treatments-xml/data/60/97/B1/6097B188B9DDC7833CE35BA10BFF00F7.xml
2024-06-21 12:38:05 +02:00

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XML

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<mods:title id="2788CD9950C17C3E697894F5CA34890A">Taxonomic Revision of the Ant-Acacias (Fabaceae, Mimosoideae, Acacia, Series Gummiferae) of the New World</mods:title>
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<mods:namePart id="EAF1848247BB9CF43334D82C6F771001">David S. Seigler</mods:namePart>
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<mods:namePart id="8A813A8BE48E8814B9434AB6FFC47A3D">John E. Ebinger</mods:namePart>
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1.
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,
<bibCitation id="9785DA793E955EDEC461BF8CA65030E3" journal="Smithsonian Contr. Bot." page="53" score="91.8138855147161" volume="13" year="1974">Smithsonian Contr. Bot. 13: 53. 1974</bibCitation>
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. TYPE:
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Osa Peninsula, bank of tributary of
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Agua Buena,
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. SW of Rincon
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)
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.
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Tree to
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tall with widely spreading branches, young twigs dark brown to dark reddish brown, lightly puberulent. Stipular spines black to dark brown, commonly grooved and with low, rounded longitudinal ridges, glabrous to lightly puberulent, nearly terete in cross section, sometimes slightly reflexed, symmetrical, broadly V-shaped with an angle of 90-120°, to
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long,
<quantity id="6375CE5A6B44480617399DFB2F226847" metricMagnitude="-3" metricUnit="m" metricValue="10.0" metricValueMax="12.0" metricValueMin="8.0" pageId="4" pageNumber="121" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8-12 mm</quantity>
wide near the base. Leaves
<quantity id="827C36C8EAF531C29D5B64ABC59EB24B" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="4" pageNumber="121" unit="mm" value="275.0" valueMax="400.0" valueMin="150.0">150-400 mm</quantity>
long, pinnae 10-22 pairs per leaf,
<quantity id="834B65DA6D1F3752F3F6D4D7CDF77766" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="4" pageNumber="121" unit="mm" value="85.0" valueMax="110.0" valueMin="60.0">60-110 mm</quantity>
long,
<quantity id="127F48B9C429F347290E5637CF6E35AF" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="4" pageNumber="121" unit="mm" value="18.5" valueMax="22.0" valueMin="15.0">15-22 mm</quantity>
between pinna pairs; rachis grooved, lightly puberulent, rachis glands puberulent, lightly striate, narrowly volcano-shaped, 1 or 2 located at the node of each pinna pair, sometimes scattered along the rachis between the pinna pairs; petiole grooved, lightly puberulent,
<quantity id="57B5BF16C945B0021F0EE8E8F8E8B182" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="3.0" metricValueMin="1.4" pageId="4" pageNumber="121" unit="mm" value="22.0" valueMax="30.0" valueMin="14.0">14-30 mm</quantity>
long. Petiolar glands narrowly volcano-shaped, usually 4-10, scattered along the petiole, puberulent, lightly striate, apex
<quantity id="A82B2742A64C5DC884B200DA7ABB9690" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="4" pageNumber="121" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4-0.7 mm</quantity>
across, base
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across. Leaflets 15-25 pairs per pinna, glabrous, linear,
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long,
<quantity id="40604967F6B1BA91670885E33CC41E9F" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.9" metricValueMin="1.8" pageId="4" pageNumber="121" unit="mm" value="2.85" valueMax="3.9" valueMin="1.8">1.8-3.9 mm</quantity>
wide, lateral veins obvious, 2-3 veins from the base, apex obtuse to rarely acute. Inflorescence a densely flowered globose head,
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across, in axillary clusters of
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the axil of reduced leaves, commonly on short axillary branches
<quantity id="876AC2A38C0899893CFA8EDDC864E004" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="4" pageNumber="121" unit="mm" value="95.0" valueMax="150.0" valueMin="40.0">40-150 mm</quantity>
long; peduncles lightly puberulent,
<quantity id="4D095DC6D7CFF5B8439F80286B94D99C" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="121" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10-15 mm</quantity>
long,
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thick, nearly the same thickness throughout; involucre located on the lower Vz of the peduncle, puberulent, 5-lobed. Floral bracts peltate, apex circular, ciliate, stalk 0.8-1.0 mm long. Flowers sessile; calyx 5-lobed, puberulent on the lobes,
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long; corolla 5-lobed, glabrous, pale yellow,
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long.. Legumes (only one specimen, Janzen 11051, with fruits seen) slightly curved, flattened,
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long,
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wide, glabrous, longitudinally striate, dark brown to black, dehiscent along two sutures, base not stipitate, the apex acute. Flowering April-July.
</paragraph>
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Distribution. Heavily shaded understory at lower elevations (to
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) in wet forests, in moist disturbed stream banks, at forest edge, and on roadsides, in Puntarenas Province, Costa Rica.
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specimens.
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.
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:
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,
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(
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<subSubSection id="5F9BDF6E2B78B2265CC683D97F9B8BD7" pageId="5" pageNumber="122" type="discussion">
<paragraph id="64C156CC4BF737060B860612BA973E0E" pageId="5" pageNumber="122" type="mainText">
The narrow geographic range of
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is typical of many of the ant-acacia species of Central America. Its normal habitat appears to be at lower elevations in heavily shaded understory of wet tropical forests. In general, it reacts fairly well to disturbances, and most present-day collections are from roadsides, recent landslides, disturbed stream banks, and secondary successional areas after logging. It is rarely found in open pastures, and according to
<bibRefCitation id="0B64B63A19138EB08BEAA317C4400DE1" author="Janzen, D. H." journalOrPublisher="Smithsonian Contr. Bot" pageId="5" pageNumber="122" pagination="1 - 131" part="13" refId="ref22088" refString="-. 1974. Swollen-thorn acacias of Central America. Smithsonian Contr. Bot. 13: 1 - 131." title="Swollen-thorn acacias of Central America" type="journal article" year="1974">Janzen (1974)</bibRefCitation>
will not survive fire.
</paragraph>
<paragraph id="BF2AF130666B133128C45E78FB60F532" pageId="5" pageNumber="122" type="mainText">
As is typical of many wet forest ant-acacias, Beltian body production in
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is not particularly extensive. However, the Beltian bodies are much larger (to
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long) than those found in dry area ant-acacias. In
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, these relatively large Beltian bodies are formed only on the lower 3-6 leaflets of each pinna, and occasionally the lower leaflet of a pinna has been replaced by a large Beltian body. None of the individuals tested positive for cyanide production.
</paragraph>
<paragraph id="F49889A513FC73B7FD13373025511D1B" pageId="5" pageNumber="122" type="mainText">
<taxonomicName id="F5FF62455691C1F7BA807F46D5095EB7" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="122" phylum="Tracheophyta" rank="species" species="allenii">Acacia allenii</taxonomicName>
is most closely related to
<taxonomicName id="B2B93C4EBEA4F4EF70C331A7454601D5" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="122" phylum="Tracheophyta" rank="species" species="melanoceras">A. melanoceras</taxonomicName>
and possibly
<taxonomicName id="3DB2DEE001DA6A49D38710381A0CCC35" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="122" phylum="Tracheophyta" rank="species" species="collinsii">A. collinsii</taxonomicName>
. The globose inflorescences and the presence of rachis glands easily separate this species from
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, which has elongated inflorescences and usually lacks rachis glands. It is easily separated from
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, since its leaflets are nearly twice as large and have obvious secondary veins. Also, well-developed leaves of
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have only 4-10 widely scattered petiolar glands, while
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commonly has 6-30 closely spaced petiolar glands on the flattened adaxial surface of the petiole. Furthermore, the fertile branches of
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are commonly less than
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long, are scattered along major branches throughout the crown, and have 5-15 inflorescences in the axis of reduced leaves. In
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, the fertile branches are to
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long, tend to be bunched near the ends of the sterile branches, and have 2-6 inflorescences at each leafless node.
</paragraph>
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</treatment>
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