888 lines
132 KiB
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888 lines
132 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.128.1804" ID-PMC="PMC3175138" ID-Pensoft-Pub="1313-2970-128-15" ID-Pensoft-UUID="6157FFBE34536B410C79CB5C11170069" ID-PubMed="21998550" ID-Zenodo-Dep="577176" ID-ZooBank="837E761ADDC84351B1080F9B9CDC7ED1" ModsDocID="1313-2970-128-15" checkinTime="1451249919653" checkinUser="pensoft" docAuthor="Valim, Ye, Kavanaugh, David H., Shi, Hongliang & Liang, Hongbin" docDate="2011" docId="63D9213A5708F24D68739AE3A3E9CCAF" docLanguage="en" docName="ZooKeys 128: 15-52" docOrigin="ZooKeys 128" docPubDate="2011-09-09" docSource="http://dx.doi.org/10.3897/zookeys.128.1804" docTitle="Lithochlaenius (Kryzhanovskij) formosanus subsp. 1976 Kryzhanovskij 1976" docType="treatment" docVersion="5" id="6157FFBE34536B410C79CB5C11170069" lastPageNumber="19" masterDocId="6157FFBE34536B410C79CB5C11170069" masterDocTitle="A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species" masterLastPageNumber="52" masterPageNumber="15" pageNumber="16" updateTime="1668152074611" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A key to species of subgenus Lithochlaenius (Coleoptera, Carabidae, Chlaeniini, Chlaenius), with descriptions of three new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Valim, Ye</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China & Key Laboratory for Plant Pests Management of Mountainous Region, Institute of Entomology, Guizhou, University, Guiyang 550025, China</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Science, San Francisco, California 94118, U. S. A.</mods:affiliation>
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<mods:nameIdentifier type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Shi, Hongliang</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2011</mods:date>
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<mods:number>2011-09-09</mods:number>
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<mods:detail type="volume">
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<mods:number>128</mods:number>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>52</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.128.1804</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.128.1804</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-128-15</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577176</mods:identifier>
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<treatment ID-GBIF-Taxon="152030676" LSID="urn:lsid:plazi:treatment:63D9213A5708F24D68739AE3A3E9CCAF" httpUri="http://treatment.plazi.org/id/63D9213A5708F24D68739AE3A3E9CCAF" lastPageId="4" lastPageNumber="19" pageId="1" pageNumber="16">
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<subSubSection pageId="1" pageNumber="16" type="nomenclature">
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<paragraph pageId="1" pageNumber="16">
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Subenus
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<taxonomicName LSID="63D9213A-5708-F24D-6873-9AE3A3E9CCAF" authority="Kryzhanovskij, 1976" authorityName="Kryzhanovskij" authorityYear="1976" class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="formosanus" subGenus="Kryzhanovskij" subSpecies="1976">Lithochlaenius Kryzhanovskij, 1976</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="16" type="reference_group">
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<paragraph pageId="1" pageNumber="16">
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<taxonomicName class="Insecta" family="Carabidae" genus="Hemichlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="genus">Hemichlaenius</taxonomicName>
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Lutshnik, 1933:169 (nec Bates, 1892:307). Type species
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<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="species" species="rambouseki">
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<emphasis italics="true" pageId="1" pageNumber="16">Chlaenius rambouseki</emphasis>
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</taxonomicName>
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Lutshnik, 1933; Kryzhanovskij, 1976:11
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</paragraph>
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<paragraph pageId="1" pageNumber="16">
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<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="genus">Lithochlaenius</taxonomicName>
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Kryzhanovskij, 1976:9. Type species
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<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="species" species="rambouseki">
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<emphasis italics="true" pageId="1" pageNumber="16">Chlaenius rambouseki</emphasis>
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</taxonomicName>
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Lutshnik, 1933; Morita, 1993:161
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</paragraph>
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<paragraph pageId="1" pageNumber="16">
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<taxonomicName authorityName="Kirschenhofer" authorityYear="1997" class="Insecta" family="Carabidae" genus="Agilochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="genus">Agilochlaenius</taxonomicName>
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Kirschenhofer, 1997:116. Type species
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<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="1" pageNumber="16" phylum="Arthropoda" rank="species" species="latro">
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<emphasis italics="true" pageId="1" pageNumber="16">Chlaenius latro</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="LaFerté-Sénectère">LaFerte-Senectere</normalizedToken>
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, 1851; Kirschenhofer, 2000:58
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16" type="diagnosis">
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<paragraph pageId="1" pageNumber="16">Diagnosis.</paragraph>
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<paragraph lastPageId="2" lastPageNumber="17" pageId="1" pageNumber="16">
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Antennomere 3 distinctly longer than antennomeres 1 and 2 combined (
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="1" pageNumber="16">Fig. 9</figureCitation>
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); pronotum distinctly cordate (
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="1" pageNumber="16">Fig. 1</figureCitation>
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), disk glabrous or sparsely pubescent, each hind angle with single seta (
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="1" pageNumber="16">Fig. 2</figureCitation>
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); scutellar setiferous pore puncture present (
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="1" pageNumber="16">Fig. 3</figureCitation>
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); venter densely pubescent laterally, sparsely pubescent or glabrous medially (
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="1" pageNumber="16">Fig. 4</figureCitation>
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);
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<pageBreakToken pageId="2" pageNumber="17" start="start">prosternal</pageBreakToken>
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intercoxal process punctate, bordered at apex; metepisterna long and narrow (
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Fig. 10</figureCitation>
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); basal margination of elytra incomplete, absent from medial portion; anterior tarsomere 4 short and deeply emarginate apically (more distinctly so in male,
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Figs 5-8</figureCitation>
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); aedeagus tubular, ejaculatory orifice long, extended to basal fifth of aedeagus, lamella short (
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Figs 11</figureCitation>
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,
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<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="2" pageNumber="17">117</figureCitation>
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-
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<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="2" pageNumber="17">145</figureCitation>
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); gonostylus of female ovipositor smooth, with one setiferous pore near apex and one slender spine at basal inner margin (
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<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Figs 12-15</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="17" type="description">
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<paragraph pageId="2" pageNumber="17">Description.</paragraph>
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<paragraph pageId="2" pageNumber="17">Length 12.0-18.0 mm, width 4.6-6.4 mm. Head and pronotum black, with green, blue or coppery metallic luster; elytra black, with blue or coppery luster in a few species; ventral surface black; legs black, brown or yellow; antennae yellow, brown, or dark brown.</paragraph>
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<paragraph pageId="2" pageNumber="17">
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Head with vertex nearly glabrous, or sparsely and coarsely punctate behind posterior level of eye; eyes moderately prominent; genae pubescent; antennae long, antennomere 1 (scape) coniform (
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<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="2" pageNumber="17">Figs 52-53, 57-59</figureCitation>
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), cylindrical (
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<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="2" pageNumber="17">Figs 56, 60</figureCitation>
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), or elongate-ovoid (
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<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="2" pageNumber="17">Figs 54-55, 61-65</figureCitation>
|
||
); antennomere 3 sparsely setose, distinctly longer than antennomeres 1 and 2 combined (
|
||
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Fig. 9</figureCitation>
|
||
); mandibles triangular, hooked at apex; labrum with six setigerous punctures near apex; maxillary and labial palpi cylindrical, slightly compressed at apex; maxillary palpi glabrous; penultimate labial palpomere with a few setae, apical labial palpomere glabrous; glossal sclerite with two subapical setae; tooth of mentum bifid or emarginate at apex, with one pair of setae near the base; gula glabrous, slightly rugose.
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="17">
|
||
Pronotum cordate (
|
||
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Fig. 1</figureCitation>
|
||
), widest at apical one-third, front angles obtuse, hind angles acute; lateral margins bordered; disk glabrous or sparsely pubescent, base longitudinally rugose, punctate; basal foveae small, deep, rugose, pubescent; hind angles each with one seta (
|
||
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="17">
|
||
Elytra moderately convex, oblong, with distinct isodiametric microsculpture at least laterally; scutellar striae long, with basal setiferous pore present (
|
||
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Fig. 3</figureCitation>
|
||
); at least outer intervals pubescent (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="2" pageNumber="17">Figs 80-93</figureCitation>
|
||
), pubescence on intervals 8 and 9 generally denser than on others; wings full-sized, functional.
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="17">
|
||
Abdominal sterna densely pubescent laterally, sparsely pubescent or glabrous medially (
|
||
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Fig. 4</figureCitation>
|
||
); sterna IV to VI with single long seta at each side; sternum VII with one seta at each side in male, two in female; apex of sternum VII more rounded in male than in female.
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="17">Pro- and mesosterna and pro-, mes-, and metepisterna densely pubescent; metasternum pubescent laterally, nearly glabrous medially; prosternal intercoxal process punctate and bordered at apex; scutellum triangular, glabrous.</paragraph>
|
||
<paragraph pageId="2" pageNumber="17">
|
||
Anterior femora without tooth; tarsomeres short, sparsely setose dorsally; anterior tarsomere 4 short and triangular, deeply emarginate (more so in male), with two rows of long setae ventrally (
|
||
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Figs 5-8</figureCitation>
|
||
); basal three anterior tarsomeres dilated in male.
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="17">
|
||
Male genitalia with aedeagus tubular, simple; ejaculatory orifice long (
|
||
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Fig. 11</figureCitation>
|
||
); apical lamella short, rounded or slightly truncated at apex. Gonostylus of female ovipositor smooth, with one setiferous pore near apex and a long slender spine at basal inner margin, outer margin without or with a very short spine (
|
||
<figureCitation captionStart="Figures 1–15" captionStartId="F1" captionText="Figures 1 - 15. Character states of Chlaenius (Lithochlaenius) spp. Figs 1 - 13 Chlaenius rambouseki Lutshnik 1 pronotum 2 site of hind angle seta 3 elytral base showing the site of basal pore 4 abdominal sterna showing the pubescence 5 male anterior tarsomere 4 in dorsal view 6 male anterior tarsomere 4 in ventral view 7 female anterior tarsomere 4 in dorsal view 8 female anterior tarsomere 4 in ventral view 9 antennomeres 1 - 5 showing antennomere 3 distinctly longer than 1 and 2 ones combined 10 metepisternum 11 aedeagus, showing the basic structure of the aedeagus in Lithochlaenius species 12 female gonostylus in ventral view 13 female gonostylus in lateral view 14 female gonostylus of Chlaenius propeagilis sp. n. in ventral view 15 female gonostylus of Chlaenius propeagilis sp. n. in lateral view." httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17">Figs 12-15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption httpUri="https://binary.pensoft.net/fig/17278" pageId="2" pageNumber="17" start="Figures 1–15" startId="F1">
|
||
<paragraph pageId="2" pageNumber="17">
|
||
<emphasis bold="true" pageId="2" pageNumber="17">Figures 1-15.</emphasis>
|
||
Character states of
|
||
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius (Lithochlaenius)</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis bold="true" pageId="2" pageNumber="17">Figs 1-13</emphasis>
|
||
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius rambouseki</emphasis>
|
||
</taxonomicName>
|
||
Lutshnik
|
||
<emphasis bold="true" pageId="2" pageNumber="17">1</emphasis>
|
||
pronotum
|
||
<emphasis bold="true" pageId="2" pageNumber="17">2</emphasis>
|
||
site of hind angle seta
|
||
<emphasis bold="true" pageId="2" pageNumber="17">3</emphasis>
|
||
elytral base showing the site of basal pore
|
||
<emphasis bold="true" pageId="2" pageNumber="17">4</emphasis>
|
||
abdominal sterna showing the pubescence
|
||
<emphasis bold="true" pageId="2" pageNumber="17">5</emphasis>
|
||
male anterior tarsomere 4 in dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">6</emphasis>
|
||
male anterior tarsomere 4 in ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">7</emphasis>
|
||
female anterior tarsomere 4 in dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">8</emphasis>
|
||
female anterior tarsomere 4 in ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">9</emphasis>
|
||
antennomeres 1-5 showing antennomere 3 distinctly longer than 1 and 2 ones combined
|
||
<emphasis bold="true" pageId="2" pageNumber="17">10</emphasis>
|
||
metepisternum
|
||
<emphasis bold="true" pageId="2" pageNumber="17">11</emphasis>
|
||
aedeagus, showing the basic structure of the aedeagus in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
<emphasis bold="true" pageId="2" pageNumber="17">12</emphasis>
|
||
female gonostylus in ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">13</emphasis>
|
||
female gonostylus in lateral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">14</emphasis>
|
||
female gonostylus of
|
||
<taxonomicName authorityName="Liu & Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="propeagilis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius propeagilis</emphasis>
|
||
</taxonomicName>
|
||
sp. n. in ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">15</emphasis>
|
||
female gonostylus of
|
||
<taxonomicName authorityName="Liu & Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="propeagilis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius propeagilis</emphasis>
|
||
</taxonomicName>
|
||
sp. n. in lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption httpUri="https://binary.pensoft.net/fig/17279" pageId="2" pageNumber="17" start="Figures 16–21" startId="F2">
|
||
<paragraph pageId="2" pageNumber="17">
|
||
<emphasis bold="true" pageId="2" pageNumber="17">Figures 16-21.</emphasis>
|
||
Habitus of
|
||
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius (Lithochlaenius)</emphasis>
|
||
</taxonomicName>
|
||
spp
|
||
<emphasis bold="true" pageId="2" pageNumber="17">16</emphasis>
|
||
<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="chuanqianensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius chuanqianensis</emphasis>
|
||
</taxonomicName>
|
||
sp. n., holotype, male, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">17</emphasis>
|
||
<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="chuanqianensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius chuanqianensis</emphasis>
|
||
</taxonomicName>
|
||
sp. n., holotype, male, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">18</emphasis>
|
||
<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="linwensini">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius linwensini</emphasis>
|
||
</taxonomicName>
|
||
sp. n., holotype, male, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">19</emphasis>
|
||
<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="linwensini">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius linwensini</emphasis>
|
||
</taxonomicName>
|
||
sp.n., holotype, male, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">20</emphasis>
|
||
<taxonomicName authorityName="Liu & Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="propeagilis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius propeagilis</emphasis>
|
||
</taxonomicName>
|
||
sp. n., holotype, male, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">21</emphasis>
|
||
<taxonomicName authorityName="Liu & Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="propeagilis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius propeagilis</emphasis>
|
||
</taxonomicName>
|
||
sp. n., holotype, male, ventral view. Scale line = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption httpUri="https://binary.pensoft.net/fig/17280" pageId="2" pageNumber="17" start="Figures 22–27" startId="F3">
|
||
<paragraph pageId="2" pageNumber="17">
|
||
<emphasis bold="true" pageId="2" pageNumber="17">Figures 22-27.</emphasis>
|
||
Habitus of
|
||
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius (Lithochlaenius)</emphasis>
|
||
</taxonomicName>
|
||
spp
|
||
<emphasis bold="true" pageId="2" pageNumber="17">22</emphasis>
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="agilis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius agilis</emphasis>
|
||
</taxonomicName>
|
||
Chaudoir, holotype, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">23</emphasis>
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="agilis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius agilis</emphasis>
|
||
</taxonomicName>
|
||
Chaudoir, holotype, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">24</emphasis>
|
||
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="formosensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius formosensis</emphasis>
|
||
</taxonomicName>
|
||
Lorenz, male, in CCCC, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">25</emphasis>
|
||
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="formosensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius formosensis</emphasis>
|
||
</taxonomicName>
|
||
Lorenz, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">26</emphasis>
|
||
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="leishanensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius leishanensis</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer, male, in IZCAS, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">27</emphasis>
|
||
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="leishanensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius leishanensis</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer, male, in IZCAS, ventral view. Scale line = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption httpUri="https://binary.pensoft.net/fig/17281" pageId="2" pageNumber="17" start="Figures 28–33" startId="F4">
|
||
<paragraph pageId="2" pageNumber="17">
|
||
<emphasis bold="true" pageId="2" pageNumber="17">Figures 28-33.</emphasis>
|
||
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius (Lithochlaenius)</emphasis>
|
||
</taxonomicName>
|
||
spp
|
||
<emphasis bold="true" pageId="2" pageNumber="17">28</emphasis>
|
||
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="noguchii">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius noguchii</emphasis>
|
||
</taxonomicName>
|
||
Bates, male, in IZCAS, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">29</emphasis>
|
||
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="noguchii">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius noguchii</emphasis>
|
||
</taxonomicName>
|
||
Bates, male, in IZCAS, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">30</emphasis>
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="wrasei">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius wrasei</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer, male, in IZCAS, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">31</emphasis>
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="wrasei">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius wrasei</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer, male, in IZCAS, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">32</emphasis>
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="agiloides">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius agiloides</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
, male, in IZCAS, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">33</emphasis>
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="agiloides">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius agiloides</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
, male, in IZCAS, ventral view. Scale line = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption httpUri="https://binary.pensoft.net/fig/17282" pageId="2" pageNumber="17" start="Figures 34–39" startId="F5">
|
||
<paragraph pageId="2" pageNumber="17">
|
||
<emphasis bold="true" pageId="2" pageNumber="17">Figures 34-39.</emphasis>
|
||
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius (Lithochlaenius)</emphasis>
|
||
</taxonomicName>
|
||
spp
|
||
<emphasis bold="true" pageId="2" pageNumber="17">34</emphasis>
|
||
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius rambouseki</emphasis>
|
||
</taxonomicName>
|
||
Lutshnik, male, collected from Ussuri region, in IZCAS, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">35</emphasis>
|
||
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius rambouseki</emphasis>
|
||
</taxonomicName>
|
||
Lutshnik, male, collected from Ussuri region, in IZCAS, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">36</emphasis>
|
||
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="anchomenoides">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius anchomenoides</emphasis>
|
||
</taxonomicName>
|
||
Bates, cotype, male, in BMNH, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">37</emphasis>
|
||
<taxonomicName authorityName="Bates" authorityYear="1889" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="anchomenoides">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius anchomenoides</emphasis>
|
||
</taxonomicName>
|
||
Bates, cotype, male, in BMNH, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">38</emphasis>
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="nuristanus">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius nuristanus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
, paratype, male, in MNHN, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">39</emphasis>
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="nuristanus">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius nuristanus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
, paratype, male, in MNHN, ventral view. Scale line = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption httpUri="https://binary.pensoft.net/fig/17283" pageId="2" pageNumber="17" start="Figures 40–45" startId="F6">
|
||
<paragraph pageId="2" pageNumber="17">
|
||
<emphasis bold="true" pageId="2" pageNumber="17">Figures 40-45.</emphasis>
|
||
<taxonomicName authority="" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="subGenus" subGenus="Lithochlaenius">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius (Lithochlaenius)</emphasis>
|
||
</taxonomicName>
|
||
spp
|
||
<emphasis bold="true" pageId="2" pageNumber="17">40</emphasis>
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="nuristanus">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius nuristanus</emphasis>
|
||
</taxonomicName>
|
||
a.
|
||
<emphasis italics="true" pageId="2" pageNumber="17">rubridipes</emphasis>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
, paratype, male, in MNHN, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">41</emphasis>
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1956" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="nuristanus">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius nuristanus</emphasis>
|
||
</taxonomicName>
|
||
a.
|
||
<emphasis italics="true" pageId="2" pageNumber="17">rubridipes</emphasis>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
, paratype, male, in MNHN, ventral view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">42</emphasis>
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="agiloides">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius agiloides</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
, holotype, male, in NMPC, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">43</emphasis>
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="formosanus">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius formosanus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
, holotype, male, in NMPC, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">44</emphasis>
|
||
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius rambouseki</emphasis>
|
||
</taxonomicName>
|
||
Lutshnik, from Taiwan, male, in IZCAS, dorsal view
|
||
<emphasis bold="true" pageId="2" pageNumber="17">45</emphasis>
|
||
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="2" pageNumber="17" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis italics="true" pageId="2" pageNumber="17">Chlaenius rambouseki</emphasis>
|
||
</taxonomicName>
|
||
Lutshnik, from Taiwan, male, in IZCAS, ventral view. Scale line = 5.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="3" lastPageNumber="18" pageId="2" pageNumber="17" type="distribution">
|
||
<paragraph pageId="2" pageNumber="17">Geographical Distribution.</paragraph>
|
||
<paragraph lastPageId="3" lastPageNumber="18" pageId="2" pageNumber="17">
|
||
China (Heilongjiang, Liaoning, Shaanxi, Gansu, Henan, Hubei, Hunan, Zhejiang, Fujian, Taiwan, Jiangxi, Guangdong, Hainan,
|
||
<pageBreakToken pageId="3" pageNumber="18" start="start">Guangxi</pageBreakToken>
|
||
, Guizhou, Sichuan, Yunnan, Tibet), North Korea, Japan, Russia (Far East), Afghanistan, Pakistan, India. The known localities of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
species are shown in
|
||
<figureCitation captionStart="Figure 158" captionStartId="F16" captionText="Figure 158. Map showing the known geographical distributions of Chlaenius (Lithochlaenius) spp." httpUri="https://binary.pensoft.net/fig/17293" pageId="3" pageNumber="18">Fig. 158</figureCitation>
|
||
. Based on the work of
|
||
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Government of India Central Publication Branch, Calcutta" pageId="23" pageNumber="38" refId="B1" refString="Andrewes, HE, 1930. Catalogue of Indian Insects. Part 18-Carabidae. Government of India Central Publication Branch, Calcutta, 389 pp." title="Catalogue of Indian Insects. Part 18 - Carabidae" year="1930">Andrewes (1930)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Paik, JK" journalOrPublisher="Journal of Asia-Pacific Entomology" pageId="23" pageNumber="38" pagination="85 - 105" refId="B23" refString="Paik, JK, Trac, DH, Will, K, 2006. Carabidae from Vietnam (Coleoptera). Journal of Asia-Pacific Entomology 9: 85 - 105" title="Carabidae from Vietnam (Coleoptera)." volume="9" year="2006">Paik et al. (2006)</bibRefCitation>
|
||
, members of this subgenus also occur in Indo-China (Laos, Cambodia, Vietnam), but we have not studied specimens from those countries.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="18" type="biology">
|
||
<paragraph pageId="3" pageNumber="18">Biology.</paragraph>
|
||
<paragraph pageId="3" pageNumber="18">
|
||
Members of this subgenus are typically collected on sandy beaches of rivers or streams (
|
||
<figureCitation captionStart="Figures 146–151" captionStartId="F14" captionText="Figures 146 - 151. Photographs of habitats for Chlaenius (Lithochlaenius) spp. 146 Chlaenius chuanqianensis sp. n., Dabaitang, Xishui county, North Guizhou, China 147 Chlaenius anchomenoides Bates, Goorais valley, Pakistan (Provided by Dr. Muhammad Abbas in Pakistan Museum of Natural History) 148 Chlaenius propeagilis sp. n., Gaoligongshan, Yunnan, China 149 Chlaenius leishanensis Kirschenhofer, Xiaodanjiang, Leigongshan, Leishan county, Southeast Guizhou, China 150 Chlaenius chuanqianensis sp. n. and Chlaenius agiloides Jedlicka, Jinshajiang, Sichuan 151 Chlaenius rambouseki Lutshnik, Ussri river, Heilongjiang, China." httpUri="https://binary.pensoft.net/fig/17291" pageId="3" pageNumber="18">Figs 146</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figures 152–157" captionStartId="F15" captionText="Figures 152 - 157. Photographs of habitats for Chlaenius (Lithochlaenius) spp. 152 Chlaenius rambouseki Lutshnik, Baisha county, Hainan Island, China 153 Chlaenius rambouseki Lutshnik, Yuanbaoshan, Guangxi, China 154 Chlaenius rambouseki Lutshnik adult walking on concrete wharf of Ussri river at night 155 Chlaenius agiloides Jedlicka adults mating at night 156 Assorted molluscs, show one food of Chlaenius rambouseki Lutshnik from Hainan Island 157 Adults of Chlaenius agiloides Jedlicka preying on dragonfly nymph at night." httpUri="https://binary.pensoft.net/fig/17292" pageId="3" pageNumber="18">153</figureCitation>
|
||
). Adults of some species (e. g.,
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="agiloides">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius agiloides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius rambouseki</emphasis>
|
||
</taxonomicName>
|
||
) have been observed feeding on mollusks, worms, and dragonfly larvae at night (
|
||
<figureCitation captionStart="Figures 152–157" captionStartId="F15" captionText="Figures 152 - 157. Photographs of habitats for Chlaenius (Lithochlaenius) spp. 152 Chlaenius rambouseki Lutshnik, Baisha county, Hainan Island, China 153 Chlaenius rambouseki Lutshnik, Yuanbaoshan, Guangxi, China 154 Chlaenius rambouseki Lutshnik adult walking on concrete wharf of Ussri river at night 155 Chlaenius agiloides Jedlicka adults mating at night 156 Assorted molluscs, show one food of Chlaenius rambouseki Lutshnik from Hainan Island 157 Adults of Chlaenius agiloides Jedlicka preying on dragonfly nymph at night." httpUri="https://binary.pensoft.net/fig/17292" pageId="3" pageNumber="18">Figs 154-157</figureCitation>
|
||
). A few species have also been collected in light traps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18" type="remarks">
|
||
<paragraph pageId="3" pageNumber="18">Remarks.</paragraph>
|
||
<paragraph pageId="3" pageNumber="18">
|
||
Based on the metallic body surface, single supraorbital seta, antennomere 3 longest and antennomeres 4-11 densely pubescent,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
species can be recognized as a member of the genus
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius</emphasis>
|
||
</taxonomicName>
|
||
. Members of this subgenus are similar to those of subgenus
|
||
<taxonomicName authorityName="Reitter" authorityYear="1908" class="Insecta" family="Carabidae" genus="Stenochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Stenochlaenius</emphasis>
|
||
</taxonomicName>
|
||
in shape of the pronotum, but the latter are much smaller and have a glabrous body surface.
|
||
</paragraph>
|
||
<paragraph pageId="3" pageNumber="18">
|
||
As presently conceived, the subgenus can be divided into two species groups: 1) the
|
||
<emphasis italics="true" pageId="3" pageNumber="18">rambouseki</emphasis>
|
||
group, members of which have all elytral intervals densely and more or less equally pubescent, and males have aedeagi slender in dorsal view; and 2) the
|
||
<emphasis italics="true" pageId="3" pageNumber="18">agilis</emphasis>
|
||
group, members of which have at least elytral intervals 1-5 glabrous medially with pubescence restricted to the strial depressions, intervals 6-9 densely pubescent, and males have aedeagi generally stouter in dorsal view. Most species of the subgenus have restricted geographical ranges, and only C.
|
||
<emphasis italics="true" pageId="3" pageNumber="18">rambouseki</emphasis>
|
||
is more broadly distributed.
|
||
</paragraph>
|
||
<paragraph lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18">
|
||
In his treatment of the North American species of genus
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation author="Bell, RT" journalOrPublisher="Miscellaneous Publications of the Entomological Society of America" pageId="23" pageNumber="38" pagination="97 - 166" refId="B5" refString="Bell, RT, 1960. A revision of the genus Chlaenius Bonelli (Coleoptera, Carabidae) in North America. Miscellaneous Publications of the Entomological Society of America 1: 97 - 166" title="A revision of the genus Chlaenius Bonelli (Coleoptera, Carabidae) in North America." volume="1" year="1960">Bell (1960)</bibRefCitation>
|
||
suggested that his
|
||
<emphasis italics="true" pageId="3" pageNumber="18">solitarius</emphasis>
|
||
species group, which included
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="cordicollis">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius cordicollis</emphasis>
|
||
</taxonomicName>
|
||
Kirby,
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="leucoscelis">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius leucoscelis</emphasis>
|
||
</taxonomicName>
|
||
Chevrolat,
|
||
<taxonomicName authorityName="Dejean" authorityYear="1826" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="prasinus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius prasinus</emphasis>
|
||
</taxonomicName>
|
||
Dejean,
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="purpureus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius purpureus</emphasis>
|
||
</taxonomicName>
|
||
Chaudoir, and
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="solitarius">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius solitarius</emphasis>
|
||
</taxonomicName>
|
||
Say, represented a distinct group within his subgenus
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius</emphasis>
|
||
</taxonomicName>
|
||
sensu stricto. He noted that the range of this group extends south into South America and that "Related forms occur in the Old World". In the paper in which he proposed
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
as a replacement name for
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Hemichlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Hemichlaenius</emphasis>
|
||
</taxonomicName>
|
||
Lutshnik,
|
||
<bibRefCitation author="Kryzhanovskij, OL" journalOrPublisher=")" pageId="23" pageNumber="38" pagination="8 - 17" refId="B16" refString="Kryzhanovskij, OL, 1976. A survey of the Callistini tribe (Coleoptera, Carabidae) in the Far East. Proceedings of the Institute of Soil Biology (N.S.) 43 (146): 8 - 17" title="A survey of the Callistini tribe (Coleoptera, Carabidae) in the Far East. Proceedings of the Institute of Soil Biology (N. S." volume="43" year="1976">Kryzhanovskij (1976: 16)</bibRefCitation>
|
||
cited Bell's paper and suggested that
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="solitarius">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Chlaenius solitarius</emphasis>
|
||
</taxonomicName>
|
||
might be a North American representative of that subgenus. Robert Davidson (personal communication) shares the view that species of Bell's
|
||
<emphasis italics="true" pageId="3" pageNumber="18">solitarius</emphasis>
|
||
group, and at least five additional species in Middle and South America, are likely related to some if not all
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
species. All of these New World species share most of the diagnostic features of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
and all of them have elytral pubescence as seen in members of the
|
||
<emphasis italics="true" pageId="3" pageNumber="18">rambouseki</emphasis>
|
||
group. However, members of all these New World species are distinguished in having the lateral and basal elytral margins smoothly continuous around the humeri without forming any trace of an angle and the elytral epipleura and basal regions smoothly continuous around the humeri, not separated by a carina of any kind. These were the main features that Bell used to distinguish members of his
|
||
<emphasis italics="true" pageId="3" pageNumber="18">solitarius</emphasis>
|
||
group. In contrast, members of all the Asian species of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
that we have studied have a distinct humeral angle formed at the junction of the lateral and basal elytral margins and a more or less distinctly carinate separation of the (lateral) epipleural from the basal (anterior vertical) elytral surfaces. Hence, Asian
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="3" pageNumber="18">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
specimens could not be identified as members of the
|
||
<emphasis italics="true" pageId="3" pageNumber="18">solitarius</emphasis>
|
||
groups using Bell's (1960) key. There is also greater variation in the development of the elytral basal margin (from complete to partially interrupted) and in the length and shape of the apica
|
||
<pageBreakToken pageId="4" pageNumber="19" start="start">l</pageBreakToken>
|
||
lamella of the male aedeagus among New World species than we have seen among the Asian
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
species. Consequently, we suggest that placement of any New World species in subgenus
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Lithochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Lithochlaenius</emphasis>
|
||
</taxonomicName>
|
||
would be premature at this time and should await a more comprehensive, worldwide treatment of genus
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius</emphasis>
|
||
</taxonomicName>
|
||
and analyses of phylogenetic relationships among the included species, species groups, and subgenera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="19" type="key to the species of subgenus lithochlaenius">
|
||
<paragraph pageId="4" pageNumber="19">Key to the species of subgenus Lithochlaenius</paragraph>
|
||
<paragraph pageId="4" pageNumber="19">
|
||
<table inLine="true" pageId="4" pageNumber="19">
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">1</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
All intervals punctate, pubescent, slightly convex (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Figs 88-89, 93</figureCitation>
|
||
); aedeagus slender, basal portion rugose (
|
||
<emphasis italics="true" pageId="4" pageNumber="19">rambouseki</emphasis>
|
||
group)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<emphasis bold="true" pageId="4" pageNumber="19">2</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
At least basal half of intervals 1-5 smooth and glabrous medially, distinctively convex (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Figs 80-87, 90-92</figureCitation>
|
||
); aedeagus usually stout, basal portion smooth (
|
||
<emphasis italics="true" pageId="4" pageNumber="19">agilis</emphasis>
|
||
group)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">3</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">2</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Intervals dull, with dense and regular punctures (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Figs 89, 93</figureCitation>
|
||
); antennomere 1 elongate ovoid (
|
||
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">Figs 61, 65</figureCitation>
|
||
); apical lamella of aedeagus thin (
|
||
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="4" pageNumber="19">Figs 137, 145</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName authorityName="Lutshnik" authorityYear="1933" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="rambouseki">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius rambouseki</emphasis>
|
||
</taxonomicName>
|
||
Lutshnik
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Intervals shining, with sparse and irregular punctures (large and small punctures mixed) (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Fig. 88</figureCitation>
|
||
); antennomere 1 cylindrical (
|
||
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">Fig. 60</figureCitation>
|
||
); apical lamella of aedeagus thick (
|
||
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="4" pageNumber="19">Figs 134-135</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="agiloides">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius agiloides</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">3</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Antennomere 1 with apical end much thicker than basal end, coniform (
|
||
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">Figs 52-53, 57-59</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">4</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Antennomere 1 with apical end as thick as basal end, elongate-ovoid (
|
||
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">Figs 54, 55, 62-64</figureCitation>
|
||
) or cylindrical (
|
||
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">Fig. 56</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">8</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">4</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Intervals 1-7 glabrous medially, with a row of pubescence laterally (near striae) (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Fig. 86</figureCitation>
|
||
); legs dark brown or brown (
|
||
<figureCitation captionStart="Figures 28–33" captionStartId="F4" captionText="Figures 28 - 33. Chlaenius (Lithochlaenius) spp 28 Chlaenius noguchii Bates, male, in IZCAS, dorsal view 29 Chlaenius noguchii Bates, male, in IZCAS, ventral view 30 Chlaenius wrasei Kirschenhofer, male, in IZCAS, dorsal view 31 Chlaenius wrasei Kirschenhofer, male, in IZCAS, ventral view 32 Chlaenius agiloides Jedlicka, male, in IZCAS, dorsal view 33 Chlaenius agiloides Jedlicka, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17281" pageId="4" pageNumber="19">Fig. 29</figureCitation>
|
||
); apex of sternum VII subtruncate (
|
||
<figureCitation captionStart="Figures 94–116" captionStartId="F11" captionText="Figures 94 - 116. Abdominal sternum VII (ventral aspect) of Chlaenius (Lithochlaenius) species 94 Chlaenius chuanqianensis sp. n., male, holotype 95 Chlaenius chuanqianensis sp. n., female, paratype 96 Chlaenius linwensini sp. n., male, holotype 97 Chlaenius linwensini sp. n., female, paratype 98 Chlaenius propeagilis sp. n., male, holotype 99 Chlaenius propeagilis sp. n., female, paratype 100 Chlaenius agilis Chaudoir, male, holotype 101 Chlaenius formosensis Lorenz, male 102 Chlaenius formosensis Lorenz, female 103 Chlaenius leishanensis Kirschenhofer, male 104 Chlaenius leishanensis Kirschenhofer, female 105 Chlaenius noguchii Bates, male 106 Chlaenius noguchii Bates, female 107 Chlaenius wrasei Kirschenhofer, male 108 Chlaenius wrasei Kirschenhofer, female 109 Chlaenius agiloides Jedlicka, male 110 Chlaenius agiloides Jedlicka, female 111 Chlaenius rambouseki Lutshnik, from Ussuri region, male 112 Chlaenius rambouseki Lutshnik from Ussuri region, female 113 Chlaenius anchomenoides Bates, male, paratype 114 Chlaenius nuristanus Jedlicka, male, paratype 115 Chlaenius nuristanus Jedlicka, female, paratype 116 Chlaenius formosanus Jedlicka from Taiwan, male. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17288" pageId="4" pageNumber="19">Figs 105, 106</figureCitation>
|
||
); aedeagus slender, depressed, apical lamella bent ventrally (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Figs 129-130</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName authorityName="Bates" authorityYear="1873" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="noguchii">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius noguchii</emphasis>
|
||
</taxonomicName>
|
||
Bates
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Intervals 6-7 pubescent, intervals 1-5 glabrous medially, with a row of pubescence laterally (near striae) (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Figs 80-81, 85, 87</figureCitation>
|
||
); aedeagus stout
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">5</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">5</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Tibiae and femora bi-colored, with tibiae black or dark brown (
|
||
<figureCitation captionStart="Figures 22–27" captionStartId="F3" captionText="Figures 22 - 27. Habitus of Chlaenius (Lithochlaenius) spp 22 Chlaenius agilis Chaudoir, holotype, dorsal view 23 Chlaenius agilis Chaudoir, holotype, ventral view 24 Chlaenius formosensis Lorenz, male, in CCCC, dorsal view 25 Chlaenius formosensis Lorenz, ventral view 26 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, dorsal view 27 Chlaenius leishanensis Kirschenhofer, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17280" pageId="4" pageNumber="19">Figs 26-27</figureCitation>
|
||
), femora yellow or light brown; lamella of aedeagus rounded at apex (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 127</figureCitation>
|
||
), thickened, and bent ventrally (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 128</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName authorityName="Kirschenhofer" authorityYear="2005" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="leishanensis">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius leishanensis</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Tibiae and femora concolorous, yellow or brown (
|
||
<figureCitation captionStart="Figures 16–21" captionStartId="F2" captionText="Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17279" pageId="4" pageNumber="19">Figs 16-19</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 28–33" captionStartId="F4" captionText="Figures 28 - 33. Chlaenius (Lithochlaenius) spp 28 Chlaenius noguchii Bates, male, in IZCAS, dorsal view 29 Chlaenius noguchii Bates, male, in IZCAS, ventral view 30 Chlaenius wrasei Kirschenhofer, male, in IZCAS, dorsal view 31 Chlaenius wrasei Kirschenhofer, male, in IZCAS, ventral view 32 Chlaenius agiloides Jedlicka, male, in IZCAS, dorsal view 33 Chlaenius agiloides Jedlicka, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17281" pageId="4" pageNumber="19">30-31</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">6</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">6</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
All antennomeres brown or dark brown (
|
||
<figureCitation captionStart="Figures 16–21" captionStartId="F2" captionText="Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17279" pageId="4" pageNumber="19">Fig. 18</figureCitation>
|
||
); hind trochanters brown, nearly the same color as hind femora (
|
||
<figureCitation captionStart="Figures 16–21" captionStartId="F2" captionText="Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17279" pageId="4" pageNumber="19">Fig. 19</figureCitation>
|
||
); elytra with basal three-fourths of intervals 1-5 glabrous medially; lamella of aedeagus sub-truncate at apex (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 119</figureCitation>
|
||
), thick (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 120</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="linwensini">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius linwensini</emphasis>
|
||
</taxonomicName>
|
||
Liu & Liang, sp. n.
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
At least antennomere 1 yellow or brown, paler than the rest (
|
||
<figureCitation captionStart="Figures 16–21" captionStartId="F2" captionText="Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17279" pageId="4" pageNumber="19">Figs 16</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 28–33" captionStartId="F4" captionText="Figures 28 - 33. Chlaenius (Lithochlaenius) spp 28 Chlaenius noguchii Bates, male, in IZCAS, dorsal view 29 Chlaenius noguchii Bates, male, in IZCAS, ventral view 30 Chlaenius wrasei Kirschenhofer, male, in IZCAS, dorsal view 31 Chlaenius wrasei Kirschenhofer, male, in IZCAS, ventral view 32 Chlaenius agiloides Jedlicka, male, in IZCAS, dorsal view 33 Chlaenius agiloides Jedlicka, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17281" pageId="4" pageNumber="19">30</figureCitation>
|
||
); hind trochanters much darker than hind femora (
|
||
<figureCitation captionStart="Figures 16–21" captionStartId="F2" captionText="Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17279" pageId="4" pageNumber="19">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 40–45" captionStartId="F6" captionText="Figures 40 - 45. Chlaenius (Lithochlaenius) spp 40 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, dorsal view 41 Chlaenius nuristanus a. rubridipes Jedlicka, paratype, male, in MNHN, ventral view 42 Chlaenius agiloides Jedlicka, holotype, male, in NMPC, dorsal view 43 Chlaenius formosanus Jedlicka, holotype, male, in NMPC, dorsal view 44 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, dorsal view 45 Chlaenius rambouseki Lutshnik, from Taiwan, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17283" pageId="4" pageNumber="19">41</figureCitation>
|
||
); elytra with entire lengths of intervals 1-5 glabrous medially; lamella of aedeagus rounded at apex (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Figs 117</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="4" pageNumber="19">131</figureCitation>
|
||
), thin (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Figs 118</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="4" pageNumber="19">132</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">7</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">7</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Antennomeres 1-3 brown, color paler than antennomeres 4-11 (
|
||
<figureCitation captionStart="Figures 28–33" captionStartId="F4" captionText="Figures 28 - 33. Chlaenius (Lithochlaenius) spp 28 Chlaenius noguchii Bates, male, in IZCAS, dorsal view 29 Chlaenius noguchii Bates, male, in IZCAS, ventral view 30 Chlaenius wrasei Kirschenhofer, male, in IZCAS, dorsal view 31 Chlaenius wrasei Kirschenhofer, male, in IZCAS, ventral view 32 Chlaenius agiloides Jedlicka, male, in IZCAS, dorsal view 33 Chlaenius agiloides Jedlicka, male, in IZCAS, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17281" pageId="4" pageNumber="19">Fig. 30</figureCitation>
|
||
); apical lamella of aedeagus bent ventrally (
|
||
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="4" pageNumber="19">Fig. 132</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="wrasei">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius wrasei</emphasis>
|
||
</taxonomicName>
|
||
Kirschenhofer
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Only antennomere 1 yellow or brown, color paler than antennomeres 2-11 (
|
||
<figureCitation captionStart="Figures 16–21" captionStartId="F2" captionText="Figures 16 - 21. Habitus of Chlaenius (Lithochlaenius) spp 16 Chlaenius chuanqianensis sp. n., holotype, male, dorsal view 17 Chlaenius chuanqianensis sp. n., holotype, male, ventral view 18 Chlaenius linwensini sp. n., holotype, male, dorsal view 19 Chlaenius linwensini sp. n., holotype, male, ventral view 20 Chlaenius propeagilis sp. n., holotype, male, dorsal view 21 Chlaenius propeagilis sp. n., holotype, male, ventral view. Scale line = 5.0 mm." httpUri="https://binary.pensoft.net/fig/17279" pageId="4" pageNumber="19">Fig. 16</figureCitation>
|
||
); apical lamella of aedeagus straight (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 118</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName authorityName="Liu & Liang" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="chuanqianensis">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius chuanqianensis</emphasis>
|
||
</taxonomicName>
|
||
Liu & Liang, sp. n.
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">8</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Intervals 1-7 glabrous medially (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Fig. 84</figureCitation>
|
||
); antennomere 1 cylindrical (
|
||
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">Fig. 56</figureCitation>
|
||
); lamella of aedeagus subtruncate at apex (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 125</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName authorityName="Lorenz" authorityYear="1998" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="formosensis">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius formosensis</emphasis>
|
||
</taxonomicName>
|
||
Lorenz
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
At most intervals 1-5 glabrous medially (
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">Figs 82-83</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 80–93" captionStartId="F10" captionText="Figures 80 - 93. Elytral intervals of Chlaenius (Lithochlaenius) spp. 80 Chlaenius chuanqianensis sp. n., holotype 81 Chlaenius linwensini sp. n., holotype 82 Chlaenius propeagilis sp. n., holotype 83 Chlaenius agilis Chaudoir, holotype 84 Chlaenius formosensis Lorenz, CCCC 85 Chlaenius leishanensis, IZCAS 86 Chlaenius noguchii Bates, IZCAS 87 Chlaenius wrasei Kirschenhofer, IZCAS 88 Chlaenius agiloides Jedlicka, IZCAS 89 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 90 Chlaenius anchomenoides Bates, paratype 91 Chlaenius nuristanus Jedlicka, paratype 92 Chlaenius nuristanus Jedlicka, paratype 93 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17287" pageId="4" pageNumber="19">90-92</figureCitation>
|
||
); antennomere 1 elongate ovoid (
|
||
<figureCitation captionStart="Figures 52–65" captionStartId="F8" captionText="Figures 52 - 65. Antennomere 1 of Chlaenius (Lithochlaenius) species 52 Chlaenius chuanqianensis sp. n., holotype 53 Chlaenius linwensini sp. n., holotype 54 Chlaenius propeagilis sp. n., holotype 55 Chlaenius agilis Chaudoir, holotype 56 Chlaenius formosensis Lorenz, CCCC 57 Chlaenius leishanensis Kirschenhofer, IZCAS 58 Chlaenius noguchii Bates, IZCAS 59 Chlaenius wrasei Kirschenhofer, IZCAS 60 Chlaenius agiloides Jedlicka, IZCAS 61 Chlaenius rambouseki Lutshnik from Ussuri region, IZCAS 62 Chlaenius anchomenoides Bates, paratype 63 Chlaenius nuristanus Jedlicka, paratype 64 Chlaenius nuristanus a. rubridipes Jedlicka, paratype 65 Chlaenius rambouseki Lutshnik from Taiwan, CCCC. Scale line = 1.0 mm." httpUri="https://binary.pensoft.net/fig/17285" pageId="4" pageNumber="19">Figs 54-55, 62-64</figureCitation>
|
||
); lamella of aedeagus round at apex (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Figs 121, 123</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="4" pageNumber="19">138, 140, 142</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">9</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">9</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Male with media lobe of aedeagus gradually bent near base (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 122</figureCitation>
|
||
), lamella triangular at apex, left side of media lobe nearly straight in dorsal view (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Fig. 121</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
<taxonomicName authorityName="Liu & Kavanaugh" authorityYear="2011" class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="propeagilis">
|
||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius propeagilis</emphasis>
|
||
</taxonomicName>
|
||
Liu & Kavanaugh, sp. n.
|
||
</td>
|
||
</tr>
|
||
<tr pageId="4" pageNumber="19">
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">-</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
|
||
Male with media lobe of aedeagus abruptly bent near the base (
|
||
<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Figs 124</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="4" pageNumber="19">139, 141, 143</figureCitation>
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), lamella rounded at apex, left side of media lobe expand laterally in dorsal view (
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<figureCitation captionStart="Figures 117–130" captionStartId="F12" captionText="Figures 117 - 130. Aedeagi of Chlaenius (Lithochlaenius) species 117 Chlaenius chuanqianensis sp. n., holotype (dorsal view) 118 Chlaenius chuanqianensis sp. n., holotype (left lateral view) 119 Chlaenius linwensini sp. n., holotype (dorsal view) 120 Chlaenius linwensini sp. n., holotype (left lateral view) 121 Chlaenius propeagilis sp. n., holotype (dorsal view) 122 Chlaenius propeagilis sp. n., holotype (left lateral view) 123 Chlaenius agilis Chaudoir, holotype (dorsal view) 124 Chlaenius agilis Chaudoir, holotype (left lateral view) 125 Chlaenius formosensis Lorenz, holotype (dorsal view) 126 Chlaenius formosensis Lorenz, holotype (left lateral view) 127 Chlaenius leishanensis Kirschenhofer (dorsal view) 128 Chlaenius leishanensis Kirschenhofer (left lateral view) 129 Chlaenius noguchii Bates (dorsal view) 130 Chlaenius noguchii Bates (left lateral view). Scale lines: A = 1.0 mm (Figs 117, 119, 121, 123, 125, 127, 129); B = 0.5 mm (Figs 118, 120, 122, 124, 126, 128, 130)." httpUri="https://binary.pensoft.net/fig/17289" pageId="4" pageNumber="19">Figs 123</figureCitation>
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,
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<figureCitation captionStart="Figures 131–145" captionStartId="F13" captionText="Figures 131 - 145. Aedeagi of Chlaenius (Lithochlaenius) species 131 Chlaenius wrasei Kirschenhofer (dorsal view) 132 Chlaenius wrasei Kirschenhofer (left lateral view) 133 Chlaenius agiloides Jedlicka (dorsal view) 134 Chlaenius agiloides Jedlicka, holotype (left lateral view) 135 Chlaenius agiloides Jedlicka (left lateral view) 136 Chlaenius rambouseki Lutshnik from Ussuri region (dorsal view) 137 Chlaenius rambouseki Lutshnik from Ussuri region (left lateral view) 138 Chlaenius anchomenoides Bates, paratype (dorsal view) 139 Chlaenius anchomenoides Bates, paratype (left lateral view) 140 Chlaenius nuristanus Jedlicka, paratype (dorsal view, black leg) 141 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 142 Chlaenius nuristanus Jedlicka, paratype (dorsal view, yellow leg) 143 Chlaenius nuristanus Jedlicka, paratype (left lateral view) 144 Chlaenius rambouseki Lutshnik from Taiwan (dorsal view) 145 Chlaenius rambouseki Lutshnik from Taiwan (left lateral view). Scale lines: A = 1.0 mm (Figs 131, 133, 136, 138, 140, 142, 144); B = 0.5 mm (Figs 132, 134, 135, 137, 139, 141, 143, 145)." httpUri="https://binary.pensoft.net/fig/17290" pageId="4" pageNumber="19">138, 140, 142</figureCitation>
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)
|
||
</td>
|
||
<td colspan="1" pageId="4" pageNumber="19" rowspan="1">
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||
<taxonomicName class="Hexapoda" family="Carabidae" genus="Chlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="agilis">
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||
<emphasis italics="true" pageId="4" pageNumber="19">Chlaenius agilis</emphasis>
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||
</taxonomicName>
|
||
Chaudoir
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="19" type="theagilisgroup">
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||
<paragraph pageId="4" pageNumber="19">Theagilisgroup</paragraph>
|
||
</subSubSection>
|
||
</treatment>
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||
</document> |