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<document ID-DOI="10.1007/s12225-021-09991-y" ID-Zenodo-Dep="7615581" approvalRequired="22" approvalRequired_for_document="1" approvalRequired_for_taxonomicNames="10" approvalRequired_for_treatments="11" checkinTime="1675780091056" checkinUser="valdenar" docAuthor="Bongcheewin, Bhanubong, Ingrouille, Martin J. &amp; Paton, Alan J." docDate="2022" docId="2733879DFFB41C310C1092541F1BFE24" docLanguage="en" docName="KewBull.77.2792.pdf.imf" docOrigin="Kew Bulletin 77 (1)" docSource="http://dx.doi.org/10.1007/s12225-021-09991-y" docTitle="Gomphostemma keralense Vioek., Gopalan" docType="treatment" docVersion="3" lastPageNumber="45" masterDocId="DB0AFFE5FFA51C230F3593701E44FFB2" masterDocTitle="A revision of Gomphostemma (Lamiaceae)" masterLastPageNumber="92" masterPageNumber="27" pageNumber="44" updateTime="1675799494316" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of Gomphostemma (Lamiaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bongcheewin, Bhanubong</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ingrouille, Martin J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paton, Alan J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Kew Bulletin</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>77</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1007/s12225-021-09991-y</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1007/s12225-021-09991-y</mods:identifier>
<mods:identifier type="Zenodo-Dep">7615581</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:2733879DFFB41C310C1092541F1BFE24" httpUri="http://treatment.plazi.org/id/2733879DFFB41C310C1092541F1BFE24" lastPageId="18" lastPageNumber="45" pageId="17" pageNumber="44">
<subSubSection box="[805,1321,292,314]" pageId="17" pageNumber="44" type="nomenclature">
<paragraph blockId="17.[805,1399,292,467]" box="[805,1321,292,314]" pageId="17" pageNumber="44">
<emphasis bold="true" box="[805,1140,292,314]" pageId="17" pageNumber="44">
3.
<taxonomicName ID-CoL="3GTCR" authority="Vioek., Gopalan" authorityName="Vioek., Gopalan" box="[840,1321,292,314]" class="Magnoliopsida" family="Lamiaceae" genus="Gomphostemma" kingdom="Plantae" order="Lamiales" pageId="17" pageNumber="44" phylum="Tracheophyta" rank="species" species="keralense">
Gomphostemma keralense
<emphasis box="[1155,1215,292,313]" italics="true" pageId="17" pageNumber="44">Vioek.</emphasis>
, Gopalan
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="44" type="reference_group">
<paragraph blockId="17.[805,1399,292,467]" pageId="17" pageNumber="44">
&amp; R. Ansari (
<bibRefCitation author="Vivekananthan, K. &amp; Gopalan, R. &amp; Ansari, R." box="[889,1241,323,345]" pageId="17" pageNumber="44" pagination="189 - 190" part="38" refId="ref79134" refString="Vivekananthan, K., Gopalan, R. &amp; Ansari, R. (1983). A new species of Gomphostemma (Labiatae) from Kerala, India. Keae 3 ull. 38: 189 - 190." title="A new species of Gomphostemma (Labiatae) from Kerala, India" type="journal article" year="1983">
Vivekananthan
<emphasis box="[1067,1120,323,345]" italics="true" pageId="17" pageNumber="44">et al.</emphasis>
1983: 189
</bibRefCitation>
).
<materialsCitation pageId="17" pageNumber="44">
Type: India, Kerala, Idikki [Idukki]?istr., way to Mangaladevi Temple, 930 m, 10 Oct. 1956,
<emphasis box="[1169,1321,384,405]" italics="true" pageId="17" pageNumber="44">Vioekananthan</emphasis>
28638 (holotype CAL (CAL0000006806);
</materialsCitation>
<materialsCitation pageId="17" pageNumber="44">isotypes K [K000826333], MH n.v.).</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="44" type="description">
<paragraph blockId="17.[805,1399,506,1414]" pageId="17" pageNumber="44">
Perennial herb.
<emphasis box="[986,1040,507,528]" italics="true" pageId="17" pageNumber="44">Stems</emphasis>
erect, woody at base, branched, obtusely quadrangular with longitudinal deep grooves, with dense sessile stellate hairs.
<emphasis box="[1209,1276,568,589]" italics="true" pageId="17" pageNumber="44">Leaoes</emphasis>
petiolate, chartaceous; blades elliptic or elliptic-ovate, 13 23 × 6 10 cm, apex acute, margin serrate-dentate, base attenuate, upper side with sparsely simple hairs and sessile stellate hairs, confined on midrib and veins; lower side pale green tomentose with sessile stellate hairs confined on midrib and veins; petioles 30 90 mm long.
<emphasis box="[957,1078,781,803]" italics="true" pageId="17" pageNumber="44">Inflorescence</emphasis>
axillary with opposite cymes sessile, congested with axis not visible between flowers, inserted at the upper nodes at which leaves are usually present; verticils 2 3-flowered; bracts rhomboid or elliptic, 3 10 × 2 5 mm, apex acute, margin shallowly serrate at the apical part and entire at the basal part, with sessile stellate hairs on both sides; bracteoles narrowly ovate, 3 9 mm long.
<emphasis box="[1057,1205,995,1016]" italics="true" pageId="17" pageNumber="44">Flowering calyx</emphasis>
infundibular, 9 11 mm long, 10-ridged, outside creamy-white tomentose with sessile stellate hairs, inside with adpressed simple hairs; tube 2 3 mm long; lobes triangular, 3 6 mm long, apex acute.
<emphasis box="[999,1133,1118,1139]" italics="true" pageId="17" pageNumber="44">Fruiting calyx</emphasis>
ovate-infundibular, 12 12 mm long; tube 6 5 mm long; lobes equal, triangular, 6 5 mm long, apex acuminate.
<emphasis box="[1152,1221,1179,1200]" italics="true" pageId="17" pageNumber="44">Corolla</emphasis>
8 10 mm long, throat narrow, not enlarged, sometimes gradually dilated towards the throat, outside glabrous below and with sessile stellate hairs above, inside glabrous; tube slender, straight, 3 5 mm long, included inside calyx.
<emphasis box="[805,986,1331,1352]" italics="true" pageId="17" pageNumber="44">Staminal filaments</emphasis>
slightly fleshy.
<emphasis box="[1151,1193,1331,1352]" italics="true" pageId="17" pageNumber="44">Style</emphasis>
slender, as long as corolla.
<emphasis box="[898,965,1362,1383]" italics="true" pageId="17" pageNumber="44">Nutlets</emphasis>
solitary, elliptic, 6 mm long, sparsely tomentose.
</paragraph>
</subSubSection>
<subSubSection box="[805,1206,1454,1476]" pageId="17" pageNumber="44" type="distribution">
<paragraph blockId="17.[805,1399,1454,1903]" box="[805,1206,1454,1476]" pageId="17" pageNumber="44">
<emphasis bold="true" box="[805,952,1454,1475]" pageId="17" pageNumber="44">DISTRIBUTION.</emphasis>
India (Kerala).
<figureCitation box="[1131,1201,1454,1475]" captionStart="Map 1" captionStartId="14.[189,236,1005,1026]" captionTargetBox="[189,1438,193,988]" captionTargetId="figure-467@14.[189,1439,192,989]" captionTargetPageId="14" captionText="Map 1. Distribution range of Gocphostecca aĸorense, G. crinituc, G. dolichoĸotrys, G. eriocarpon and G. keralense." figureDoi="http://doi.org/10.5281/zenodo.7615600" httpUri="https://zenodo.org/record/7615600/files/figure.png" pageId="17" pageNumber="44">Map 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="44" type="materials_examined">
<paragraph blockId="17.[805,1399,1454,1903]" pageId="17" pageNumber="44">
<emphasis bold="true" box="[805,1134,1484,1506]" pageId="17" pageNumber="44">
SPECIMENS EXAMINED.
<materialsCitation pageId="17" pageNumber="44">
INDIA. Kerala: Idukki?istr., Paermade [Peermade, 9.3665°N 56.9833°E],
<emphasis box="[1320,1399,1515,1536]" italics="true" pageId="17" pageNumber="44">Meebold</emphasis>
12921 (K);
</materialsCitation>
</emphasis>
<materialsCitation pageId="17" pageNumber="44">
Udumbansholay [Udumbanchola, 9.9°, 55.1665°],
<emphasis box="[933,1012,1576,1597]" italics="true" pageId="17" pageNumber="44">Meebold</emphasis>
13013 (K);
</materialsCitation>
<materialsCitation pageId="17" pageNumber="44">
way to Mangaladevi Temple, 930 m, 10 Oct. 1956,
<emphasis box="[1169,1321,1606,1627]" italics="true" pageId="17" pageNumber="44">Vioekananthan</emphasis>
28638 (holotype CAL; isotypes K).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="44" type="biology_ecology">
<paragraph blockId="17.[805,1399,1454,1903]" pageId="17" pageNumber="44">
<emphasis bold="true" box="[805,900,1667,1689]" pageId="17" pageNumber="44">HABITAT.</emphasis>
In evergreen or semi-evergreen forest; alt. 930 m.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="44" type="conservation">
<paragraph blockId="17.[805,1399,1454,1903]" pageId="17" pageNumber="44">
<emphasis bold="true" box="[805,1077,1729,1750]" pageId="17" pageNumber="44">CONSERVATION STATUS.</emphasis>
<taxonomicName box="[1094,1365,1728,1749]" class="Magnoliopsida" family="Lamiaceae" genus="Gomphostemma" kingdom="Plantae" order="Lamiales" pageId="17" pageNumber="44" phylum="Tracheophyta" rank="species" species="keralense">
<emphasis box="[1094,1365,1728,1749]" italics="true" pageId="17" pageNumber="44">Gomphostemma keralense</emphasis>
</taxonomicName>
is known from three collections collected from Idukki?istr. and has an EOO of 231.01 km
<superScript attach="left" box="[1209,1219,1784,1798]" fontSize="6" pageId="17" pageNumber="44">2</superScript>
and AOO of 12 km
<superScript attach="right" box="[840,850,1814,1828]" fontSize="6" pageId="17" pageNumber="44">2</superScript>
. Google Earth imagery shows some evidence of habitat loss. Hence, the Vulnerable (VU) category is applied (VU?2).
</paragraph>
</subSubSection>
<subSubSection box="[189,713,201,223]" pageId="18" pageNumber="45" type="biology_ecology">
<paragraph blockId="18.[189,783,201,406]" box="[189,713,201,223]" pageId="18" pageNumber="45">
<emphasis bold="true" box="[189,315,202,222]" pageId="18" pageNumber="45">PHENOLOGY</emphasis>
. Flowering and fruiting in October.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="45" type="discussion">
<paragraph blockId="18.[189,783,201,406]" pageId="18" pageNumber="45">
<emphasis bold="true" box="[189,263,232,253]" pageId="18" pageNumber="45">NOTES.</emphasis>
<taxonomicName box="[277,531,231,252]" class="Magnoliopsida" family="Lamiaceae" genus="Gomphostemma" kingdom="Plantae" order="Lamiales" pageId="18" pageNumber="45" phylum="Tracheophyta" rank="species" species="keralense">
<emphasis box="[277,531,231,252]" italics="true" pageId="18" pageNumber="45">Gomphostemma keralense</emphasis>
</taxonomicName>
is recognised by the axillary inflorescence with a short corolla included in the calyx. It is morphologically similar to
<taxonomicName box="[640,782,292,314]" class="Magnoliopsida" family="Lamiaceae" genus="Gomphostemma" kingdom="Plantae" order="Lamiales" pageId="18" pageNumber="45" phylum="Tracheophyta" rank="species" species="heyneanum">
<emphasis box="[640,782,292,314]" italics="true" pageId="18" pageNumber="45">G. heyneanum</emphasis>
</taxonomicName>
in having an included corolla and is found in the same area of South India. But it differs by having axillary inflorescences.
</paragraph>
</subSubSection>
</treatment>
</document>