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<document id="9FEBDA001F260791A07C9EA9192622F3" ID-DOI="10.5281/zenodo.193523" ID-GBIF-Dataset="d1368e6e-3dea-4d66-bacd-94c02d3805fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="193523" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460152417717" checkinUser="plazi" docAuthor="Köhler, Gunther" docDate="2010" docId="602D8784834FEC69FF41FA3AD964F91E" docLanguage="en" docName="zt02354p018.pdf" docOrigin="Zootaxa 2354" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anolis charlesmyersi Köhler, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="9C14FFFC834AEC61FFD6FF8FDD27FFA0" masterDocTitle="A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="6" updateTime="1698235538294" updateUser="plazi">
<mods:mods id="68531668B4E2EA95B4443DF6FEE5415D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E528844C68119F2B1FF735B45848A993">A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae)</mods:title>
</mods:titleInfo>
<mods:name id="4211470BEF8C366BC0910796D7F65EED" type="personal">
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<mods:roleTerm id="A38CF14D66B0909A2624E73F975CE4F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9954CAA219A1DE39E8B6BE382660FCB">Köhler, Gunther</mods:namePart>
</mods:name>
<mods:typeOfResource id="F5520FBB5B8ADFE4B903C42602434197">text</mods:typeOfResource>
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<mods:title id="5E16EA5D9FB78F2CA16998669BD290BE">Zootaxa</mods:title>
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<mods:part id="AA8FDD77D07EF3993459360D0E4E1B18">
<mods:date id="D81B686C2EAF29FF7F7F0631599A0879">2010</mods:date>
<mods:detail id="CDF685BD3BC619C48E2180EBCD06CB05" type="volume">
<mods:number id="BE4540B21320F17F2641E9BA7907619D">2354</mods:number>
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<mods:classification id="0809DDB346EC035DAEA43D1D439A9B29">journal article</mods:classification>
<mods:identifier id="81C93A00B81990C62581F7373B5D299B" type="DOI">10.5281/zenodo.193523</mods:identifier>
<mods:identifier id="A85E4ABB6E66078B972983B26961689A" type="GBIF-Dataset">d1368e6e-3dea-4d66-bacd-94c02d3805fa</mods:identifier>
<mods:identifier id="BCE95E57313107005534E1A46D92B20C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A86D7EF2D9FBF175F07ADE87A03D67F8" type="Zenodo-Dep">193523</mods:identifier>
</mods:mods>
<treatment id="602D8784834FEC69FF41FA3AD964F91E" ID-DOI="http://doi.org/10.5281/zenodo.6201487" ID-GBIF-Taxon="119383282" ID-Zenodo-Dep="6201487" LSID="urn:lsid:plazi:treatment:602D8784834FEC69FF41FA3AD964F91E" httpUri="http://treatment.plazi.org/id/602D8784834FEC69FF41FA3AD964F91E" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
<subSubSection id="A09E6519834FEC64FF41FA3ADC74FA55" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="E83B3692834FEC64FF41FA3ADCDEFA6F" blockId="5.[151,505,1461,1525]" box="[151,505,1461,1487]" pageId="5" pageNumber="6">
<heading id="B37381FE834FEC64FF41FA3ADCDEFA6F" bold="true" box="[151,505,1461,1487]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="DAF0EA80834FEC64FF41FA3ADCDEFA6F" bold="true" box="[151,505,1461,1487]" pageId="5" pageNumber="6">
<taxonomicName id="2F844D11834FEC64FF41FA3ADCB3FA6F" box="[151,404,1461,1487]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="charlesmyersi" status="sp. nov.">
<emphasis id="DAF0EA80834FEC64FF41FA3ADCB3FA6F" bold="true" box="[151,404,1461,1487]" italics="true" pageId="5" pageNumber="6">Anolis charlesmyersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1C357FB834FEC64FE4DFA3ADCDEFA6F" box="[411,505,1461,1487]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E83B3692834FEC64FF41FA54DC74FA55" blockId="5.[151,505,1461,1525]" box="[151,339,1499,1525]" pageId="5" pageNumber="6">
<figureCitation id="70BF2A17834FEC64FF41FA54DDCEFA55" box="[151,233,1499,1525]" captionStart="FIGURE 4" captionStartId="8.[151,254,1033,1057]" captionTargetBox="[264,1323,306,1011]" captionTargetId="figure@8.[264,1323,306,1011]" captionTargetPageId="8" captionText="FIGURE 4. Holotype of Anolis charlesmyersi (SMF 89688) in life. SVL = 65.0 mm. Photo by S. Lotzkat." httpUri="https://zenodo.org/record/193528/files/figure.png" pageId="5" pageNumber="6">Figs. 4</figureCitation>
,
<figureCitation id="70BF2A17834FEC64FF23FA54DC22FA55" box="[245,261,1499,1525]" captionStart="FIGURE 5" captionStartId="9.[151,254,2005,2029]" captionTargetBox="[346,1244,209,1972]" captionTargetId="figure@9.[322,1278,183,1989]" captionTargetPageId="9" captionText="FIGURE 5. Head of holotype of Anolis charlesmyersi (SMF 89688). Scale bars equal 1.0 mm." httpUri="https://zenodo.org/record/193529/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
, and 6
</paragraph>
</subSubSection>
<subSubSection id="A09E6519834FEC64FF41F9A9DFD1F9FC" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="E83B3692834FEC64FF41F9A9DFD1F9FC" blockId="5.[151,1413,1573,1629]" pageId="5" pageNumber="6">
<taxonomicName id="2F844D11834FEC64FF41F9A9DF3EF99C" authority="Wettstein (1934)" authorityName="Wettstein" authorityYear="1934" box="[151,537,1573,1597]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA80834FEC64FF41F9A9DC70F99D" box="[151,343,1574,1597]" italics="true" pageId="5" pageNumber="6">Anolis pentaprion</emphasis>
:
<bibRefCitation id="8C154B63834FEC64FEB2F9AADF3EF99C" author="Wettstein" box="[356,537,1573,1596]" pageId="5" pageNumber="17" refString="Wettstein, O. (1934) Ergebnisse der osterreichischen biologischen Costa Rica-Expedition 1930. Die Amphibien und Reptilien. Aus den Sitzungsberichten der Akademie der Wissenschaften in Wien Mathematischnaturwissenschaftliche Klasse, Abteilung I, 143, 1 - 39." type="journal article" year="1934">Wettstein (1934)</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="8C154B63834FEC64FDF2F9AADE2BF99C" author="Taylor" box="[548,780,1573,1596]" pageId="5" pageNumber="17" refString="Taylor, E. H. (1956) A review of the lizards of Costa Rica. - University of Kansas Science Bulletin, 38, 1 - 322." type="journal article" year="1956">Taylor (1956; in part)</bibRefCitation>
,
<bibRefCitation id="8C154B63834FEC64FCC1F9AAD916F99C" author="Williams" box="[791,1073,1573,1596]" pageId="5" pageNumber="17" refString="Williams, K. L. &amp; Smith, H. M. (1968) Noteworthy lizards of the genus Anolis from Costa Rica. Caribbean Journal of Science, 6, 163 - 166. (&quot; 1966 &quot;)." type="journal article" year="1968">Williams and Smith (1968</bibRefCitation>
“1966”),
<bibRefCitation id="8C154B63834FEC64FB4CF9AAD8A6F99C" author="Myers" box="[1178,1409,1573,1596]" lastPageId="5" lastPageNumber="17" pageId="5" pageNumber="16" refString="Myers, C. W. (1971) Central American lizards related to Anolis pentaprion: Two new species from the Cordillera de Talamanca. American Museum Novitates, 2471, 1 - 40." type="journal article" year="1971">Myers (1971; in part)</bibRefCitation>
,
<taxonomicName id="2F844D11834FEC64FF41F9C9DF6FF9FC" authority="Etheridge" authorityName="Etheridge" box="[151,584,1605,1629]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="subSpecies" species="pentaprion" subSpecies="pentaprion">
<emphasis id="DAF0EA80834FEC64FF41F9C9DCF5F9FD" box="[151,466,1606,1629]" italics="true" pageId="5" pageNumber="6">Anolis pentaprion pentaprion</emphasis>
:
<bibRefCitation id="8C154B63834FEC64FE36F9CADF6FF9FC" author="Etheridge" box="[480,584,1605,1628]" pageId="5" pageNumber="16" refString="Etheridge, R. (1959) The Relationships of the Anoles (Reptilia: Sauria: Iguanidae). An Interpretation Based on Skeletal Morphology. Ph. D. Dissertation, University of Michigan, Ann Arbor, Michigan, U. S. A." type="book" year="1959">Etheridge</bibRefCitation>
</taxonomicName>
(1959; in part.),
</paragraph>
</subSubSection>
<subSubSection id="A09E6519834FEC64FF41F905DC76F89A" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="E83B3692834FEC64FF41F905D8BCF96A" blockId="5.[151,1437,1674,2037]" pageId="5" pageNumber="6">
<emphasis id="DAF0EA80834FEC64FF41F905DC37F904" bold="true" box="[151,272,1674,1700]" pageId="5" pageNumber="6">
<typeStatus id="373F8830834FEC64FF41F905DC2BF904" box="[151,268,1674,1700]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="8E95AE57834FEC64FECFF904DC73F905" LSID="urn:lsid:biocol.org:col:34838" box="[281,340,1675,1701]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
89688, adult male, from trail to Río Majagua, Los Algarrobos,
<geoCoordinate id="8DB05055834FEC64FB9EF904D9ECF905" box="[1096,1227,1675,1701]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="8.48927">8.48927°N</geoCoordinate>
,
<geoCoordinate id="8DB05055834FEC64FB0EF904D849F905" box="[1240,1390,1675,1701]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="-82.43333">82.43333°W</geoCoordinate>
, ca.
<quantity id="2F7C9B77834FEC64FF41F93FDDC6F96A" box="[151,225,1712,1738]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="6" unit="m" value="130.0">130 m</quantity>
a.s.l., Province Chiriquí,
<collectingCountry id="90937602834FEC64FDDBF93FDF49F96A" box="[525,622,1712,1738]" name="Panama" pageId="5" pageNumber="6">Panama</collectingCountry>
; collected by Sebastian Lotzkat and Andreas Hertz on
<date id="9C3A1052834FEC64FB39F93FD8B0F96A" box="[1263,1431,1712,1738]" pageId="5" pageNumber="6" value="2009-04-23">23 April 2009</date>
.
</paragraph>
<paragraph id="E83B3692834FEC64FF12F95ADC76F89A" blockId="5.[151,1437,1674,2037]" pageId="5" pageNumber="6">
<emphasis id="DAF0EA80834FEC64FF12F95ADC69F94F" bold="true" box="[196,334,1749,1775]" pageId="5" pageNumber="6">
<typeStatus id="373F8830834FEC64FF12F95ADC6EF94F" box="[196,329,1749,1775]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="8E95AE57834FEC64FE80F959DCB5F950" LSID="urn:lsid:biocol.org:col:34838" box="[342,402,1750,1776]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="6">SMF</collectionCode>
89508, 90053, adult males, from trail to Río Majagua, Los Algarrobos,
<geoCoordinate id="8DB05055834FEC64FAC6F959D8BFF950" box="[1296,1432,1750,1776]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="1" value="8.48456">8.48456°N</geoCoordinate>
,
<geoCoordinate id="8DB05055834FEC64FF41F974DC0AF8B5" box="[151,301,1787,1813]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="1" value="-82.43417">82.43417°W</geoCoordinate>
, ca.
<quantity id="2F7C9B77834FEC64FEB5F974DC8AF8B5" box="[355,429,1787,1813]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="6" unit="m" value="110.0">110 m</quantity>
a.s.l., Province Chiriquí,
<collectingCountry id="90937602834FEC64FD08F974DE18F8B5" box="[734,831,1787,1813]" name="Panama" pageId="5" pageNumber="6">Panama</collectingCountry>
; collected by Sebastian Lotzkat and Andreas Hertz on
<date id="9C3A1052834FEC64FF6AF8AFDC6BF89A" box="[188,332,1824,1850]" pageId="5" pageNumber="6" value="2008-06-07">7 June 2008</date>
.
</paragraph>
</subSubSection>
<subSubSection id="A09E6519834FEC67FF12F8CADECAFD37" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="E83B3692834FEC64FF12F8CAD8BBF855" blockId="5.[151,1437,1674,2037]" pageId="5" pageNumber="6">
<emphasis id="DAF0EA80834FEC64FF12F8CADC62F8FF" bold="true" box="[196,325,1861,1887]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
A medium-sized species (SVL in largest specimen examined 78.0 mm in males,
<quantity id="2F7C9B77834FEC64FAC5F8C9D85AF8C0" box="[1299,1405,1862,1888]" metricMagnitude="-2" metricUnit="m" metricValue="7.67" pageId="5" pageNumber="6" unit="mm" value="76.7">76.7 mm</quantity>
in females) of the genus
<taxonomicName id="2F844D11834FEC64FE7AF8E4DCDCF824" box="[428,507,1899,1924]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="DAF0EA80834FEC64FE7AF8E4DCDCF824" box="[428,507,1899,1924]" italics="true" pageId="5" pageNumber="6">Anolis</emphasis>
</taxonomicName>
(sensu
<bibRefCitation id="8C154B63834FEC64FD8AF8E4DFFCF825" author="Poe" box="[604,731,1899,1925]" pageId="5" pageNumber="17" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe, 2004</bibRefCitation>
) that differs from all Central American anoles except
<emphasis id="DAF0EA80834FEC64FA54F8E4DEF6F80A" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="2F844D11834FEC64FA54F8E4DDD7F80A" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="beckeri">A. beckeri</taxonomicName>
,
<taxonomicName id="2F844D11834FEC64FF2AF81EDC5DF80A" box="[252,378,1937,1962]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="cristifer">A. cristifer</taxonomicName>
,
<taxonomicName id="2F844D11834FEC64FE50F81EDF35F80A" box="[390,530,1937,1962]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fungosus">A. fungosus</taxonomicName>
,
<taxonomicName id="2F844D11834FEC64FDC8F81EDFE4F80A" box="[542,707,1937,1962]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="pentaprion">A. pentaprion</taxonomicName>
,
<taxonomicName id="2F844D11834FEC64FD19F81EDE66F80A" box="[719,833,1937,1962]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="salvini">A. salvini</taxonomicName>
,
<taxonomicName id="2F844D11834FEC64FC9BF81EDEF6F80A" box="[845,977,1937,1962]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="utilensis">A. utilensis</taxonomicName>
</emphasis>
by (1) having (1) extremely short hind legs (fourth toe of adpressed hindlimb usually reaching to level of tympanum; ratio shank length / head length 0.49–0.84); (2) smooth, juxtaposed ventral scales; (3) a relatively short tail (ratio tail length / SVL 1.0–1.5).
</paragraph>
<paragraph id="E83B3692834CEC67FF41FF17DECAFD37" blockId="6.[151,1437,152,663]" pageId="6" pageNumber="7">
<taxonomicName id="2F844D11834CEC67FF41FF17DCB6FF11" box="[151,401,152,177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA80834CEC67FF41FF17DCB6FF11" box="[151,401,152,177]" italics="true" pageId="6" pageNumber="7">Anolis charlesmyersi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D11834CEC67FDF9FF17DF8FFF11" box="[559,680,152,177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA80834CEC67FDF9FF17DF8FFF11" box="[559,680,152,177]" italics="true" pageId="6" pageNumber="7">A. beckeri</emphasis>
</taxonomicName>
by having a pronounced serrated caudal crest (versus lacking a pronounced serrated caudal crest).
<taxonomicName id="2F844D11834CEC67FDECFF32DE11FF76" box="[570,822,189,214]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA80834CEC67FDECFF32DE11FF76" box="[570,822,189,214]" italics="true" pageId="6" pageNumber="7">Anolis charlesmyersi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D11834CEC67FC00FF32D970FF76" box="[982,1111,189,214]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cristifer">
<emphasis id="DAF0EA80834CEC67FC00FF32D970FF76" box="[982,1111,189,214]" italics="true" pageId="6" pageNumber="7">A. cristifer</emphasis>
</taxonomicName>
by having a double row of slightly enlarged middorsal scales, not forming a serrated crest (versus a serrated middorsal crest, at least on posterior portion of dorsum, present in
<taxonomicName id="2F844D11834CEC67FDB3FE87DFC3FE81" box="[613,740,264,289]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cristifer">
<emphasis id="DAF0EA80834CEC67FDB3FE87DFC3FE81" box="[613,740,264,289]" italics="true" pageId="6" pageNumber="7">A. cristifer</emphasis>
</taxonomicName>
).
<taxonomicName id="2F844D11834CEC67FD2AFE87DED3FE81" box="[764,1012,264,289]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA80834CEC67FD2AFE87DED3FE81" box="[764,1012,264,289]" italics="true" pageId="6" pageNumber="7">Anolis charlesmyersi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D11834CEC67FB59FE87D83EFE81" box="[1167,1305,264,289]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="fungosus">
<emphasis id="DAF0EA80834CEC67FB59FE87D83EFE81" box="[1167,1305,264,289]" italics="true" pageId="6" pageNumber="7">A. fungosus</emphasis>
</taxonomicName>
by lacking bony parietal protuberances (versus having a pair of small bony parietal protuberances posterior and lateral to interparietal plate).
<taxonomicName id="2F844D11834CEC67FEAAFEDDDF56FECB" box="[380,625,338,363]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA80834CEC67FEAAFEDDDF56FECB" box="[380,625,338,363]" italics="true" pageId="6" pageNumber="7">Anolis charlesmyersi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D11834CEC67FCDCFEDDDE8CFECB" box="[778,939,338,363]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA80834CEC67FCDCFEDDDE8CFECB" box="[778,939,338,363]" italics="true" pageId="6" pageNumber="7">A. pentaprion</emphasis>
</taxonomicName>
by having a brick red (in life) dewlap with few large widely spaced gorgetal scales scattered across the dewlap or in loosely arranged rows with 4–9 widely spaced scales (versus a pink dewlap with purple pigment between rows of gorgetal scales and with regular rows of 17–25 small narrowly spaced gorgetal scales).
<taxonomicName id="2F844D11834CEC67FCA0FE4DD94AFE7B" box="[886,1133,450,475]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA80834CEC67FCA0FE4DD94AFE7B" box="[886,1133,450,475]" italics="true" pageId="6" pageNumber="7">Anolis charlesmyersi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D11834CEC67FADEFE4DD850FE7B" box="[1288,1399,450,475]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="salvini">
<emphasis id="DAF0EA80834CEC67FADEFE4DD850FE7B" box="[1288,1399,450,475]" italics="true" pageId="6" pageNumber="7">A. salvini</emphasis>
</taxonomicName>
by having smooth, obliquely conical ventral scales (versus keeled), lacking a dark interorbital bar (versus present), and lacking enlarged postcloacal scales (versus males with a pair of greatly enlarged postcloacal scales).
<taxonomicName id="2F844D11834CEC67FF2BFDBDDF23FDEB" box="[253,516,562,587]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA80834CEC67FF2BFDBDDF23FDEB" box="[253,516,562,587]" italics="true" pageId="6" pageNumber="7">Anolis charlesmyersi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="2F844D11834CEC67FD7AFDBDDE1EFDEB" box="[684,825,562,587]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="utilensis">
<emphasis id="DAF0EA80834CEC67FD7AFDBDDE1EFDEB" box="[684,825,562,587]" italics="true" pageId="6" pageNumber="7">A. utilensis</emphasis>
</taxonomicName>
by having the proximal subdigital scales of toes differentiated as slightly broadened lamellae (versus those scales granular) and by having a pronounced serrated caudal crest (versus lacking a pronounced serrated caudal crest).
</paragraph>
</subSubSection>
<caption id="BCFB661A834CEC67FF41F9C6DE15F960" httpUri="https://zenodo.org/record/193527/files/figure.png" pageId="6" pageNumber="7" targetBox="[161,1430,732,1582]" targetPageId="6">
<paragraph id="E83B3692834CEC67FF41F9C6DE15F960" blockId="6.[151,1437,1609,1729]" pageId="6" pageNumber="7">
<emphasis id="DAF0EA80834CEC67FF41F9C6DC3DF9C1" bold="true" box="[151,282,1609,1633]" pageId="6" pageNumber="7">FIGURE 3.</emphasis>
Map indicating collecting localities of
<taxonomicName id="2F844D11834CEC67FD12F9C5DEA2F9C1" box="[708,901,1610,1633]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA80834CEC67FD12F9C5DEA2F9C1" box="[708,901,1610,1633]" italics="true" pageId="6" pageNumber="7">Anolis pentaprion</emphasis>
</taxonomicName>
and related taxa in Mexico and Central America. Each symbol can represent one or more adjacent localities. Areas above sea level, 500 and 1000 m are shaded pale, medium dark, and dark gray, respectively. Red triangles:
<taxonomicName id="2F844D11834CEC67FD2DF905DEFCF901" box="[763,987,1674,1697]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA80834CEC67FD2DF905DEFCF901" box="[763,987,1674,1697]" italics="true" pageId="6" pageNumber="7">Anolis charlesmyersi</emphasis>
</taxonomicName>
; blue circles:
<taxonomicName id="2F844D11834CEC67FBA2F905D9C7F901" box="[1140,1248,1674,1697]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="beckeri">
<emphasis id="DAF0EA80834CEC67FBA2F905D9C7F901" box="[1140,1248,1674,1697]" italics="true" pageId="6" pageNumber="7">A. beckeri</emphasis>
</taxonomicName>
; green circles:
<taxonomicName id="2F844D11834CEC67FA53F905DDCCF961" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="cristifer">
<emphasis id="DAF0EA80834CEC67FA53F905DDCCF961" italics="true" pageId="6" pageNumber="7">A. cristifer</emphasis>
</taxonomicName>
; black squares:
<taxonomicName id="2F844D11834CEC67FE4EF925DF0CF961" box="[408,555,1706,1729]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA80834CEC67FE4EF925DF0CF961" box="[408,555,1706,1729]" italics="true" pageId="6" pageNumber="7">A. pentaprion</emphasis>
</taxonomicName>
; red square:
<taxonomicName id="2F844D11834CEC67FD62F925DE0CF961" box="[692,811,1706,1729]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="utilensis">
<emphasis id="DAF0EA80834CEC67FD62F925DE0CF961" box="[692,811,1706,1729]" italics="true" pageId="6" pageNumber="7">A. utilensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="A09E6519834CEC69FF12F961D923F9EE" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="E83B3692834CEC66FF12F961D903FBF7" blockId="6.[151,1436,1774,2025]" lastBlockId="7.[151,1437,152,2007]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="DAF0EA80834CEC67FF12F961DCD5F8A8" bold="true" box="[196,498,1774,1800]" pageId="6" pageNumber="7">
Description of
<typeStatus id="373F8830834CEC67FEA8F961DCCBF8A8" box="[382,492,1774,1800]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male as indicated by everted hemipenes; SVL 65.0 mm; tail length 93.0 mm, tail complete; tail length/SVL ratio 1.43; tail compressed in cross section, tail height
<quantity id="2F7C9B77834CEC67FB14F89BD838F88E" box="[1218,1311,1812,1838]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="7" unit="mm" value="4.4">4.4 mm</quantity>
, tail width
<quantity id="2F7C9B77834CEC67FF41F8B5DDD3F8F4" box="[151,244,1850,1876]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="7" unit="mm" value="3.1">3.1 mm</quantity>
; axilla to groin distance
<quantity id="2F7C9B77834CEC67FDC5F8B5DF58F8F4" box="[531,639,1850,1876]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="6" pageNumber="7" unit="mm" value="24.9">24.9 mm</quantity>
; head length
<quantity id="2F7C9B77834CEC67FCCCF8B5DEA2F8F4" box="[794,901,1850,1876]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="6" pageNumber="7" unit="mm" value="17.5">17.5 mm</quantity>
, head length/SVL ratio 0.27; snout length
<quantity id="2F7C9B77834CEC67FAA1F8B5DDEDF8D9" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="7" unit="mm" value="8.5">8.5 mm</quantity>
; head width
<quantity id="2F7C9B77834CEC67FEBEF8D0DCFDF8D9" box="[360,474,1887,1913]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="6" pageNumber="7" unit="mm" value="10.4">10.4 mm</quantity>
; longest toe of adpressed hind limb reaching to a point between shoulder and tympanum; shank length
<quantity id="2F7C9B77834CEC67FE13F80BDF14F83E" box="[453,563,1924,1950]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="6" pageNumber="7" unit="mm" value="13.6">13.6 mm</quantity>
, shank length/head length ratio 0.78; longest finger of extended forelimb reaching to a point between eye and nostril; longest finger of adpressed forelimb reaches to a point
<quantity id="2F7C9B77834CEC67FA80F825D8BBF864" box="[1366,1436,1962,1988]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="mm" value="2.0">2 mm</quantity>
anterior to insertion of hind limbs. Scales on snout tuberculate to weakly keeled; 10 postrostrals; 7 scales between nasals; scales in distinct prefrontal depression slightly wrinkled to tuberculate; supraorbital semicircles well developed, 2 (right)–3 (left) scales broadly in contact; supraorbital disc composed of 11–14 distinctly enlarged smooth scales; three short superciliaries, posterior one shortest; about 4 rows of small keeled scales extending between enlarged supraorbitals and superciliaries; interparietal scale well developed, 3.1 x
<quantity id="2F7C9B77834DEC66FF37FEA2DC65FEE7" box="[225,322,301,327]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="8" unit="mm" value="1.3">1.3 mm</quantity>
(length x width), surrounded by scales of moderate size; interparietal and supraorbital semicircles in broad contact; canthal ridge distinct, composed of three large and four small anterior canthal scales; 6 scales present between second canthals; 8 scales present between posterior canthals; 19 (right)–20 (left) loreal scales in a maximum of 3 (right)–4 (left) horizontal rows, with the scales of lower rows mostly smooth, and those of upper rows mostly weakly keeled; subocular scales smooth to wrinkled, five subocular scales broadly in contact with five supralabials; 8 supralabials to level below center of eye; ear opening 0.8 x
<quantity id="2F7C9B77834DEC66FF41FD82DDD0FD87" box="[151,247,525,551]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="mm" value="0.9">0.9 mm</quantity>
(length x height); mental 2.0 x
<quantity id="2F7C9B77834DEC66FD5AFD82DFCBFD87" box="[652,748,525,551]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="mm" value="0.9">0.9 mm</quantity>
(width x length), almost completely divided medially, bordered posteriorly by 6 postmentals (outer pair largest); 8 infralabials to level below center of eye; sublabials undifferentiated; smooth granular scales present on chin and throat; dewlap extending well onto chest, from level below mideye to
<quantity id="2F7C9B77834DEC66FDE5FDF2DFA8FD37" box="[563,655,637,663]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="8" unit="mm" value="3.5">3.5 mm</quantity>
beyond level of axilla; dorsum of body with smooth conical nonimbricate scales, 2 medial rows of slightly enlarged scales, largest dorsal scales about 0.35 x
<quantity id="2F7C9B77834DEC66FB0FFD2DD866FD1C" box="[1241,1345,674,700]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="8" unit="mm" value="0.29">0.29 mm</quantity>
(length x width); about 51 medial dorsal scales in one head length; about 75 medial dorsal scales between axilla and groin; lateral scales smooth, granular and homogeneous, average size
<quantity id="2F7C9B77834DEC66FBD2FD62D955FCA7" box="[1028,1138,749,775]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="8" unit="mm" value="0.24">0.24 mm</quantity>
in diameter; ventrals at midbody smooth, bulging, nonimbricate, about 0.34 x
<quantity id="2F7C9B77834DEC66FCF3FC9DDEB7FC8C" box="[805,912,786,812]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="8" unit="mm" value="0.31">0.31 mm</quantity>
(length x width); about 52 ventral scales in one head length; about 77 ventral scales between axilla and groin; 154 scales around midbody; ventral caudal scales strongly keeled, forming longitudinal ridges; caudal middorsal scales enlarged, forming a serrated crest; lateral caudal scales mostly smooth, without whorls of enlarged scales, although an indistinct division in segments is discernible; precloacal scales pointed and granular; postcloacal scales not enlarged; no tube-like axillary pocket present; scales on dorsal surface of forelimb smooth, conical, subimbricate, about 0.25 x
<quantity id="2F7C9B77834DEC66FABFFC42DDE2FBAC" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="8" unit="mm" value="0.24">0.24 mm</quantity>
(length x width); digital pads dilated, dilated pad about three times width of non-dilated scales on distal phalanx; distal phalanx narrower than and raised from dilated pad; 33 (right)–34 (left) lamellae under phalanges II–IV of fourth toe; seven scales under distal phalanx of fourth toe.
</paragraph>
<paragraph id="E83B3692834DEC66FF12FBEDD962FB67" blockId="7.[151,1437,152,2007]" pageId="7" pageNumber="8">The completely everted left hemipenis is a large bilobate organ; sulcus spermaticus bordered by well developed sulcal lips and bifurcates at base of apex, branches continue to tip of lobes; a small asulcate processus present; apex covered with fine calyces, truncus with transverse folds.</paragraph>
<paragraph id="E83B3692834DEC66FF12FB5DDEC3FA97" blockId="7.[151,1437,152,2007]" pageId="7" pageNumber="8">The extended dewlap has four horizontal gorgetal-sternal rows with 8–10 large scales per row; about 75 scales in marginal series, modal number of anterior marginal pairs 4; about 35 apicogorgetals; gorgetals widely spaced, no scales between gorgetal rows; about 20 apicosternals.</paragraph>
<paragraph id="E83B3692834DEC66FF12FACDD94BF957" blockId="7.[151,1437,152,2007]" pageId="7" pageNumber="8">
In life, dorsal and lateral surfaces Dark Drab (119B), with diffuse spots and blotches of Vandyke Brown (221) and circular dirty white blotches with a suggestion of Pearl Gray (81); a dirty white middorsal stripe with a suggestion of Pearl Gray (81) interrupted by Sepia (119) blotches; a diffuse dirty white lateral stripe with a suggestion of Pearl Gray (81) extending from shoulder to groin; tail Beige (219D), except for its base, with broad Brussels Brown (121B) transverse bands and fine Sepia (119) mottling; a diffuse Vandyke Brown (221) pre- and postorbital stripe extending about
<quantity id="2F7C9B77834DEC66FD2FFA72DE6AF9B7" box="[761,845,1533,1559]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="cm" value="0.5">0.5 cm</quantity>
each anteriorly and posteriorly of orbit; ventral surfaces of body, limbs, and base of tail dirty white with a suggestion of Pearl Gray (81), with reticulate Dark Drab (119B) mottling; chin and gular region Flaxflower Blue (170C) with a suggestion of Light Sky Blue (168D), with reticulate Hair Brown (119A) mottling; iris Cinnamon (123A); throat Blackish Neutral Gray (82); tongue Cream Color (54); dewlap Poppy Red (108A) with circular Crimson (108) blotches between gorgetal rows; dewlap scales dirty white with a suggestion of Lavender Blue (170D). Color in preservative (70% ethanol) similar to color in life except for chin and gular region is dirty white.
</paragraph>
<paragraph id="E83B3692834DEC69FF12F88DDE9AFF5C" blockId="7.[151,1437,152,2007]" lastBlockId="8.[151,1437,152,252]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="DAF0EA80834DEC66FF12F88DDC61F8BC" bold="true" box="[196,326,1794,1820]" pageId="7" pageNumber="8">Variation.</emphasis>
The
<typeStatus id="373F8830834DEC66FE5DF88DDF24F8BC" box="[395,515,1794,1820]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
agree well with the
<typeStatus id="373F8830834DEC66FCD1F88DDE53F8BC" box="[775,884,1794,1820]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
in general appearance, morphometrics, and scalation (see Table 1). The coloration in life of SMF 89508 was recorded as follows: Dorsal and lateral surfaces of body, limbs and anterior half of tail Smoke Gray (45) suffused with Cinnamon-Drab (219C), sparse Sepia (219) mottling and scattered flecks of dirty white with a suggestion of Pearl Gray (81); posterior half of tail Drab-Gray (119D) with broad Raw Sienna (136) transverse bands; ventral surfaces of body, limbs, and anterior half of tail dirty white with a suggestion of Opaline Green (162D); dorsal and lateral surfaces of head Drab-Gray (119D) with Sepia (219) mottling on supraorbital semicircles, snout, and parietal region; ventral surface of head Light Sky Blue (168D) with Hair Brown (119A) spots on anterior half; dewlap Spectrum Red (11) with Carmine (8) blotches and dirty white scales.
</paragraph>
<caption id="BCFB661A8342EC69FF41FB86D821FB80" box="[151,1286,1033,1057]" httpUri="https://zenodo.org/record/193528/files/figure.png" pageId="8" pageNumber="9" targetBox="[264,1323,306,1011]" targetPageId="8">
<paragraph id="E83B36928342EC69FF41FB86D821FB80" blockId="8.[151,1286,1033,1057]" box="[151,1286,1033,1057]" pageId="8" pageNumber="9">
<emphasis id="DAF0EA808342EC69FF41FB86DC3FFB81" bold="true" box="[151,280,1033,1057]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
Holotype of
<taxonomicName id="2F844D118342EC69FE70FB86DFA0FB80" box="[422,647,1033,1056]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA808342EC69FE70FB86DFA0FB80" box="[422,647,1033,1056]" italics="true" pageId="8" pageNumber="9">Anolis charlesmyersi</emphasis>
</taxonomicName>
(SMF 89688) in life. SVL = 65.0 mm. Photo by S. Lotzkat.
</paragraph>
</caption>
<paragraph id="E83B36928342EC69FF12FBC1DE19F9A3" blockId="8.[151,1437,1102,1726]" pageId="8" pageNumber="9">
<emphasis id="DAF0EA808342EC69FF12FBC1DCF0FBC8" bold="true" box="[196,471,1102,1128]" pageId="8" pageNumber="9">Natural history notes.</emphasis>
In the area of Hacienda Barú (near Dominical,
<collectingCountry id="909376028342EC69FBC6FBC1D9B4FBC8" box="[1040,1171,1102,1128]" name="Costa Rica" pageId="8" pageNumber="9">Costa Rica</collectingCountry>
)
<taxonomicName id="2F844D118342EC69FB75FBC0D8BBFBC8" box="[1187,1436,1103,1128]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA808342EC69FB75FBC0D8BBFBC8" box="[1187,1436,1103,1128]" italics="true" pageId="8" pageNumber="9">Anolis charlesmyersi</emphasis>
</taxonomicName>
appears to be fairly common, especially along the beach and flat areas where there are many
<taxonomicName id="2F844D118342EC69FAC1FBFBDDD1FB12" class="Magnoliopsida" family="Combretaceae" genus="Terminalia" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="catappa">
<emphasis id="DAF0EA808342EC69FAC1FBFBDDD1FB12" italics="true" pageId="8" pageNumber="9">Terminalia catappa</emphasis>
</taxonomicName>
trees (Almendro de playa) (R. Mason pers. communication
<date id="9C3A10528342EC69FC69FB16D9AFFB13" box="[959,1160,1177,1203]" pageId="8" pageNumber="9" value="2009-01-23">23 January 2009</date>
). This statement is also supported by the relative large number of specimens collected from this site (see Appendix I). However, I failed to detect this species during a two days stay at Hacienda Barú during the dry season in
<date id="9C3A10528342EC69FAD5FB6BD8BEFB5E" box="[1283,1433,1252,1278]" pageId="8" pageNumber="9" value="2009-03">March 2009</date>
. Whether my lack of spotting this species indicates some kind of seasonality remains to be investigated.
<bibRefCitation id="8C154B638342EC69FF41FAA1DCA0FAE8" author="Barbour" box="[151,391,1326,1352]" pageId="8" pageNumber="16" refString="Barbour, T. (1934) The anoles. II. The mainland species from Mexico southward. Bulletin of the Museum of Comparative Zoology, Harvard University, 77, 121 - 155." type="journal article" year="1934">Barbour (1934:145)</bibRefCitation>
stated that
<taxonomicName id="2F844D118342EC69FDC2FAA0DFC9FAE8" box="[532,750,1327,1352]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA808342EC69FDC2FAA0DFC9FAE8" box="[532,750,1327,1352]" italics="true" pageId="8" pageNumber="9">Anolis pentaprion</emphasis>
</taxonomicName>
was “not uncommon in the Canal Zone during the rainy season, excessively rare during the dry months.” At the
<typeStatus id="373F88308342EC69FCFDFADBDE79FACE" box="[811,862,1364,1390]" pageId="8" pageNumber="9">type</typeStatus>
locality near Los Algarrobos, Chiriquí,
<collectingCountry id="909376028342EC69FAE1FADBD8B0FACE" box="[1335,1431,1364,1390]" name="Panama" pageId="8" pageNumber="9">Panama</collectingCountry>
, a similar seasonality seems to occur with several observations during a single search in the rainy season and none but the
<typeStatus id="373F88308342EC69FEE1FA11DC85FA18" box="[311,418,1438,1464]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
during numerous searches at the peak of the dry season (Lotzkat pers. comm.
<date id="9C3A10528342EC69FABFFA11DDFEFA7E" pageId="8" pageNumber="9" value="2009-10">Oct. 2009</date>
). The
<typeStatus id="373F88308342EC69FEF3FA4BDCA8FA7E" box="[293,399,1476,1502]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
was collected at 22:40 hours sleeping on a vertical twig of a small tree, about
<quantity id="2F7C9B778342EC69FA80FA4BD8BBFA7E" box="[1366,1436,1476,1502]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="9" unit="m" value="2.5">2.5 m</quantity>
above the ground in relatively open secondary vegetation.
</paragraph>
<paragraph id="E83B36928342EC69FF12F981D923F9EE" blockId="8.[151,1437,1102,1726]" pageId="8" pageNumber="9">
<emphasis id="DAF0EA808342EC69FF12F981DF35F988" bold="true" box="[196,530,1550,1576]" pageId="8" pageNumber="9">Geographic distribution.</emphasis>
<collectingCountry id="909376028342EC69FDCAF981DF18F988" box="[540,575,1550,1576]" name="American Samoa" pageId="8" pageNumber="9">As</collectingCountry>
currently understood,
<taxonomicName id="2F844D118342EC69FCB7F980D949F988" box="[865,1134,1551,1576]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="charlesmyersi">
<emphasis id="DAF0EA808342EC69FCB7F980D949F988" box="[865,1134,1551,1576]" italics="true" pageId="8" pageNumber="9">Anolis charlesmyersi</emphasis>
</taxonomicName>
is distributed along the lowlands of the Pacific versant in
<collectingCountry id="909376028342EC69FDF3F9BBDF83F9EE" box="[549,676,1588,1614]" name="Costa Rica" pageId="8" pageNumber="9">Costa Rica</collectingCountry>
and western
<collectingCountry id="909376028342EC69FC96F9BBDEBAF9EE" box="[832,925,1588,1614]" name="Panama" pageId="8" pageNumber="9">Panama</collectingCountry>
(
<figureCitation id="70BF2A178342EC69FC7BF9BBDED2F9EE" box="[941,1013,1588,1614]" captionStart="FIGURE 3" captionStartId="6.[151,255,1609,1633]" captionTargetBox="[161,1430,732,1582]" captionTargetId="figure@6.[151,1436,717,1588]" captionTargetPageId="6" captionText="FIGURE 3. Map indicating collecting localities of Anolis pentaprion and related taxa in Mexico and Central America. Each symbol can represent one or more adjacent localities. Areas above sea level, 500 and 1000 m are shaded pale, medium dark, and dark gray, respectively. Red triangles: Anolis charlesmyersi; blue circles: A. beckeri; green circles: A. cristifer; black squares: A. pentaprion; red square: A. utilensis." httpUri="https://zenodo.org/record/193527/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A09E65198342EC69FF12F9D7D964F91E" pageId="8" pageNumber="9" type="etymology">
<paragraph id="E83B36928342EC69FF12F9D7D964F91E" blockId="8.[151,1437,1102,1726]" pageId="8" pageNumber="9">
<emphasis id="DAF0EA808342EC69FF12F9D7DC73F9D2" bold="true" box="[196,340,1624,1650]" pageId="8" pageNumber="9">Etymology.</emphasis>
The specific name is a patronym for my colleague Charles W. Myers in recognition of his important contributions to the herpetology of Lower Central
<collectingCountry id="909376028342EC69FCBDF9F1DEF7F938" box="[875,976,1662,1688]" name="United States of America" pageId="8" pageNumber="9">America</collectingCountry>
in general and to the understanding of the taxonomy and sytematics of the anoles of the
<taxonomicName id="2F844D118342EC69FD0DF92BDE7BF91D" box="[731,860,1700,1725]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="pentaprion">
<emphasis id="DAF0EA808342EC69FD0DF92BDE7BF91D" box="[731,860,1700,1725]" italics="true" pageId="8" pageNumber="9">pentaprion</emphasis>
</taxonomicName>
group in particular.
</paragraph>
</subSubSection>
</treatment>
</document>
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