152 lines
17 KiB
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152 lines
17 KiB
XML
<document id="EB3CE5E6844881C139B7045E65CCA864" ID-CLB-Dataset="28466" ID-DOI="http://dx.doi.org/10.3897/dez.65.27424" ID-GBIF-Dataset="290e099d-cb77-4ad7-975e-32ce4a51926f" ID-Pensoft-Pub="1860-1324-2-177" ID-ZooBank="A047B48DD161424FB8800428DCC5888A" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1860-1324-2-177" ModsDocOrigin="Deutsche Entomologische Zeitschrift 65 (2)" ModsDocTitle="Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M.lophoides (Chaudoir) species complex" checkinTime="1552154842744" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="978329A8C4E10E907B23378CE17F27CB" docLanguage="en" docName="DeutEntomolZeit 65(2): 177-224" docOrigin="Deutsche Entomologische Zeitschrift 65 (2)" docSource="http://dx.doi.org/10.3897/dez.65.27424" docTitle="Mecyclothorax punctatus" docType="treatment" docVersion="7" lastPageNumber="177" masterDocId="FFB33A56FFD86772FFAAFFA9FF882245" masterDocTitle="Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex" masterLastPageNumber="224" masterPageNumber="177" pageNumber="177" updateTime="1701378747773" updateUser="ExternalLinkService">
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<mods:title id="6054FEE26E5AA8CBF61253EF9A46BD47">Taxonomic review of Australian Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) with special emphasis on the M. lophoides (Chaudoir) species complex</mods:title>
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<mods:namePart id="42CF154B5930229CD0A71A68F2D1BCFF">Liebherr, James K.</mods:namePart>
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<mods:title id="C41CD1674CC4C07643D16B8623F9317B">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="EC3121C6A78151878C51A74927E2CAC1">2018</mods:date>
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<mods:number id="C45F12B374371C457A762C332B7B630C">65</mods:number>
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<mods:url id="8CAAC9EBC0D6E21FC72A6A7333991D37">http://dx.doi.org/10.3897/dez.65.27424</mods:url>
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<mods:classification id="82FACCB866085C8C81E3B5D580278D35">journal article</mods:classification>
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<mods:identifier id="52C3B4F4D335A138DB74010B8C3A26C5" type="DOI">http://dx.doi.org/10.3897/dez.65.27424</mods:identifier>
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<treatment id="978329A8C4E10E907B23378CE17F27CB" ID-GBIF-Taxon="154365983" LSID="urn:lsid:plazi:treatment:978329A8C4E10E907B23378CE17F27CB" httpUri="http://treatment.plazi.org/id/978329A8C4E10E907B23378CE17F27CB" lastPageNumber="177" pageId="0" pageNumber="177">
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<subSubSection id="642DC4B1AFD2A9417CD59CF882F0C6A4" pageId="0" pageNumber="177" type="nomenclature">
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<paragraph id="4790185FA72F4BB75C10F7E090C12E5E" pageId="0" pageNumber="177">
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<taxonomicName id="A606064409A05128D51930453D952D05" ID-CoL="8MNLN" authority="Sloane" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax punctatus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="punctatus">Mecyclothorax punctatus (Sloane)</taxonomicName>
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Figures 2B, 3B, 7C, 8B, 9B, 10B, 11A
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</paragraph>
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</subSubSection>
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<subSubSection id="CAC3438CA1955B33E6E387EC2FF94EBA" pageId="0" pageNumber="177" type="reference_group">
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<paragraph id="6D901C48352FD9E7637E038BB4B096CB" pageId="0" pageNumber="177">
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<taxonomicName id="B083ACD1854B23143B98F4D4EEAA5808" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax punctatus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="punctatus">Cyclothorax punctatus</taxonomicName>
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Sloane, 1895: 449.
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</paragraph>
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<paragraph id="C949B3D1122DEBC1297AF90948A95214" pageId="0" pageNumber="177">
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<taxonomicName id="D361083EE8B7A3D2593CB23C75F8A454" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax punctatus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="punctatus">Mecyclothorax punctatus</taxonomicName>
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, Sloane, 1903: 585;
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<bibRefCitation id="D8FD22869655F17AE3915F2298391D93" author="Moore, BP" journalOrPublisher="Journal of the Australian Entomological Society" pageId="0" pageNumber="177" pagination="161 - 166" title="Taxonomic notes on some Australasian Mecyclothorax Sharp (Coleoptera: Carabidae: Psydrinae) and descriptions of new species." url="https://doi.org/10.1111/j.1440-6055.1984.tb01935.x" volume="23" year="1984">Moore 1984</bibRefCitation>
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: 163, fig. 12.
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</paragraph>
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<paragraph id="D0915209FEA819C8F199EA0B232A27DB" pageId="0" pageNumber="177">
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<taxonomicName id="A3252E2AA26617B94EDB69CF9AA58489" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax punctatus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="subSpecies" species="punctatus" subSpecies="peckorum">Mecyclothorax punctatus peckorum</taxonomicName>
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Baehr, 2016a: 94.
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</paragraph>
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</subSubSection>
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<subSubSection id="E8E6F215AC1D8820F4AC9584F98D688E" pageId="0" pageNumber="177" type="diagnosis">
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<paragraph id="6B8F2BB7D1BA4C69CC005F77DBA9C06E" pageId="0" pageNumber="177">Diagnosis</paragraph>
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<paragraph id="21492601A15306F5549F3257A5F4A36C" pageId="0" pageNumber="177">
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(n = 5). This species is most similar to
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<taxonomicName id="2AC5C86CE0860A6712DAF76A2EB48611" lsidName="M. moorei" pageId="0" pageNumber="177" rank="species" species="moorei">M. moorei</taxonomicName>
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, with several diagnostic characters differentiating the two listed there. The eyes are large and convex, largely covering the ocular lobes; EyL/OLL = 0.92-0.96. The elytra are broad, with broadly rounded humeri (Fig. 3B). The single available female specimen assignable to
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<taxonomicName id="0B6421DB2B5FED9F312A4EC2492A34B3" lsidName="M. p." pageId="0" pageNumber="177" rank="subSpecies" species="p." subSpecies="peckorum">M. p. peckorum</taxonomicName>
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has the elytral base broad, slightly different from the nominate form. However the pronotal and elytral strial punctures are deeper and less dense in
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<taxonomicName id="F3014D8A317F2877837C94E39090068F" lsidName="M. p." pageId="0" pageNumber="177" rank="subSpecies" species="p." subSpecies="peckorum">M. p. peckorum</taxonomicName>
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-as reported in
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<bibRefCitation id="128F9B09328D6445DE4D2CB72A6B91EF" author="Baehr, M" journalOrPublisher="Spixiana" pageId="0" pageNumber="177" pagination="93 - 97" title="A new subspecies of Mecyclothoraxpunctatus (Sloane) from south-western Australia." url="http://www.zobodat.at/pdf/Spixiana_039_0093-0097.pdf" volume="39" year="2016 a">Baehr (2016a)</bibRefCitation>
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-resulting in recognition of the two forms here. In addition, the parascutellar seta is much less in evidence among individuals of this species. When present it is short and fine, and it is certainly absent from the majority of examined specimens. Standardized body length 2.7-3.2 mm. Setal formula ++/++/
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<normalizedToken id="777196E14334055D032A193587B32DA2" originalValue="±">+/-</normalizedToken>
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2++.
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</paragraph>
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<caption id="0DC37D835D0AD82122A1F04A1A7EB8EB" pageId="0" pageNumber="177">
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<paragraph id="59A46FEA4F51D1F8FDB962E4F52A1797" pageId="0" pageNumber="177">
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Figure 3.
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<taxonomicName id="D108A40A8E8712F08383124D44B36E29" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Eucyclothorax)" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="subGenus" subGenus="Eucyclothorax">Mecyclothorax (Eucyclothorax)</taxonomicName>
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spp., dorsal view: A,
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<taxonomicName id="72EB7E94B47394A40597308E777C1703" lsidName="M. moorei" pageId="0" pageNumber="177" rank="species" species="moorei">M. moorei</taxonomicName>
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; B,
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<taxonomicName id="FC0E658F30DCEC94F54FA1926041B6D0" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
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, NSW; C,
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<taxonomicName id="4E5FC60E6A5D8658FFDD4567334BA06B" lsidName="M. curtus" pageId="0" pageNumber="177" rank="species" species="curtus">M. curtus</taxonomicName>
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; D,
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<taxonomicName id="C349B829FF3FF5F3AE34BC22E126B546" lsidName="M. blackburni" pageId="0" pageNumber="177" rank="species" species="blackburni">M. blackburni</taxonomicName>
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; E,
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<taxonomicName id="A33CA3383091C461151C21C40420568F" lsidName="M. isolatus" pageId="0" pageNumber="177" rank="species" species="isolatus">M. isolatus</taxonomicName>
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holotype.
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</paragraph>
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</caption>
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<paragraph id="1A744459E9B3EFC4F4EBA2EC04AD6865" pageId="0" pageNumber="177">
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Male genitalia (n = 1). Aedeagal median lobe apex narrowly rounded and not projected beyond apical ostial margin (Fig. 7C); right paramere narrow, elongate, with ~10 setae along the ventral margin, 2 apical setae, and the dorsal margin with 3 small setae (Fig. 8B); left paramere broadly expanded basally, its dorsal margin convex, the apical 1/3 of length very narrow in contrast to broad base.
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<bibRefCitation id="C893FC593B12EDD8F6972DA1EBABD3D8" author="Baehr, M" journalOrPublisher="Spixiana" pageId="0" pageNumber="177" pagination="93 - 97" title="A new subspecies of Mecyclothoraxpunctatus (Sloane) from south-western Australia." url="http://www.zobodat.at/pdf/Spixiana_039_0093-0097.pdf" volume="39" year="2016 a">Baehr (2016a</bibRefCitation>
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, fig. 2) documents a short leftward expansion on the median lobe apex of
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<taxonomicName id="2485BDA82737CD7DC65948124D2F2505" lsidName="M. p." pageId="0" pageNumber="177" rank="subSpecies" species="p." subSpecies="peckorum">M. p. peckorum</taxonomicName>
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. The parameral setation of the two subspecific forms is identical (Fig. 8B;
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<bibRefCitation id="539B19AF20DC01A27EF45F845A2506AF" author="Baehr, M" journalOrPublisher="Spixiana" pageId="0" pageNumber="177" pagination="93 - 97" title="A new subspecies of Mecyclothoraxpunctatus (Sloane) from south-western Australia." url="http://www.zobodat.at/pdf/Spixiana_039_0093-0097.pdf" volume="39" year="2016 a">Baehr 2016a</bibRefCitation>
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, fig. 2).
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</paragraph>
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<paragraph id="A5267AAF3A2BA52E36EED88276701810" pageId="0" pageNumber="177">
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Female reproductive tract (n = 1). Bursa copulatrix elongate, columnar (Fig. 9B); helminthoid sclerite elongate; spermathecal duct basally expanded and sclerotized, the more apical portion of duct membranous and of lesser diameter; basal gonocoxite with 3 larger setae laterally along apical margin (Fig. 10B); apical gonocoxite broad basally, breadth more than half length; lateral ensiform setae broad, elongate, length about 0.5
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<normalizedToken id="5F70F5D50C6CFA9B52AF5BB5E10976BF" originalValue="×">x</normalizedToken>
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length of apical gonocoxite, setal surface longitudinal striate; apical nematiform setae in subbasal sensory furrow.
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</paragraph>
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</subSubSection>
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<subSubSection id="889D12DE1BB6E078C6E37795ED780855" pageId="0" pageNumber="177" type="type information">
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<paragraph id="80B11C6B87694D92E960E95C3BB9500D" pageId="0" pageNumber="177">Type information.</paragraph>
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<paragraph id="7F73CE11860D1BF3357D4AAE307C513E" pageId="0" pageNumber="177">
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For
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<taxonomicName id="94238F68F4EDB8688D33B9807AAA4B95" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="species" species="punctatus">M. punctatus</taxonomicName>
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, Lectotype female (SAMA) hereby designated: white card with
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<normalizedToken id="FE9455649C9F32450E55E2096B00544B" originalValue="“5104”">"5104"</normalizedToken>
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and
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<normalizedToken id="0E68B970E1A5A632C756FF4E3575B609" originalValue="“N.S.W.”">"N.S.W."</normalizedToken>
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in red ink // N.S. Wales //
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<taxonomicName id="F6B251838E7C0929251449252062E04F" class="Insecta" family="Carabidae" genus="Moriomorphina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Moriomorphina punctatus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="punctatus">punctatus</taxonomicName>
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Sloane // J. 7252 /
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<taxonomicName id="0B28014BDBC62FDA80571FEC65515970" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
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/
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<taxonomicName id="A5BF5B7DA26B77B225837866B00198DD" class="Insecta" family="Carabidae" genus="Moriomorphina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Moriomorphina punctatus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="punctatus">punctatus</taxonomicName>
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/ M.S. Wales / Cotype [red ink] // Lectotype ♀ /
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<taxonomicName id="16B333A2F91B1A8812A5767BA5477681" class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclothorax" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="genus">Cyclothorax</taxonomicName>
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/
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<taxonomicName id="DA03797A11F780F5967DD11CFAC0FA84" class="Insecta" family="Carabidae" genus="Moriomorphina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Moriomorphina punctatus" order="Coleoptera" pageId="0" pageNumber="177" phylum="Arthropoda" rank="species" species="punctatus">punctatus</taxonomicName>
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Sloane / det. J.K. Liebherr 2004 [red label with black border]. For
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<taxonomicName id="4F0E8B2091CC4D23A1557FAABB4B3758" lsidName="M. punctatus" pageId="0" pageNumber="177" rank="subSpecies" species="punctatus" subSpecies="peckorum">M. punctatus peckorum</taxonomicName>
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, Holotype male (ANIC, not seen): Pemberton Warren N.P. WA 12 Jul. 1980 S. & J. Peck SBP106 // Berlesate karri litter and moss (
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<bibRefCitation id="DD3BAA1B254DBADCCD0D73B9D1398FD9" author="Baehr, M" journalOrPublisher="Spixiana" pageId="0" pageNumber="177" pagination="93 - 97" title="A new subspecies of Mecyclothoraxpunctatus (Sloane) from south-western Australia." url="http://www.zobodat.at/pdf/Spixiana_039_0093-0097.pdf" volume="39" year="2016 a">Baehr 2016a</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="3C5A7CD3374B054ABEEC8F825D525616" pageId="0" pageNumber="177" type="distribution">
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<paragraph id="405A94FAD6F5979533EDD8EA14EDE429" pageId="0" pageNumber="177">Distribution and habitat.</paragraph>
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<paragraph id="F0AC752D84C30D9743930E7ED2BBA06F" pageId="0" pageNumber="177">
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Following
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<normalizedToken id="4E6FC52D97DA3BFF408BFDA516DAA63D" originalValue="Baehrʼs">Baehrʼs</normalizedToken>
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(2016a) circumscription of this species, it is distributed across an easterly set of populations in southern New South Wales and northern Victoria (Fig. 11A), plus a disjunct set of western localities in the southwest region of Western Australia. Sloane wrote of the type series, "I have found it in considerable numbers under logs and the leaves of saplings at a place about twenty-five miles north-west from the town of Urana. It is sluggish in its movements (Sloane, 1895: 449
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<normalizedToken id="38502277EB242470FD11EB980ABEE879" originalValue="–450).”">-450)."</normalizedToken>
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Thus the type locality is very close to one of the localities where he collected types of
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<taxonomicName id="AB5EB6105D0420FC028197390C4A31A4" lsidName="M. cordicollis" pageId="0" pageNumber="177" rank="species" species="cordicollis">M. cordicollis</taxonomicName>
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(
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<bibRefCitation id="32D9FCD4F1C3CAEFBBD12CB5AA1E00C1" author="Sloane, TG" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="0" pageNumber="177" pagination="553 - 584" title="Studies in Australian Entomology. no. IX. New species of Carabidae (with notes on some previously described species, and synoptic lists of species)." url="https://doi.org/10.5962/bhl.part.7682" volume="24" year="1900">Sloane 1900</bibRefCitation>
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). More recently, specimens from the western localities have been collected via litter sifting of jarrah forest litter (
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<taxonomicName id="87FC4CDD238E1021C5AD98414C537D5B" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus marginata" order="Myrtales" pageId="0" pageNumber="177" phylum="Tracheophyta" rank="species" species="marginata">Eucalyptus marginata</taxonomicName>
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Donn ex Sm.). All specimens are vestigially winged.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |