pageNumber="231">T. bicolor</emphasis>
</taxonomicName>
: elytra mainly yellow) and the reddish abdomen (
<taxonomicName id="E6C54D336A75E97AFD21B4C9FCCC1749" baseAuthorityName="CAMERON" baseAuthorityYear="1943" box="[671,771,1528,1550]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="13B1EAA26A75E97AFD21B4C9FCCC1749" box="[671,771,1528,1550]" italics="true" pageId="2" pageNumber="231">T. bicolor</emphasis>
</taxonomicName>
: abdomen blackish-brown); from 
<taxonomicName id="E6C54D336A74E97BFF4FB56CFDE61536" authority="(CAMERON, 1943)" baseAuthorityName="CAMERON" baseAuthorityYear="1943" box="[241,553,1116,1138]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="13B1EAA26A74E97BFF4FB56CFE931535" box="[241,348,1116,1137]" italics="true" pageId="3" pageNumber="232">T. bryanti</emphasis>
(CAMERON, 1943)
</taxonomicName>
by the uniformly reddish abdomen (
<taxonomicName id="E6C54D336A74E97BFC74B56CFBF91535" baseAuthorityName="CAMERON" baseAuthorityYear="1943" box="[970,1078,1116,1137]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="13B1EAA26A74E97BFC74B56CFBF91535" box="[970,1078,1116,1137]" italics="true" pageId="3" pageNumber="232">T. bryanti</emphasis>
</taxonomicName>
: abdominal segments IV-VIII blackish-brown);
</paragraph>
<caption id="75BA66386A74E97BFF0CB518FDB81579" ID-DOI="http://doi.org/10.5281/zenodo.5416219" ID-Zenodo-Dep="5416219" box="[178,631,1064,1086]" httpUri="https://zenodo.org/record/5416219/files/figure.png" pageId="3" pageNumber="232" startId="3.[178,223,1064,1085]" targetBox="[178,1054,211,1054]" targetPageId="3">
<paragraph id="217A36B06A74E97BFF0CB518FDB81579" blockId="3.[178,631,1064,1086]" box="[178,631,1064,1086]" pageId="3" pageNumber="232">
<emphasis id="13B1EAA26A74E97BFF0CB518FF3F1579" bold="true" box="[178,240,1064,1085]" pageId="3" pageNumber="232">Map 1</emphasis>
: Distribution of 
<taxonomicName id="E6C54D336A74E97BFE36B519FDE01579" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[392,559,1064,1086]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
<emphasis id="13B1EAA26A74E97BFE36B519FDE01579" box="[392,559,1064,1086]" italics="true" pageId="3" pageNumber="232">Tetrasticta laotica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="088257D96A74E97BFD8BB518FDB81579" box="[565,631,1064,1085]" pageId="3" pageNumber="232" rank="species">nov.sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="217A36B06A74E97BFF0CB5ADFD5A158B" blockId="3.[178,1085,1181,1231]" pageId="3" pageNumber="232">
from 
<taxonomicName id="E6C54D336A74E97BFF54B5ADFD8915F7" authority="CAMERON, 1939" authorityName="CAMERON" authorityYear="1939" box="[234,582,1181,1203]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="13B1EAA26A74E97BFF54B5ADFE9C15F6" box="[234,339,1181,1202]" italics="true" pageId="3" pageNumber="232">T. elegans</emphasis>
CAMERON, 1939 (Java)
</taxonomicName>
by much darker coloration of the forebody alone (
<taxonomicName id="E6C54D336A74E97BFF04B58AFEED158A" authorityName="CAMERON" authorityYear="1939" box="[186,290,1209,1231]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="13B1EAA26A74E97BFF04B58AFEED158A" box="[186,290,1209,1231]" italics="true" pageId="3" pageNumber="232">T. elegans</emphasis>
</taxonomicName>
: head and pronotum yellowish-red);
</paragraph>
<paragraph id="217A36B06A74E97BFF0CB5EEFD1E1469" blockId="3.[178,1085,1246,1325]" pageId="3" pageNumber="232">
from 
<taxonomicName id="E6C54D336A74E97BFF55B5EEFD8F15B0" authority="CAMERON, 1939" authorityName="CAMERON" authorityYear="1939" box="[235,576,1246,1268]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="13B1EAA26A74E97BFF55B5EEFE8315B7" box="[235,332,1246,1267]" italics="true" pageId="3" pageNumber="232">T. javana</emphasis>
CAMERON, 1939 (Java)
</taxonomicName>
by the uniformly black pronotum and elytra, and by the pale-reddish abdomen (
<taxonomicName id="E6C54D336A74E97BFDB6B5CBFDBE1454" authorityName="CAMERON" authorityYear="1939" box="[520,625,1275,1296]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="13B1EAA26A74E97BFDB6B5CBFDBE1454" box="[520,625,1275,1296]" italics="true" pageId="3" pageNumber="232">T. javana</emphasis>
</taxonomicName>
: pronotum red; humeral angles of elytra yellowish-red; abdominal segments V-VIII blackish);
</paragraph>
<paragraph id="217A36B06A74E97BFF0CB40BFEF214CE" blockId="3.[178,1085,1339,1418]" pageId="3" pageNumber="232">
from 
<taxonomicName id="E6C54D336A74E97BFF50B40CFD491415" authority="PACE, 2008" authorityName="PACE" authorityYear="2008" box="[238,646,1339,1361]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="13B1EAA26A74E97BFF50B40CFE561414" box="[238,409,1339,1361]" italics="true" pageId="3" pageNumber="232">T. kinabaluensis</emphasis>
<bibRefCitation id="45544B416A74E97BFE1BB40CFDEF1414" author="PACE R" box="[421,544,1339,1361]" pageId="3" pageNumber="232" pagination="157 - 183" refId="ref1948" refString="PACE R. (2008): Le specie di Thamiaraeini, Oxypodini, Hoplandriini e Aleocharini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 157 - 183." type="journal article" year="2008">PACE, 2008</bibRefCitation>
(Borneo)
</taxonomicName>
by much stouter antennae, a completely reddish abdomen (
<taxonomicName id="E6C54D336A74E97BFEC9B469FDEE1429" authorityName="PACE" authorityYear="2008" box="[375,545,1368,1390]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="13B1EAA26A74E97BFEC9B469FDEE1429" box="[375,545,1368,1390]" italics="true" pageId="3" pageNumber="232">T. kinabaluensis</emphasis>
</taxonomicName>
: segments IV-VII brown), and a pronotum of different shape);
</paragraph>
<paragraph id="217A36B06A74E97BFF0CB4A9FDE0148F" blockId="3.[178,1085,1433,1483]" pageId="3" pageNumber="232">
from 
<taxonomicName id="E6C54D336A74E97BFF52B4AAFD4114EB" authority="PACE, 2013" authorityName="PACE" authorityYear="2013" box="[236,654,1433,1455]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="caputcyrneum">
<emphasis id="13B1EAA26A74E97BFF52B4AAFE5814EB" box="[236,407,1434,1455]" italics="true" pageId="3" pageNumber="232">T. caputcyrneum</emphasis>
PACE, 2013 (
<collectingCountry id="59D276206A74E97BFD99B4A9FD4814EB" box="[551,647,1433,1455]" name="Malaysia" pageId="3" pageNumber="232">Malaysia</collectingCountry>
)
</taxonomicName>
by the reddish abdomen and the absence of microsculpture on the whole body;
</paragraph>
<paragraph id="217A36B06A74E97BFF0CB4EAFEA4176D" blockId="3.[178,1085,1498,1577]" pageId="3" pageNumber="232">
from 
<taxonomicName id="E6C54D336A74E97BFF53B4EBFCFD14B4" authority="YAMAMOTO &amp; MARUYAMA, 2013" authorityName="YAMAMOTO &amp; MARUYAMA" authorityYear="2013" box="[237,818,1498,1520]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="gnatha">
<emphasis id="13B1EAA26A74E97BFF53B4EBFE9F14AB" box="[237,336,1498,1520]" italics="true" pageId="3" pageNumber="232">T. gnatha</emphasis>
YAMAMOTO &amp; MARUYAMA, 2013 (
<collectingCountry id="59D276206A74E97BFD75B4EAFCE414B4" box="[715,811,1498,1520]" name="Malaysia" pageId="3" pageNumber="232">Malaysia</collectingCountry>
)
</taxonomicName>
by the blackish antennomere IV (reddish in 
<taxonomicName id="E6C54D336A74E97BFE3FB4C7FE2D174F" authorityName="YAMAMOTO &amp; MARUYAMA" authorityYear="2013" box="[385,482,1526,1548]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="gnatha">
<emphasis id="13B1EAA26A74E97BFE3FB4C7FE2D174F" box="[385,482,1526,1548]" italics="true" pageId="3" pageNumber="232">T. gnatha</emphasis>
</taxonomicName>
) and the uniformly reddish abdomen (tergites V-VII infuscate in 
<taxonomicName id="E6C54D336A74E97BFF45B724FE93176C" authorityName="YAMAMOTO &amp; MARUYAMA" authorityYear="2013" box="[251,348,1555,1577]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="gnatha">
<emphasis id="13B1EAA26A74E97BFF45B724FE93176C" box="[251,348,1555,1577]" italics="true" pageId="3" pageNumber="232">T. gnatha</emphasis>
</taxonomicName>
);
</paragraph>
<paragraph id="217A36B06A73E97CFF0CB1EAFE761026" blockId="4.[178,1085,218,354]" pageId="4" pageNumber="233">
from 
<taxonomicName id="E6C54D336A73E97CFF54B1EBFDFF11AB" authority="ZHENG &amp; ZHAO, 2014" authorityName="ZHENG &amp; ZHAO" authorityYear="2014" box="[234,560,218,240]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="bobbii">
<emphasis id="13B1EAA26A73E97CFF54B1EBFE8811AB" box="[234,327,218,240]" italics="true" pageId="4" pageNumber="233">T. bobbii</emphasis>
ZHENG &amp; ZHAO, 2014
</taxonomicName>
(
<collectingCountry id="59D276206A73E97CFD81B1EAFD4E11B4" box="[575,641,218,240]" name="China" pageId="4" pageNumber="233">China</collectingCountry>
: 
<collectingRegion id="E301F8526A73E97CFD35B1EBFD1211B4" box="[651,733,219,240]" country="China" name="Yunnan" pageId="4" pageNumber="233">Yunnan</collectingRegion>
) by larger and more bulging eyes, a blackish antennomere IV (reddish in 
<taxonomicName id="E6C54D336A73E97CFDF9B1C7FD6A1048" authorityName="ZHENG &amp; ZHAO" authorityYear="2014" box="[583,677,247,268]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="bobbii">
<emphasis id="13B1EAA26A73E97CFDF9B1C7FD6A1048" box="[583,677,247,268]" italics="true" pageId="4" pageNumber="233">T. bobbii</emphasis>
</taxonomicName>
), the uniformly blackish pronotum and elytra, a uniformly reddish abdomen (tergites V-VII at least partly infuscate in 
<taxonomicName id="E6C54D336A73E97CFC6CB024FBE1106C" authorityName="ZHENG &amp; ZHAO" authorityYear="2014" box="[978,1070,275,297]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="bobbii">
<emphasis id="13B1EAA26A73E97CFC6CB024FBE1106C" box="[978,1070,275,297]" italics="true" pageId="4" pageNumber="233">T. bobbii</emphasis>
</taxonomicName>
), the presence of six coarse macropunctures on the pronotum (absent in 
<taxonomicName id="E6C54D336A73E97CFC3FB000FC121001" authorityName="ZHENG &amp; ZHAO" authorityYear="2014" box="[897,989,304,325]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="bobbii">
<emphasis id="13B1EAA26A73E97CFC3FB000FC121001" box="[897,989,304,325]" italics="true" pageId="4" pageNumber="233">T. bobbii</emphasis>
</taxonomicName>
), and the shape of the spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="69DF653B6A73E97CFF0CB041FDD11098" pageId="4" pageNumber="233" type="description">
<paragraph id="217A36B06A73E97CFF0CB041FDD11098" blockId="4.[178,1086,369,476]" pageId="4" pageNumber="233">
For illustrations of 
<taxonomicName id="E6C54D336A73E97CFEC1B041FDDC10C2" baseAuthorityName="BERNHAUER" baseAuthorityYear="1903" box="[383,531,369,390]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="13B1EAA26A73E97CFEC1B041FDDC10C2" box="[383,531,369,390]" italics="true" pageId="4" pageNumber="233">T. brevipennis</emphasis>
</taxonomicName>
, 
<taxonomicName id="E6C54D336A73E97CFD9DB041FD4410C2" baseAuthorityName="CAMERON" baseAuthorityYear="1943" box="[547,651,369,390]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="13B1EAA26A73E97CFD9DB041FD4410C2" box="[547,651,369,390]" italics="true" pageId="4" pageNumber="233">T. bicolor</emphasis>
</taxonomicName>
, 
<taxonomicName id="E6C54D336A73E97CFD25B041FCCD10C2" baseAuthorityName="CAMERON" baseAuthorityYear="1943" box="[667,770,369,390]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="13B1EAA26A73E97CFD25B041FCCD10C2" box="[667,770,369,390]" italics="true" pageId="4" pageNumber="233">T. bryanti</emphasis>
</taxonomicName>
, 
<taxonomicName id="E6C54D336A73E97CFCACB041FCAC10C2" authorityName="MARUYAMA &amp; SUGAYA" authorityYear="2002" box="[786,867,369,390]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="laeta">
<emphasis id="13B1EAA26A73E97CFCACB041FCAC10C2" box="[786,867,369,390]" italics="true" pageId="4" pageNumber="233">T. laeta</emphasis>
</taxonomicName>
, 
<taxonomicName id="E6C54D336A73E97CFCCDB041FBD710C2" authorityName="PACE" authorityYear="2000" box="[883,1048,369,390]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="13B1EAA26A73E97CFCCDB041FBD710C2" box="[883,1048,369,390]" italics="true" pageId="4" pageNumber="233">T. thailandensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="E6C54D336A73E97CFB96B041FEF210E6" authorityName="PACE" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="13B1EAA26A73E97CFB96B041FEF210E6" italics="true" pageId="4" pageNumber="233">T. kinabaluensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="E6C54D336A73E97CFEECB0BEFDCC10E7" authorityName="PACE" authorityYear="2013" box="[338,515,398,419]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="caputcyrneum">
<emphasis id="13B1EAA26A73E97CFEECB0BEFDCC10E7" box="[338,515,398,419]" italics="true" pageId="4" pageNumber="233">T. caputcyrneum</emphasis>
</taxonomicName>
, 
<taxonomicName id="E6C54D336A73E97CFDA6B0BEFD4E10E6" authorityName="YAMAMOTO &amp; MARUYAMA" authorityYear="2013" box="[536,641,397,419]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="gnatha">
<emphasis id="13B1EAA26A73E97CFDA6B0BEFD4E10E6" box="[536,641,397,419]" italics="true" pageId="4" pageNumber="233">T. gnatha</emphasis>
</taxonomicName>
, and 
<taxonomicName id="E6C54D336A73E97CFD76B0BEFCE410E6" authorityName="ZHENG &amp; ZHAO" authorityYear="2014" box="[712,811,397,419]" class="Insecta" family="Staphylinidae" genus="Tetrasticta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="bobbii">
<emphasis id="13B1EAA26A73E97CFD76B0BEFCE410E6" box="[712,811,397,419]" italics="true" pageId="4" pageNumber="233">T. bobbii</emphasis>
</taxonomicName>
see 
<bibRefCitation id="45544B416A73E97CFCD9B0BEFBF610E7" author="MARUYAMA M" box="[871,1081,397,419]" pageId="4" pageNumber="233" pagination="621 - 637" refId="ref1834" refString="MARUYAMA M. (2004): Redescription of the genus Creochara (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini) and its systematic position. - The Canadian Entomologist 136: 621 - 637." type="journal article" year="2004">MARUYAMA (2004)</bibRefCitation>
, MARUYAMA &amp; SUGAYA (2002), 
<bibRefCitation id="45544B416A73E97CFDBCB09AFDB110FB" author="PACE R" box="[514,638,426,447]" pageId="4" pageNumber="233" pagination="39 - 86" refId="ref1913" refString="PACE R. (2000): Aleocharinae della Thailandia (Coleoptera, Staphylinidae). - Bollettino del Museo Regionale di Scienze Naturali, Torino 17 (1): 39 - 86." type="journal article" year="2000">PACE (2000</bibRefCitation>
, 
<bibRefCitation id="45544B416A73E97CFD34B09AFD7010FB" author="PACE R" box="[650,703,426,447]" pageId="4" pageNumber="233" pagination="157 - 183" refId="ref1948" refString="PACE R. (2008): Le specie di Thamiaraeini, Oxypodini, Hoplandriini e Aleocharini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 157 - 183." type="journal article" year="2008">2008</bibRefCitation>
, 
<bibRefCitation id="45544B416A73E97CFD75B09AFCC11084" author="PACE R" box="[715,782,426,448]" pageId="4" pageNumber="233" pagination="33 - 63" refId="ref2065" refString="PACE R. (2013 b): New distributional data, new species and three new genera of Aleocharinae from Malaysia, Vietnam and Taiwan (Coleoptera: Staphylinidae). - Tropical Zoology 26 (1): 33 - 63." type="journal article" year="2013">2013b</bibRefCitation>
), 
<bibRefCitation id="45544B416A73E97CFC9CB09AFF381098" author="YAMAMOTO S" firstAuthor="YAMAMOTO" pageId="4" pageNumber="233" pagination="39 - 46" refId="ref2168" refString="YAMAMOTO S. &amp; M. MARUYAMA (2013): A peculiar new species of the genus Tetrasticta KRAATZ (Coleoptera, Staphylinidae, Aleocharinae) from Peninsular Malaysia. - ZooKeys 336: 39 - 46." type="journal article" year="2013">YAMAMOTO &amp; MARUYAMA (2013)</bibRefCitation>
, and 
<bibRefCitation id="45544B416A73E97CFE93B0F7FDD61098" author="ZHENG D. - L. &amp; M. - J. ZHAO" box="[301,537,454,476]" firstAuthor="ZHENG" journalOrPublisher="ZooKeys" pageId="4" pageNumber="233" pagination="113 - 116" part="404" refId="ref2207" refString="ZHENG D. - L. &amp; M. - J. ZHAO (2014): A new species of genus [sic] Tetrasticta KRAATZ (Coleoptera, Staphylinidae, Aleocharinae) from Xishuangbanna, Southwest China. - ZooKeys 404: 113 - 116." title="A new species of genus [sic] Tetrasticta KRAATZ (Coleoptera, Staphylinidae, Aleocharinae) from Xishuangbanna, Southwest China" type="journal article" year="2014">ZHENG &amp; ZHAO (2014)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="69DF653B6A73E97CFF0CB0DCFC331378" pageId="4" pageNumber="233" type="distribution">
<paragraph id="217A36B06A73E97CFF0CB0DCFC331378" blockId="4.[178,1085,492,572]" pageId="4" pageNumber="233">
Distribution and natural history: This species is currently known from two localities in 
<collectingRegion id="E301F8526A73E97CFE5EB339FD85135B" box="[480,586,521,543]" country="Laos" name="Phongsali" pageId="4" pageNumber="233">Phongsaly</collectingRegion>
and Oudom Xai provinces, North 
<collectingCountry id="59D276206A73E97CFC12B339FC12135A" box="[940,989,521,542]" name="Laos" pageId="4" pageNumber="233">Laos</collectingCountry>
(
<figureCitation id="B9FE2A356A73E97CFC55B339FBE1135B" box="[1003,1070,521,543]" captionStart="Map 1" captionStartId="3.[178,223,1064,1085]" captionTargetBox="[178,1054,211,1054]" captionTargetId="figure-1@3.[178,1054,211,1054]" captionTargetPageId="3" captionText="Map 1: Distribution of Tetrasticta laotica nov.sp." figureDoi="http://doi.org/10.5281/zenodo.5416219" httpUri="https://zenodo.org/record/5416219/files/figure.png" pageId="4" pageNumber="233">Map 1</figureCitation>
). The 
<typeStatus id="FE7E88126A73E97CFF5EB317FEC31378" box="[224,268,551,572]" pageId="4" pageNumber="233">type</typeStatus>
specimens were collected at altitudes of approximately 1100 and 
<quantity id="E63D9B556A73E97CFC16B316FC371378" box="[936,1016,550,572]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="233" unit="m" value="1500.0">1500 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>