276 lines
35 KiB
XML
276 lines
35 KiB
XML
<document id="D87FEE2B3319495E8B902A700992549C" ID-DOI="10.5281/zenodo.193523" ID-GBIF-Dataset="d1368e6e-3dea-4d66-bacd-94c02d3805fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="193523" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460152417717" checkinUser="plazi" docAuthor="Köhler, Gunther" docDate="2010" docId="602D87848341EC6DFF41FEF8D9EAFA52" docLanguage="en" docName="zt02354p018.pdf" docOrigin="Zootaxa 2354" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Anolis pentaprion COPE 1862" docType="treatment" docVersion="8" lastPageNumber="13" masterDocId="9C14FFFC834AEC61FFD6FF8FDD27FFA0" masterDocTitle="A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="12" updateTime="1698235538294" updateUser="plazi">
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<mods:title id="845F0A5197DDDEC24424B86847F0C3B2">A revision of the Central American species related to Anolis pentaprion with the resurrection of A. beckeri and the description of a new species (Squamata: Polychrotidae)</mods:title>
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<mods:namePart id="158D28D939482EFB9724A0DE120886F6">Köhler, Gunther</mods:namePart>
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<mods:title id="9E080FC2424583182BBB54D4F4B61C98">Zootaxa</mods:title>
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<mods:date id="86C944EFE3D0B0A7C49F741178E08D8B">2010</mods:date>
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<mods:detail id="C58E8BA2166B53BB37CE2FC4808B5C61" type="volume">
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<mods:number id="FBF5E7BF476E5A5070C75749AA9ABB02">2354</mods:number>
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<mods:identifier id="1114AD624935814ACA0B997FDB10126D" type="DOI">10.5281/zenodo.193523</mods:identifier>
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<mods:identifier id="8AC8E53836033AAC6A360EB38F696BAE" type="GBIF-Dataset">d1368e6e-3dea-4d66-bacd-94c02d3805fa</mods:identifier>
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<mods:identifier id="9330974FFA6356F9605004847A4149F1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F9B1ACC70DC060B7F5E047CB412677E8" type="Zenodo-Dep">193523</mods:identifier>
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<treatment id="602D87848341EC6DFF41FEF8D9EAFA52" ID-DOI="http://doi.org/10.5281/zenodo.6201493" ID-GBIF-Taxon="119383284" ID-Zenodo-Dep="6201493" LSID="urn:lsid:plazi:treatment:602D87848341EC6DFF41FEF8D9EAFA52" httpUri="http://treatment.plazi.org/id/602D87848341EC6DFF41FEF8D9EAFA52" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<subSubSection id="A09E65198341EC6AFF41FEF8DCD9FE31" box="[151,510,375,401]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="E83B36928341EC6AFF41FEF8DCD9FE31" blockId="11.[151,510,375,401]" box="[151,510,375,401]" pageId="11" pageNumber="12">
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<heading id="B37381FE8341EC6AFF41FEF8DCD9FE31" bold="true" box="[151,510,375,401]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
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<taxonomicName id="2F844D118341EC6AFF41FEF8DCD9FE31" authority="COPE 1862" authorityName="COPE" authorityYear="1862" box="[151,510,375,401]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
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<emphasis id="DAF0EA808341EC6AFF41FEF8DCD9FE31" bold="true" box="[151,510,375,401]" pageId="11" pageNumber="12">
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<emphasis id="DAF0EA808341EC6AFF41FEF8DC48FE31" bold="true" box="[151,367,375,401]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
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||
<bibRefCitation id="8C154B638341EC6AFEA1FEF8DC9BFE31" author="COPE" box="[375,444,375,401]" pageId="11" pageNumber="16" refString="Cope, E. D. (1862) Contributions to Neotropical saurology. Proceedings of the Academy of Natural Sciences of Philadelphia, 14, 176 - 188." type="journal article" year="1862">COPE</bibRefCitation>
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||
1862
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="A09E65198341EC6AFF41FE4FDC24FCB4" pageId="11" pageNumber="12" type="reference_group">
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<paragraph id="E83B36928341EC6AFF41FE4FDFC3FE57" blockId="11.[151,1436,448,567]" pageId="11" pageNumber="12">
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<taxonomicName id="2F844D118341EC6AFF41FE4FDCD6FE77" authority="Dunn (1930)" authorityName="Dunn" authorityYear="1930" box="[151,497,448,471]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
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<emphasis id="DAF0EA808341EC6AFF41FE4FDC70FE77" box="[151,343,448,471]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
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||
:
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<bibRefCitation id="8C154B638341EC6AFEB3FE4FDCD6FE77" author="Dunn" box="[357,497,448,471]" pageId="11" pageNumber="16" refString="Dunn, E. R. (1930) Notes on Central American Anolis. Proceedings of the New England Zoological Club, 12, 15 - 24." type="journal article" year="1930">Dunn (1930)</bibRefCitation>
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||
</taxonomicName>
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,
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||
<bibRefCitation id="8C154B638341EC6AFE2AFE4FDF82FE77" author="Schmidt" box="[508,677,448,471]" pageId="11" pageNumber="17" refString="Schmidt, K. P. (1933) Amphibians and reptiles collected by the Smithsonian Biological Survey of the Panama Canal Zone. Smithsonian Miscellaneous Collections, 89, 1 - 20." type="journal article" year="1933">Schmidt (1933)</bibRefCitation>
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||
,
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<bibRefCitation id="8C154B638341EC6AFD66FE4FDE7FFE77" author="Barbour" box="[688,856,448,471]" pageId="11" pageNumber="16" refString="Barbour, T. (1934) The anoles. II. The mainland species from Mexico southward. Bulletin of the Museum of Comparative Zoology, Harvard University, 77, 121 - 155." type="journal article" year="1934">Barbour (1934)</bibRefCitation>
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,
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<bibRefCitation id="8C154B638341EC6AFCB5FE4FD909FE77" box="[867,1070,448,471]" pageId="11" pageNumber="16" refString="Gaige, H. T., Hartweg, N. & Stuart, L. C. (1937) Notes on a collection of amphibians and reptiles from eastern Nicaragua. Occasional Papers Museum of Zoology University of Michigan, 357, 1 - 18." type="journal article">Gaige et al. (1937)</bibRefCitation>
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||
,
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<bibRefCitation id="8C154B638341EC6AFBEFFE4FD80EFE77" author="Taylor" box="[1081,1321,448,471]" pageId="11" pageNumber="17" refString="Taylor, E. H. (1956) A review of the lizards of Costa Rica. - University of Kansas Science Bulletin, 38, 1 - 322." type="journal article" year="1956">Taylor (1956; in part.)</bibRefCitation>
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||
,
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||
<bibRefCitation id="8C154B638341EC6AFAE2FE4FDC44FE57" author="Etheridge" pageId="11" pageNumber="16" refString="Etheridge, R. (1959) The Relationships of the Anoles (Reptilia: Sauria: Iguanidae). An Interpretation Based on Skeletal Morphology. Ph. D. Dissertation, University of Michigan, Ann Arbor, Michigan, U. S. A." type="book" year="1959">Etheridge (1959; in part)</bibRefCitation>
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,
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<bibRefCitation id="8C154B638341EC6AFEB8FE6FDF25FE57" author="Myers" box="[366,514,480,503]" lastPageId="11" lastPageNumber="17" pageId="11" pageNumber="16" refString="Myers, C. W. (1971) Central American lizards related to Anolis pentaprion: Two new species from the Cordillera de Talamanca. American Museum Novitates, 2471, 1 - 40." type="journal article" year="1971">Myers (1971)</bibRefCitation>
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,
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<bibRefCitation id="8C154B638341EC6AFDDBFE6FDFC7FE57" author="Poe" box="[525,736,480,503]" pageId="11" pageNumber="17" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe (2004; in part.)</bibRefCitation>
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||
.
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||
</paragraph>
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<paragraph id="E83B36928341EC6AFF41FD8FDF40FD97" blockId="11.[151,1436,448,567]" pageId="11" pageNumber="12">
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||
<treatmentCitationGroup id="C89411BC8341EC6AFF41FD8FDF40FD97" pageId="11" pageNumber="12">
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||
<taxonomicName id="2F844D118341EC6AFF41FD8FDF76FDB7" authority="BOULENGER, 1890:81" authorityName="BOULENGER" authorityPageNumber="81" authorityYear="1890" box="[151,593,512,535]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="4" phylum="Chordata" rank="species" species="panamensis">
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<emphasis id="DAF0EA808341EC6AFF41FD8FDC47FDB7" box="[151,352,512,535]" italics="true" pageId="11" pageNumber="12">Anolis panamensis</emphasis>
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||
<treatmentCitation id="692510838341EC6AFEB0FD8FDCCDFDB7" author="BOULENGER" box="[358,490,512,535]" page="81" pageId="11" pageNumber="12" year="1890">
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||
<bibRefCitation id="8C154B638341EC6AFEB0FD8FDCCDFDB7" author="BOULENGER" box="[358,490,512,535]" pageId="11" pageNumber="16" refString="Boulenger, G. A. (1890) First report on additions to the lizard collection in the British Museum (Natural History). Proceedings of the Zoological Society London, 1890, 77 - 86." type="proceedings paper" year="1890">BOULENGER</bibRefCitation>
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||
</treatmentCitation>
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||
, 1890:81
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||
</taxonomicName>
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||
; two
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<typeStatus id="373F88308341EC6AFD5DFD8FDFCFFDB7" box="[651,744,512,535]" pageId="11" pageNumber="12" type="syntype">syntypes</typeStatus>
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||
BMNH 1946.8.13.7–8 from “
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<collectingCountry id="909376028341EC6AFBFCFD8FD9A1FDB7" box="[1066,1158,512,535]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
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.”
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<taxonomicName id="2F844D118341EC6AFF41FDAFDC9FFD97" authority="Kohler" authorityName="Kohler" box="[151,440,544,567]" class="Squamata" family="Dactyloidae" genus="Norops" higherTaxonomySource="CoL" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
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<emphasis id="DAF0EA808341EC6AFF41FDAFDC47FD97" box="[151,352,544,567]" italics="true" pageId="11" pageNumber="12">Norops pentaprion</emphasis>
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:
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<bibRefCitation id="8C154B638341EC6AFEB8FDAFDC9FFD97" author="Kohler" box="[366,440,544,567]" pageId="11" pageNumber="16" refString="Kohler, G. (2001) Anfibios y Reptiles de Nicaragua. Herpeton, Offenbach, Germany, 208 pp." type="book" year="2001">Köhler</bibRefCitation>
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</taxonomicName>
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(2001; in part.).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="E83B36928341EC6AFF41FDEADC24FCB4" blockId="11.[151,1437,613,2020]" pageId="11" pageNumber="12">
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||
<emphasis id="DAF0EA808341EC6AFF41FDEADCBCFDDF" bold="true" box="[151,411,613,639]" pageId="11" pageNumber="12">
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||
<typeStatus id="373F88308341EC6AFF41FDEADDD9FDDF" box="[151,254,613,639]" pageId="11" pageNumber="12" type="neotype">Neotype</typeStatus>
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||
designation.
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</emphasis>
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The
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<typeStatus id="373F88308341EC6AFE00FDEADF1BFDDF" box="[470,572,613,639]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
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||
of
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<emphasis id="DAF0EA808341EC6AFDB5FDEADE1BFDDE" box="[611,828,613,638]" italics="true" pageId="11" pageNumber="12">
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||
<taxonomicName id="2F844D118341EC6AFDB5FDEADE1FFDDE" box="[611,824,613,638]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">Anolis pentaprion</taxonomicName>
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,
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</emphasis>
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supposedly designated to be deposited at USNM, is lost (
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<bibRefCitation id="8C154B638341EC6AFF04FD05DC5CFD04" author="Barbour" box="[210,379,650,676]" pageId="11" pageNumber="16" refString="Barbour, T. (1934) The anoles. II. The mainland species from Mexico southward. Bulletin of the Museum of Comparative Zoology, Harvard University, 77, 121 - 155." type="journal article" year="1934">Barbour 1934</bibRefCitation>
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,
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<bibRefCitation id="8C154B638341EC6AFE5EFD05DF3BFD04" author="Myers" box="[392,540,650,676]" lastPageId="11" lastPageNumber="17" pageId="11" pageNumber="16" refString="Myers, C. W. (1971) Central American lizards related to Anolis pentaprion: Two new species from the Cordillera de Talamanca. American Museum Novitates, 2471, 1 - 40." type="journal article" year="1971">Myers 1971</bibRefCitation>
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). K.A. Tighe (pers. comm.
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<date id="9C3A10528341EC6AFCB2FD05D918FD04" box="[868,1087,650,676]" pageId="11" pageNumber="12" value="2009-09-02">2 September 2009</date>
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) wrote that “the
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<typeStatus id="373F88308341EC6AFADFFD05D856FD04" box="[1289,1393,650,676]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
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[of
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<taxonomicName id="2F844D118341EC6AFF41FD3FDC4AFD69" box="[151,365,688,713]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
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<emphasis id="DAF0EA808341EC6AFF41FD3FDC4AFD69" box="[151,365,688,713]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
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</taxonomicName>
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] was apparently never returned to the USNM after Cope described it. There is an old lost
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<typeStatus id="373F88308341EC6AFF41FD5ADDECFD4F" box="[151,203,725,751]" pageId="11" pageNumber="12">type</typeStatus>
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card in Cochran's handwriting that has the information from the original description, but no catalogue number.”
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</paragraph>
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</subSubSection>
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<subSubSection id="A09E65198341EC6AFF12FCAFDC49FB9F" pageId="11" pageNumber="12" type="description">
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<paragraph id="E83B36928341EC6AFF12FCAFDC49FB9F" blockId="11.[151,1437,613,2020]" pageId="11" pageNumber="12">
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To avoid future uncertainties I herewith designate SMF 83608 as the
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<typeStatus id="373F88308341EC6AFC20FCAFD974FC9A" box="[1014,1107,800,826]" pageId="11" pageNumber="12" type="neotype">neotype</typeStatus>
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||
of
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<taxonomicName id="2F844D118341EC6AFBADFCAFD8BBFC9A" authority="Cope." authorityName="Cope." box="[1147,1436,800,826]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
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<emphasis id="DAF0EA808341EC6AFBADFCAFD868FC99" box="[1147,1359,800,825]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
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Cope.
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</taxonomicName>
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SMF 83608 was collected on
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<date id="9C3A10528341EC6AFE22FCCADF88FCFF" box="[500,687,837,863]" pageId="11" pageNumber="12" value="2009-08-12">12 August 2009</date>
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by Johannes Köhler and Arne Schulze at San Rafael, ca.
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<quantity id="2F7C9B778341EC6AFA87FCCAD8BBFCFF" box="[1361,1436,837,863]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="km" value="15.0">15 km</quantity>
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||
S Los Chiles,
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||
<geoCoordinate id="8DB050558341EC6AFEEFFCE5DCECFC24" box="[313,459,874,900]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="1" value="10.73719">10.73719°N</geoCoordinate>
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||
,
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||
<geoCoordinate id="8DB050558341EC6AFE00FCE5DF4BFC24" box="[470,620,874,900]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="1" value="-84.49378">84.49378°W</geoCoordinate>
|
||
,
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||
<quantity id="2F7C9B778341EC6AFDA0FCE5DF95FC24" box="[630,690,874,900]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="m" value="60.0">60 m</quantity>
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elevation, Alajuela Province,
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<collectingCountry id="909376028341EC6AFBC7FCE5D9B3FC24" box="[1041,1172,874,900]" name="Costa Rica" pageId="11" pageNumber="12">Costa Rica</collectingCountry>
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||
. Original field number JJK 182. SMF 83608 is an adult male as indicated by its partially everted hemipenes; SVL 61.0 mm; tail length 79.0 mm (tail complete); head length
|
||
<quantity id="2F7C9B778341EC6AFD7BFC3ADE3CFC6F" box="[685,795,949,975]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="11" pageNumber="12" unit="mm" value="15.4">15.4 mm</quantity>
|
||
; head width
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||
<quantity id="2F7C9B778341EC6AFC65FC3AD934FC6F" box="[947,1043,949,975]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="11" pageNumber="12" unit="mm" value="9.5">9.5 mm</quantity>
|
||
; snout length
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||
<quantity id="2F7C9B778341EC6AFB6CFC3AD83EFC6F" box="[1210,1305,949,975]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="12" unit="mm" value="7.4">7.4 mm</quantity>
|
||
; ear length
|
||
<quantity id="2F7C9B778341EC6AFF41FC55DDD1FC54" box="[151,246,986,1012]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="12" unit="mm" value="0.8">0.8 mm</quantity>
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||
; ear height
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||
<quantity id="2F7C9B778341EC6AFE55FC55DCC4FC54" box="[387,483,986,1012]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="mm" value="1.5">1.5 mm</quantity>
|
||
; interparietal length
|
||
<quantity id="2F7C9B778341EC6AFD0AFC55DE1BFC54" box="[732,828,986,1012]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="12" unit="mm" value="1.3">1.3 mm</quantity>
|
||
; interparietal width
|
||
<quantity id="2F7C9B778341EC6AFBF8FC55D9A9FC54" box="[1070,1166,986,1012]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="12" unit="mm" value="1.1">1.1 mm</quantity>
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||
; axilla–groin–distance 26.0 mm; shank length
|
||
<quantity id="2F7C9B778341EC6AFE6FFB8FDF0DFBBA" box="[441,554,1024,1050]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="11" pageNumber="12" unit="mm" value="12.5">12.5 mm</quantity>
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. Scalation characteristics of SMF 83608 are given in parentheses in the description below.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A09E65198341EC6AFF12FBC5DECAF88A" pageId="11" pageNumber="12" type="diagnosis">
|
||
<paragraph id="E83B36928341EC6AFF12FBC5DECAF88A" blockId="11.[151,1437,613,2020]" pageId="11" pageNumber="12">
|
||
<emphasis id="DAF0EA808341EC6AFF12FBC5DC6FFBC4" bold="true" box="[196,328,1098,1124]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
|
||
A medium-sized species (SVL in largest specimen examined 77.0 mm in males,
|
||
<quantity id="2F7C9B778341EC6AFAF1FBC5D85BFBC4" box="[1319,1404,1098,1124]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="12" unit="mm" value="74.0">74 mm</quantity>
|
||
in females) of the genus
|
||
<taxonomicName id="2F844D118341EC6AFE7AFBFFDCDCFB29" box="[428,507,1136,1161]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="DAF0EA808341EC6AFE7AFBFFDCDCFB29" box="[428,507,1136,1161]" italics="true" pageId="11" pageNumber="12">Anolis</emphasis>
|
||
</taxonomicName>
|
||
(sensu
|
||
<bibRefCitation id="8C154B638341EC6AFD8AFBFFDFFCFB2A" author="Poe" box="[604,731,1136,1162]" pageId="11" pageNumber="17" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89." type="journal article" year="2004">Poe, 2004</bibRefCitation>
|
||
) that differs from all Central American anoles except
|
||
<emphasis id="DAF0EA808341EC6AFA54FBFFDEDFFB0E" italics="true" pageId="11" pageNumber="12">
|
||
<taxonomicName id="2F844D118341EC6AFA54FBFFDDD7FB0E" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="beckeri">A. beckeri</taxonomicName>
|
||
,
|
||
<taxonomicName id="2F844D118341EC6AFF2AFB1ADCE1FB0E" box="[252,454,1173,1198]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="charlesmyersi">A. charlesmyersi</taxonomicName>
|
||
,
|
||
<taxonomicName id="2F844D118341EC6AFE04FB1ADF77FB0E" box="[466,592,1173,1198]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cristifer">A. cristifer</taxonomicName>
|
||
,
|
||
<taxonomicName id="2F844D118341EC6AFD8AFB1ADFCEFB0E" box="[604,745,1173,1198]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="fungosus">A. fungosus</taxonomicName>
|
||
,
|
||
<taxonomicName id="2F844D118341EC6AFD20FB1ADE4FFB0E" box="[758,872,1173,1198]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="salvini">A. salvini</taxonomicName>
|
||
,
|
||
<taxonomicName id="2F844D118341EC6AFCA3FB1ADEDFFB0E" box="[885,1016,1173,1198]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="utilensis">A. utilensis</taxonomicName>
|
||
</emphasis>
|
||
by (1) having extremely short hind legs (fourth toe of adpressed hindlimb usually reaching to level of tympanum; ratio shank length / head length 0.49–0.84); (2) smooth, juxtaposed ventral scales; (3) a relatively short tail (ratio tail length / SVL 1.0–1.5).
|
||
<taxonomicName id="2F844D118341EC6AFF41FA8ADC55FABE" box="[151,370,1285,1310]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
|
||
<emphasis id="DAF0EA808341EC6AFF41FA8ADC55FABE" box="[151,370,1285,1310]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2F844D118341EC6AFDC0FA8ADFB6FABE" box="[534,657,1285,1310]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="beckeri">
|
||
<emphasis id="DAF0EA808341EC6AFDC0FA8ADFB6FABE" box="[534,657,1285,1310]" italics="true" pageId="11" pageNumber="12">A. beckeri</emphasis>
|
||
</taxonomicName>
|
||
by having a pronounced serrated caudal crest (versus lacking a pronounced serrated caudal crest) and by having gorgetal rows with 17–25 narrowly spaced scales (versus gorgetal rows with 4–10 widely spaced scales).
|
||
<taxonomicName id="2F844D118341EC6AFD06FADFDE83FAC9" box="[720,932,1360,1385]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
|
||
<emphasis id="DAF0EA808341EC6AFD06FADFDE83FAC9" box="[720,932,1360,1385]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2F844D118341EC6AFB94FADFD82EFAC9" box="[1090,1289,1360,1385]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="charlesmyersi">
|
||
<emphasis id="DAF0EA808341EC6AFB94FADFD82EFAC9" box="[1090,1289,1360,1385]" italics="true" pageId="11" pageNumber="12">A. charlesmyersi</emphasis>
|
||
</taxonomicName>
|
||
by having a pink dewlap in life with purple pigment between rows of gorgetal scales and regular rows with 17–25 small narrowly spaced gorgetal scales (versus a brick red dewlap with few large widely spaced gorgetal scales scattered across the dewlap or in loosely arranged rows with 4–9 widely spaced scales).
|
||
<taxonomicName id="2F844D118341EC6AFB12FA4FD8BBFA79" box="[1220,1436,1472,1497]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
|
||
<emphasis id="DAF0EA808341EC6AFB12FA4FD8BBFA79" box="[1220,1436,1472,1497]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2F844D118341EC6AFEFAFA6ADC8CFA5E" box="[300,427,1509,1534]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cristifer">
|
||
<emphasis id="DAF0EA808341EC6AFEFAFA6ADC8CFA5E" box="[300,427,1509,1534]" italics="true" pageId="11" pageNumber="12">A. cristifer</emphasis>
|
||
</taxonomicName>
|
||
by having a double row of slightly enlarged middorsal scales, not forming a serrated crest (versus a serrated middorsal crest, at least on posterior portion of dorsum, present in
|
||
<taxonomicName id="2F844D118341EC6AFB6DF984D81EF984" box="[1211,1337,1547,1572]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="cristifer">
|
||
<emphasis id="DAF0EA808341EC6AFB6DF984D81EF984" box="[1211,1337,1547,1572]" italics="true" pageId="11" pageNumber="12">A. cristifer</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="2F844D118341EC6AFA87F984DC3CF9E9" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
|
||
<emphasis id="DAF0EA808341EC6AFA87F984DC3CF9E9" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2F844D118341EC6AFE6BF9BFDF6DF9E9" box="[445,586,1584,1609]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="fungosus">
|
||
<emphasis id="DAF0EA808341EC6AFE6BF9BFDF6DF9E9" box="[445,586,1584,1609]" italics="true" pageId="11" pageNumber="12">A. fungosus</emphasis>
|
||
</taxonomicName>
|
||
by lacking bony parietal protuberances (versus having a pair of small bony parietal protuberances posterior and lateral to interparietal plate).
|
||
<taxonomicName id="2F844D118341EC6AFBD5F9DAD9F9F9CE" box="[1027,1246,1621,1646]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
|
||
<emphasis id="DAF0EA808341EC6AFBD5F9DAD9F9F9CE" box="[1027,1246,1621,1646]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2F844D118341EC6AFA54F9DADDCFF934" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="salvini">
|
||
<emphasis id="DAF0EA808341EC6AFA54F9DADDCFF934" italics="true" pageId="11" pageNumber="12">A. salvini</emphasis>
|
||
</taxonomicName>
|
||
by having smooth, obliquely conical ventral scales (versus keeled), lacking a dark interorbital bar (versus present), and lacking enlarged postcloacal scales (versus males with a pair of greatly enlarged postcloacal scales).
|
||
<taxonomicName id="2F844D118341EC6AFEA9F94ADF76F97E" box="[383,593,1733,1758]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="pentaprion">
|
||
<emphasis id="DAF0EA808341EC6AFEA9F94ADF76F97E" box="[383,593,1733,1758]" italics="true" pageId="11" pageNumber="12">Anolis pentaprion</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2F844D118341EC6AFD3CF94ADE4CF97E" box="[746,875,1733,1758]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="utilensis">
|
||
<emphasis id="DAF0EA808341EC6AFD3CF94ADE4CF97E" box="[746,875,1733,1758]" italics="true" pageId="11" pageNumber="12">A. utilensis</emphasis>
|
||
</taxonomicName>
|
||
by having the proximal subdigital scales of toes differentiated as slightly broadened lamellae (versus those scales granular) and by having a pronounced serrated caudal crest (versus lacking a pronounced serrated caudal crest).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A09E65198341EC6DFF12F8BAD9EAFA52" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="E83B36928341EC6DFF12F8BAD9ABFB67" blockId="11.[151,1437,613,2020]" lastBlockId="12.[151,1438,152,1522]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
<emphasis id="DAF0EA808341EC6AFF12F8BADC46F8EF" bold="true" box="[196,353,1845,1871]" pageId="11" pageNumber="12">Description.</emphasis>
|
||
SVL 52.0–77.0 (62.6±5.7) mm in males, 46.0–64.0 (56.4±5.4) mm in females; for other morphometric data see Table 1; dorsal head scales in internasal, prefrontal, and frontal areas rugose; scales in parietal area rugose to smooth; shallow frontal and parietal depressions present; 5–8 postrostrals (7); anterior nasal divided, lower scale contacting rostral and first supralabial; 5–9 internasals (6); canthal ridge sharply defined; scales comprising supraorbital semicircles ridged, especially anterior ones, largest scale in semicircles larger than largest supraocular scale; supraorbital semicircles well defined, broadly in contact with each other or separated by one scale at narrowest point (in contact); 1–4 scales separating supraorbital semicircles and interparietal at narrowest point (2); interparietal well defined, irregular in outline, longer than wide, larger than size of ear opening; about 6–16 enlarged, smooth to faintly keeled supraocular scales on each side (9), in 2–3 rows; enlarged supraoculars completely separated from supraorbital semicircles; three short superciliaries, posterior one shortest; usually 3 enlarged canthals; 5–9 scales between second canthals (8); 6–15 scales present between posterior canthals (6); loreal region slightly concave, 13–29 mostly smooth or rugose (some keeled) loreal scales (16) in a maximum of 3–6 horizontal rows (3); 7–10 supralabials (9) and 7–10 infralabials (9) to level below center of eye; suboculars smooth to rugose, in broad contact with supralabials; ear opening vertically oval; scales anterior to ear opening slightly larger than those posterior to ear opening; usually 6, occasionally 5 postmentals (6), outer pair largest; gular scales not keeled; male dewlap moderately large, extending to beyond level of axillae onto chest; male dewlap with 5–7 horizontal gorgetalsternal scale rows, about 16–18 scales per row; female dewlap well developed, extending to level of axillae; female dewlap with 4–7 horizontal gorgetal-sternal scale rows, with about 9–15 scales per row; about 2 middorsal scale rows slightly enlarged, mostly smooth, dorsal scales lateral to middorsal series grading into granular lateral scales; no enlarged scales among granular laterals; lateral scales granular, slightly smaller than largest dorsal scales; about 73–108 dorsal scales along vertebral midline between levels of axillae and groin in males (73), about
|
||
<quantity id="2F7C9B778346EC6DFEBCFC9DDCF8FC8C" box="[362,479,786,812]" metricMagnitude="0" metricUnit="m" metricValue="2.5146" metricValueMax="2.7939999999999996" metricValueMin="2.2352" pageId="12" pageNumber="13" unit="in" value="99.0" valueMax="110.0" valueMin="88.0">88–110 in</quantity>
|
||
females; about 56–80 dorsal scales along vertebral midline contained in one head length in males (52), about
|
||
<quantity id="2F7C9B778346EC6DFE36FCB7DF6EFCF2" box="[480,585,824,850]" metricMagnitude="0" metricUnit="m" metricValue="1.4605" metricValueMax="1.778" metricValueMin="1.143" pageId="12" pageNumber="13" unit="in" value="57.5" valueMax="70.0" valueMin="45.0">45–70 in</quantity>
|
||
females; ventral scales on midsection slightly larger than largest dorsal scales; ventral body scales smooth, obliquely conical, juxtaposed; about 67–86 ventral scales along midventral line between levels of axilla and groin in males (84), about
|
||
<quantity id="2F7C9B778346EC6DFC01FC0DD918FC3C" box="[983,1087,898,924]" metricMagnitude="0" metricUnit="m" metricValue="2.0065999999999997" metricValueMax="2.3622" metricValueMin="1.6509999999999998" pageId="12" pageNumber="13" unit="in" value="79.0" valueMax="93.0" valueMin="65.0">65–93 in</quantity>
|
||
females; about 34–62 ventral scales contained in one head length in males (44), about
|
||
<quantity id="2F7C9B778346EC6DFC84FC27DE98FC62" box="[850,959,936,962]" metricMagnitude="0" metricUnit="m" metricValue="1.1176" metricValueMax="1.3208" metricValueMin="0.9144" pageId="12" pageNumber="13" unit="in" value="44.0" valueMax="52.0" valueMin="36.0">36–52 in</quantity>
|
||
females; about 134–190 scales around midbody in males (154), about
|
||
<quantity id="2F7C9B778346EC6DFDC1FC42DF86FC47" box="[535,673,973,999]" metricMagnitude="0" metricUnit="m" metricValue="3.7845999999999997" metricValueMax="4.114800000000001" metricValueMin="3.4543999999999997" pageId="12" pageNumber="13" unit="in" value="149.0" valueMax="162.0" valueMin="136.0">136–162 in</quantity>
|
||
females; tubelike axillary pocket absent; precloacal scales not keeled; enlarged postcloacal scales absent; tail oval at about midlength; basal subcaudal scales smooth to faintly keeled; lateral caudal scales keeled, homogeneous, although an indistinct division in segments is discernible; dorsal medial caudal scale row enlarged, keeled, not forming a crest; 2 median subcaudal scale rows distinctly enlarged, keeled; most scales on lateral surface of antebrachium smooth; 29–33 subdigital lamellae on Phalanges II–IV of Toe IV of hind limbs (29); 6–9 subdigital scales on Phalanx I of Toe IV of hind limbs (6); longest toe of adpressed hind limb usually reaching to shoulder region.
|
||
</paragraph>
|
||
<paragraph id="E83B36928346EC6DFF12FB5DDF21FA07" blockId="12.[151,1438,152,1522]" pageId="12" pageNumber="13">
|
||
The coloration in life of an adult male (SMF 80961) from Bartola, Department Río San Juan,
|
||
<collectingCountry id="909376028346EC6DFACAFB5DD8BFFB4C" box="[1308,1432,1234,1260]" name="Nicaragua" pageId="12" pageNumber="13">Nicaragua</collectingCountry>
|
||
, was recorded as follows: Dorsal and lateral surfaces of body, limbs and anterior half of tail Clay Color (123B) with Raw Umber (223) and Beige (219D) blotches; dorsal surface of head Cinnamon Drab (119C); dewlap Vinaceous (4) with rows of Magenta (2) blotches and dirty white scales. The coloration of the dewlap in life of another adult male (SMF 82100) from the same locality was recorded as Pink (7) with rows of Magenta (2) blotches and dirty white scales.
|
||
</paragraph>
|
||
<paragraph id="E83B36928346EC6DFF12FA3DD9EAFA52" blockId="12.[151,1438,152,1522]" pageId="12" pageNumber="13">
|
||
<emphasis id="DAF0EA808346EC6DFF12FA3DDF33FA6C" bold="true" box="[196,532,1458,1484]" pageId="12" pageNumber="13">Geographic distribution:</emphasis>
|
||
Along the Caribbean versant of Lower Central
|
||
<collectingCountry id="909376028346EC6DFBA0FA3DD9C5FA6C" box="[1142,1250,1458,1484]" name="United States of America" pageId="12" pageNumber="13">America</collectingCountry>
|
||
from southern
|
||
<collectingCountry id="909376028346EC6DFF41FA57DC28FA52" box="[151,271,1496,1522]" name="Nicaragua" pageId="12" pageNumber="13">Nicaragua</collectingCountry>
|
||
to northwestern
|
||
<collectingCountry id="909376028346EC6DFE03FA57DF6AFA52" box="[469,589,1496,1522]" name="Colombia" pageId="12" pageNumber="13">Colombia</collectingCountry>
|
||
; also on Pacific versant in central and eastern
|
||
<collectingCountry id="909376028346EC6DFBBFFA57D9EDFA52" box="[1129,1226,1496,1522]" name="Panama" pageId="12" pageNumber="13">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |