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<document ID-DOI="http://doi.org/10.5281/zenodo.4762843" ID-GBIF-Dataset="97d9d905-b4c2-4be0-9886-eb21c200d2b8" ID-GBIF-Taxon="182073706" ID-ISSN="1854-0392" ID-Zenodo-Dep="4762843" checkinTime="1620936722352" checkinUser="felipe" docAuthor="Baumann, Richard W. &amp; Lee, Jonathan J." docDate="2009" docId="2373BB7A3F7B3F018EFE1C22FE5EF250" docLanguage="en" docName="Illiesia.5.1.1-10.pdf" docOrigin="Illiesia 5 (1)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Isoperla karuk Baumann &amp; Lee 2009, sp.n." docType="treatment" docVersion="5" lastPageId="8" lastPageNumber="9" masterDocId="DF4AC3023F7F3F098D0B1979FFB9F722" masterDocTitle="Two Interesting New Species Of Isoperla From Northern California (Plecoptera: Perlodidae)" masterLastPageNumber="10" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1643565812222" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two Interesting New Species Of Isoperla From Northern California (Plecoptera: Perlodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baumann, Richard W.</mods:namePart>
<mods:affiliation>Monte L. Bean Life Science Museum, Department of Biology, Brigham Young University, Provo, UT 84602 E-mail: richard _ baumann @ byu. edu</mods:affiliation>
<mods:nameIdentifier type="email">richard_baumann@byu.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Jonathan J.</mods:namePart>
<mods:affiliation>2337 15 Street Eureka, California 95501 E-mail: jlee @ humboldt 1. com</mods:affiliation>
<mods:nameIdentifier type="email">jlee@humboldt</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:end>10</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758990</mods:identifier>
<mods:identifier type="GBIF-Dataset">97d9d905-b4c2-4be0-9886-eb21c200d2b8</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4758990</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4762843" ID-GBIF-Taxon="182073706" ID-Zenodo-Dep="4762843" LSID="urn:lsid:plazi:treatment:2373BB7A3F7B3F018EFE1C22FE5EF250" httpUri="http://treatment.plazi.org/id/2373BB7A3F7B3F018EFE1C22FE5EF250" lastPageId="8" lastPageNumber="9" pageId="4" pageNumber="5">
<subSubSection box="[1013,1265,1371,1395]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph blockId="4.[1013,1265,1371,1395]" box="[1013,1265,1371,1395]" pageId="4" pageNumber="5">
<heading bold="true" box="[1013,1265,1371,1395]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="5" reason="2">
<emphasis bold="true" box="[1013,1265,1371,1395]" pageId="4" pageNumber="5">
<taxonomicName authority="Baumann &amp; Lee, 2009" authorityName="Baumann &amp; Lee" authorityYear="2009" box="[1013,1191,1371,1395]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="karuk" status="sp.n.">
<emphasis bold="true" box="[1013,1191,1371,1395]" italics="true" pageId="4" pageNumber="5">Isoperla karuk</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1205,1265,1371,1395]" pageId="4" pageNumber="5" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1066,1212,1407,1431]" pageId="4" pageNumber="5" type="description">
<paragraph blockId="4.[1066,1212,1407,1431]" box="[1066,1212,1407,1431]" pageId="4" pageNumber="5">
(
<figureCitation box="[1075,1203,1407,1431]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="5.[189,240,1796,1820]" captionStartId-1="6.[189,240,1796,1820]" captionTargetBox-0="[249,1383,189,1746]" captionTargetBox-1="[249,1383,189,1748]" captionTargetId-0="figure-42@5.[249,1383,189,1748]" captionTargetId-1="figure-42@6.[249,1383,189,1748]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figs. 1724. Isoperla karuk: 17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up." captionText-1="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi-0="http://doi.org/10.5281/zenodo.4758996" figureDoi-1="http://doi.org/10.5281/zenodo.4758998" httpUri-0="https://zenodo.org/record/4758996/files/figure.png" httpUri-1="https://zenodo.org/record/4758998/files/figure.png" pageId="4" pageNumber="5">Figs. 1732</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="materials_examined">
<paragraph blockId="4.[835,1443,1477,1928]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[835,1079,1478,1502]" pageId="4" pageNumber="5">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111527304" collectingDate="2006-03-29" collectorName="J. J. Lee" county="Humboldt County" location="California" municipality="Aikens Creek" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[1090,1203,1478,1502]" pageId="4" pageNumber="5">Holotype</typeStatus>
<specimenCount box="[1214,1234,1477,1502]" pageId="4" pageNumber="5" type="male"></specimenCount>
from
<location LSID="urn:lsid:plazi:treatment:2373BB7A3F7B3F018EFE1C22FE5EF250:AE055CB73F7B3F0D88281CBFFA27F2FC" box="[1315,1438,1478,1502]" county="Humboldt County" municipality="Aikens Creek" name="California" pageId="4" pageNumber="5">California</location>
,
<collectingCounty box="[835,1065,1514,1538]" pageId="4" pageNumber="5">Humboldt County</collectingCounty>
,
<locationDeviation pageId="4" pageNumber="5">Klamath River @ junction of Aikens Creek</locationDeviation>
,
<date box="[1009,1113,1549,1573]" pageId="4" pageNumber="5" value="2006-03-29">
<collectingDate box="[1009,1113,1549,1573]" pageId="4" pageNumber="5" value="2006-03-29">29III06</collectingDate>
</date>
,
<collectorName box="[1122,1206,1550,1573]" pageId="4" pageNumber="5">J.J. Lee</collectorName>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111527302" collectingDate="2006-03-29" collectorName="J. J. Lee" county="Humboldt Co." location="California" municipality="Aikens Creek" pageId="4" pageNumber="5" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus box="[1216,1335,1550,1574]" pageId="4" pageNumber="5">Paratypes</typeStatus>
(
<collectorName box="[1349,1438,1549,1573]" pageId="4" pageNumber="5">J. J. Lee</collectorName>
, collector):
<location LSID="urn:lsid:plazi:treatment:2373BB7A3F7B3F018EFE1C22FE5EF250:AE055CB73F7B3F0D8EC51F48FBF0F16B" box="[974,1097,1585,1609]" county="Humboldt Co." municipality="Aikens Creek" name="California" pageId="4" pageNumber="5">California</location>
,
<collectingCounty box="[1108,1282,1585,1609]" pageId="4" pageNumber="5">Humboldt Co.</collectingCounty>
, same data as
<typeStatus box="[835,944,1620,1644]" pageId="4" pageNumber="5">holotype</typeStatus>
,
<date box="[957,1060,1620,1644]" pageId="4" pageNumber="5" value="2006-03-29">
<collectingDate box="[957,1060,1620,1644]" pageId="4" pageNumber="5" value="2006-03-29">29III06</collectingDate>
</date>
,
<specimenCount box="[1073,1114,1619,1644]" pageId="4" pageNumber="5" type="male">2 ♂</specimenCount>
</materialsCitation>
; 27II08,
<specimenCount box="[1236,1277,1619,1644]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
; 6III08,
<specimenCount box="[1394,1436,1619,1644]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
;
<materialsCitation ID-GBIF-Occurrence="3111527305" collectingDate="2006-04-19" location="Mad River" municipality="Mad River" pageId="4" pageNumber="5" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:2373BB7A3F7B3F018EFE1C22FE5EF250:AE055CB73F7B3F0D8E481F01FC7AF1B2" box="[835,963,1656,1680]" municipality="Mad River" name="Mad River" pageId="4" pageNumber="5">Mad River</location>
@
<collectingMunicipality box="[996,1123,1656,1680]" pageId="4" pageNumber="5">Mad River</collectingMunicipality>
Fish Hatchery,
<date box="[1314,1418,1656,1680]" pageId="4" pageNumber="5" value="2006-04-19">
<collectingDate box="[1314,1418,1656,1680]" pageId="4" pageNumber="5" value="2006-04-19">19IV06</collectingDate>
</date>
,
<specimenCount pageId="4" pageNumber="5" type="male">2 ♂</specimenCount>
,
<specimenCount box="[868,907,1690,1715]" pageId="4" pageNumber="5" type="female">3 ♀</specimenCount>
</materialsCitation>
; 7IV07
<specimenCount box="[1013,1052,1690,1715]" pageId="4" pageNumber="5" type="male">2 ♂</specimenCount>
; 18IV07,
<specimenCount box="[1179,1218,1690,1715]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
,
<specimenCount box="[1231,1270,1690,1715]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
; 19IV07,
<specimenCount box="[1397,1436,1690,1715]" pageId="4" pageNumber="5" type="male">6 ♂</specimenCount>
; 5V07,
<specimenCount box="[927,966,1726,1751]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
; 8IV08,
<specimenCount box="[1081,1120,1726,1751]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
; 12V08,
<specimenCount box="[1239,1279,1726,1751]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
;
<materialsCitation ID-GBIF-Occurrence="3111527310" collectingDate="2006-03-20" location="Mad River" municipality="Mad River Rd." pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:2373BB7A3F7B3F018EFE1C22FE5EF250:AE055CB73F7B3F0D88071FC6FA33F1F5" box="[1292,1418,1727,1751]" municipality="Mad River Rd." name="Mad River" pageId="4" pageNumber="5">Mad River</location>
@ 1
<superScript attach="left" box="[849,861,1763,1777]" fontSize="6" pageId="4" pageNumber="5">st</superScript>
curve
<collectingMunicipality box="[943,1121,1762,1786]" pageId="4" pageNumber="5">Mad River Rd.</collectingMunicipality>
,
<date box="[1133,1236,1762,1786]" pageId="4" pageNumber="5" value="2006-03-20">
<collectingDate box="[1133,1236,1762,1786]" pageId="4" pageNumber="5" value="2006-03-20">20III06</collectingDate>
</date>
,
<specimenCount box="[1247,1287,1761,1786]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
</materialsCitation>
; 29,30III07,
<specimenCount box="[835,888,1797,1822]" pageId="4" pageNumber="5" type="male">12 ♂</specimenCount>
,
<specimenCount box="[902,955,1797,1822]" pageId="4" pageNumber="5" type="female">10 ♀</specimenCount>
; 1IV07,
<specimenCount box="[1069,1109,1797,1822]" pageId="4" pageNumber="5" type="male">2 ♂</specimenCount>
,
<specimenCount box="[1123,1162,1797,1822]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
; 3IV07,
<specimenCount box="[1277,1316,1797,1822]" pageId="4" pageNumber="5" type="male">3 ♂</specimenCount>
,
<specimenCount box="[1330,1369,1797,1822]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
; 4IV 07,
<specimenCount box="[875,913,1832,1857]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
(reared); 7IV07,
<specimenCount box="[1123,1161,1832,1857]" pageId="4" pageNumber="5" type="male">4 ♂</specimenCount>
; 28IV07,
<specimenCount box="[1286,1325,1832,1857]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
; 17III08,
<specimenCount box="[835,874,1868,1893]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
.
<materialsCitation ID-GBIF-Occurrence="3111527306" location="California Academy of Sciences" pageId="4" pageNumber="5" specimenCount="1" typeStatus="holotype">
<typeStatus box="[887,1000,1869,1893]" pageId="4" pageNumber="5">Holotype</typeStatus>
deposited at the California Academy of Sciences
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758996" ID-Zenodo-Dep="4758996" httpUri="https://zenodo.org/record/4758996/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,240,1796,1820]" targetBox="[249,1383,189,1746]" targetPageId="5">
<paragraph blockId="5.[189,1443,1796,1892]" pageId="5" pageNumber="6">
Figs. 1724.
<emphasis box="[331,492,1796,1820]" italics="true" pageId="5" pageNumber="6">
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[331,486,1796,1820]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="karuk">Isoperla karuk</taxonomicName>
:
</emphasis>
17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758998" ID-Zenodo-Dep="4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,240,1796,1820]" targetBox="[249,1383,189,1748]" targetPageId="6">
<paragraph blockId="6.[189,1443,1796,1892]" pageId="6" pageNumber="7">
Figs. 2532.
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[332,487,1796,1820]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="karuk">
<emphasis box="[332,487,1796,1820]" italics="true" pageId="6" pageNumber="7">Isoperla karuk</emphasis>
</taxonomicName>
: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hair like addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up.
</paragraph>
</caption>
<subSubSection pageId="7" pageNumber="8" type="description">
<paragraph blockId="7.[189,798,198,1928]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[189,258,198,222]" pageId="7" pageNumber="8">Male.</emphasis>
Macropterous. Length of forewings
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="7" pageNumber="8" unit="mm" value="9.5" valueMax="10.5" valueMin="8.5">8.510.5 mm</quantity>
; length of body 8.09.0 mm. General body color medium brown. Head with ocelli and base of antennae connected by dark brown M shaped pigment band; yellow patch forward of anterior ocellus and between lateral ocelli and compound eye; interocellar space ranging from mostly yellow to an oval, central yellow spot; occiput light brown, with lateral light oval patch broken by brown reticulations, light patch broadly to barely connected to yellow patch adjacent to eyes; antennae brown. Pronotum with wide, median light stripe; disks light brown to lateral margins, rugosities darker brown; anterior and posterior margins cream to light brown. Meso and metanota dark brown, large yellow patch anteriorly between mesonotal wing bases. Wings hyaline, veins brown, fumose in coastal space. Legs light brown, femur with proximal dorsal brown mark and longitudinal dorsal brown stripe, tibia with incomplete narrow brown band in proximal onefourth. Abdominal terga medium brown dorsally, pale laterally, a pair of median light marks on first few terga, broken brown stripes appearing weakly laterally; abdominal sterna cream colored with brown marks at pleural folds, medial and mediolateral pair of brown spots evident to varying degrees. Vesicle short, broadly rounded (
<figureCitation box="[700,785,1122,1146]" captionStart="Figs" captionStartId="6.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@6.[249,1383,189,1748]" captionTargetPageId="6" captionText="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi="http://doi.org/10.5281/zenodo.4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="7" pageNumber="8">Fig. 29</figureCitation>
). Cerci brown, segments with one long posteroventral seta. Paraprocts short and stout, apically pointed, barely curving over tenth tergum (
<figureCitation box="[586,664,1229,1253]" captionStart="Figs" captionStartId="6.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@6.[249,1383,189,1748]" captionTargetPageId="6" captionText="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi="http://doi.org/10.5281/zenodo.4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="7" pageNumber="8">Fig. 30</figureCitation>
). Aedeagus (
<figureCitation box="[197,378,1264,1288]" captionStart="Figs" captionStartId="5.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1746]" captionTargetId="figure-42@5.[249,1383,189,1748]" captionTargetPageId="5" captionText="Figs. 1724. Isoperla karuk: 17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up." figureDoi="http://doi.org/10.5281/zenodo.4758996" httpUri="https://zenodo.org/record/4758996/files/figure.png" pageId="7" pageNumber="8">Figs. 17 &amp; 19</figureCitation>
) with a large posteromedial bihemispherical lobe bearing long, rust colored anteriorly directed spinules (
<figureCitation box="[537,629,1335,1359]" captionStart="Figs" captionStartId="5.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1746]" captionTargetId="figure-42@5.[249,1383,189,1748]" captionTargetPageId="5" captionText="Figs. 1724. Isoperla karuk: 17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up." figureDoi="http://doi.org/10.5281/zenodo.4758996" httpUri="https://zenodo.org/record/4758996/files/figure.png" pageId="7" pageNumber="8">Figs. 18</figureCitation>
&amp;
<figureCitation box="[668,697,1335,1359]" captionStart="Figs" captionStartId="6.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@6.[249,1383,189,1748]" captionTargetPageId="6" captionText="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi="http://doi.org/10.5281/zenodo.4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="7" pageNumber="8">25</figureCitation>
); a thin, sclerotized, clavate process (lateral view), projecting from midventral margin of bi hemispherical lobe, keeled dorsally and ventrally, flattening apically, heavily sculptured and spinulate (
<figureCitation box="[587,713,1477,1501]" captionStart="Figs" captionStartId="5.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1746]" captionTargetId="figure-42@5.[249,1383,189,1748]" captionTargetPageId="5" captionText="Figs. 1724. Isoperla karuk: 17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up." figureDoi="http://doi.org/10.5281/zenodo.4758996" httpUri="https://zenodo.org/record/4758996/files/figure.png" pageId="7" pageNumber="8">Figs. 2124</figureCitation>
); a pair of lobes positioned laterally on either side of bihemispherical lobe, bearing rust colored spinules directed posteriorly (
<figureCitation box="[434,555,1584,1608]" captionStart="Figs" captionStartId="5.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1746]" captionTargetId="figure-42@5.[249,1383,189,1748]" captionTargetPageId="5" captionText="Figs. 1724. Isoperla karuk: 17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up." figureDoi="http://doi.org/10.5281/zenodo.4758996" httpUri="https://zenodo.org/record/4758996/files/figure.png" pageId="7" pageNumber="8">Figs.17, 20</figureCitation>
&amp;
<figureCitation box="[585,615,1584,1608]" captionStart="Figs" captionStartId="6.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@6.[249,1383,189,1748]" captionTargetPageId="6" captionText="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi="http://doi.org/10.5281/zenodo.4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="7" pageNumber="8">26</figureCitation>
); a patch of thin light colored spinules located below clavate process, narrowed and fringed apically (
<figureCitation box="[584,679,1655,1679]" captionStart="Figs" captionStartId="5.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1746]" captionTargetId="figure-42@5.[249,1383,189,1748]" captionTargetPageId="5" captionText="Figs. 1724. Isoperla karuk: 17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up." figureDoi="http://doi.org/10.5281/zenodo.4758996" httpUri="https://zenodo.org/record/4758996/files/figure.png" pageId="7" pageNumber="8">Figs. 22</figureCitation>
&amp;
<figureCitation box="[723,753,1655,1679]" captionStart="Figs" captionStartId="6.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@6.[249,1383,189,1748]" captionTargetPageId="6" captionText="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi="http://doi.org/10.5281/zenodo.4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="7" pageNumber="8">27</figureCitation>
). A band of small, light brown spinules encircling base, with areas projecting forward laterally, spinules short and rounded at apex, each with a lateral hair like process (
<figureCitation box="[292,382,1797,1821]" captionStart="Figs" captionStartId="5.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1746]" captionTargetId="figure-42@5.[249,1383,189,1748]" captionTargetPageId="5" captionText="Figs. 1724. Isoperla karuk: 17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up." figureDoi="http://doi.org/10.5281/zenodo.4758996" httpUri="https://zenodo.org/record/4758996/files/figure.png" pageId="7" pageNumber="8">Figs. 19</figureCitation>
&amp;
<figureCitation box="[416,445,1797,1821]" captionStart="Figs" captionStartId="6.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@6.[249,1383,189,1748]" captionTargetPageId="6" captionText="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi="http://doi.org/10.5281/zenodo.4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="7" pageNumber="8">28</figureCitation>
); anterior surface of aedeagus void of spinules (
<figureCitation box="[392,544,1832,1857]" captionStart="Figs" captionStartId="5.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1746]" captionTargetId="figure-42@5.[249,1383,189,1748]" captionTargetPageId="5" captionText="Figs. 1724. Isoperla karuk: 17. Aedeagus, dorsal, 18. Aedeagus, dorsal, close up, 19. Aedeagus, lateral, 20. Aedeagus, lateral, close up, 21. Aedeagus process, dorsal, close up, 22. Aedeagus process, ventral, 23. Aedeagus process, lateral, 24. Aedeagus process, lateral, close up." figureDoi="http://doi.org/10.5281/zenodo.4758996" httpUri="https://zenodo.org/record/4758996/files/figure.png" pageId="7" pageNumber="8">Figs. 17 &amp; 19</figureCitation>
).
</paragraph>
<paragraph blockId="7.[189,798,198,1928]" lastBlockId="7.[835,1444,198,1929]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[189,282,1868,1892]" pageId="7" pageNumber="8">Female.</emphasis>
Macropterous. Length of forewings 10.0
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="7" pageNumber="8" unit="mm" value="10.5">10.5 mm</quantity>
; length of body 9.010.0 mm. General body color and head and pronotal pigmentation patterns similar to male. Subgenital plate brown, darker than rest of segment, triangular, terminating in broad triangular point, produced ¼ to ½ over length of sternum 9.
</paragraph>
<paragraph blockId="7.[835,1444,198,1929]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[835,887,341,365]" pageId="7" pageNumber="8">Egg.</emphasis>
Shape oval, round in cross section, narrower toward poles (
<figureCitation box="[1001,1080,376,400]" captionStart="Figs" captionStartId="6.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@6.[249,1383,189,1748]" captionTargetPageId="6" captionText="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi="http://doi.org/10.5281/zenodo.4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="7" pageNumber="8">Fig. 31</figureCitation>
). Collar well developed, higher and with broader flanges (
<figureCitation box="[1147,1226,412,436]" captionStart="Figs" captionStartId="6.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@6.[249,1383,189,1748]" captionTargetPageId="6" captionText="Figs. 2532. Isoperla karuk: 25. Aedeagus, spinules, dorsal patch, 26. Dorsal and lateral aedeagus spinule patches, 27. Spinules, at base of aedeagus, 28. Spinules with hairlike addition, encircling aedeagus, 29. Vesicle, male. 30, Male terminalia, dorsal, showing paraprocts, 31. Egg, lateral, 32. Egg collar, close up." figureDoi="http://doi.org/10.5281/zenodo.4758998" httpUri="https://zenodo.org/record/4758998/files/figure.png" pageId="7" pageNumber="8">Fig. 32</figureCitation>
) than shown for
<taxonomicName class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulva">
<emphasis italics="true" pageId="7" pageNumber="8">I. fulva</emphasis>
</taxonomicName>
and
<taxonomicName box="[949,1090,448,471]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis box="[949,1090,448,471]" italics="true" pageId="7" pageNumber="8">I. marmorata</emphasis>
</taxonomicName>
in
<bibRefCitation author="Szczytko, S. W. &amp; K. W. Stewart" box="[1128,1439,447,471]" pageId="7" pageNumber="8" pagination="1 - 120" refId="ref4166" refString="Szczytko, S. W. and K. W. Stewart. 1979. The genus Isoperla (Plecoptera) of Western North America; holomorphology and systematics, and a new stonefly genus Cascadoperla. Memoirs of the Entomological Society of America, 32: 1 - 120." type="journal article" year="1979">Szczytko &amp; Stewart (1979)</bibRefCitation>
. Chorion with thickened ridges, forming pentagon shaped depressions, proteinaceous bodies sometimes present. Micropyles appearing on the bottom half of the egg. Color dark brown and opaque.
</paragraph>
<paragraph blockId="7.[835,1444,198,1929]" box="[835,1042,625,649]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[835,910,625,649]" pageId="7" pageNumber="8">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="etymology">
<paragraph blockId="7.[835,1444,198,1929]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[835,973,661,685]" pageId="7" pageNumber="8">Etymology.</emphasis>
The species name is a noun in apposition and honors the Karuk people of the Klamath River in northern California. The common name Klamath Stripetail is suggested.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="diagnosis">
<paragraph blockId="7.[835,1444,198,1929]" pageId="7" pageNumber="8">
<emphasis bold="true" box="[835,957,803,827]" pageId="7" pageNumber="8">Diagnosis</emphasis>
.
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[977,1136,803,827]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="karuk">
<emphasis box="[977,1136,803,827]" italics="true" pageId="7" pageNumber="8">Isoperla karuk</emphasis>
</taxonomicName>
is placed in the
<taxonomicName class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis italics="true" pageId="7" pageNumber="8">Isoperla marmorata</emphasis>
</taxonomicName>
complex which includes
<taxonomicName authority="(Needham and Claassen)" baseAuthorityName="Needham and Claassen" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis box="[1292,1443,840,863]" italics="true" pageId="7" pageNumber="8">I. marmorata</emphasis>
(Needham and Claassen)
</taxonomicName>
and
<taxonomicName authority="Claassen. It" authorityName="Claassen. It" box="[1210,1443,874,898]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulva">
<emphasis box="[1210,1292,874,898]" italics="true" pageId="7" pageNumber="8">I. fulva</emphasis>
Claassen. It
</taxonomicName>
shares the following characteristics (
<bibRefCitation author="Szczytko, S. W. &amp; K. W. Stewart" pageId="7" pageNumber="8" pagination="1 - 120" refId="ref4166" refString="Szczytko, S. W. and K. W. Stewart. 1979. The genus Isoperla (Plecoptera) of Western North America; holomorphology and systematics, and a new stonefly genus Cascadoperla. Memoirs of the Entomological Society of America, 32: 1 - 120." type="journal article" year="1979">Szczytko &amp; Stewart 1979</bibRefCitation>
): 1. male aedeagus bearing a spinose, club shaped, sclerotized process; 2. broadly rounded, shallow male vesicle; 3. angulate female subgenital plate (shared with
<taxonomicName box="[1086,1173,1052,1076]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulva">
<emphasis box="[1086,1173,1052,1076]" italics="true" pageId="7" pageNumber="8">I. fulva</emphasis>
</taxonomicName>
). The male is easily distinguished from other members of the complex by the unique aedeagus with a bi hemispherical lobe bearing reddish brown spinulae directed anteriorly and two smaller lobes bearing reddish brown spinulae directed posteriorly. The
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[1263,1352,1229,1253]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="karuk">
<emphasis box="[1263,1352,1229,1253]" italics="true" pageId="7" pageNumber="8">I. karuk</emphasis>
</taxonomicName>
female cannot presently be distinguished from the female of
<taxonomicName box="[835,914,1301,1325]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulva">
<emphasis box="[835,914,1301,1325]" italics="true" pageId="7" pageNumber="8">I. fulva</emphasis>
</taxonomicName>
by the shape of the subgenital plate but the egg has a higher and more broadened collar. Nymphs that we believe are
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[1243,1340,1372,1396]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="karuk">
<emphasis box="[1243,1340,1372,1396]" italics="true" pageId="7" pageNumber="8">I. karuk</emphasis>
</taxonomicName>
fit the description for
<taxonomicName authority="(Szczytko &amp; Stewart 1979)" baseAuthorityName="Szczytko &amp; Stewart" baseAuthorityYear="1979" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis box="[1029,1177,1408,1431]" italics="true" pageId="7" pageNumber="8">I. marmorata</emphasis>
(
<bibRefCitation author="Szczytko, S. W. &amp; K. W. Stewart" pageId="7" pageNumber="8" pagination="1 - 120" refId="ref4166" refString="Szczytko, S. W. and K. W. Stewart. 1979. The genus Isoperla (Plecoptera) of Western North America; holomorphology and systematics, and a new stonefly genus Cascadoperla. Memoirs of the Entomological Society of America, 32: 1 - 120." type="journal article" year="1979">Szczytko &amp; Stewart 1979</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
<paragraph blockId="7.[835,1444,198,1929]" lastBlockId="8.[189,798,980,1394]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis bold="true" box="[835,950,1478,1502]" pageId="7" pageNumber="8">Remarks.</emphasis>
This species was collected from medium sized rivers close to sea level. Adults were found from February into May. The peak
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[1233,1313,1549,1573]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="karuk">
<emphasis box="[1233,1313,1549,1573]" italics="true" pageId="7" pageNumber="8">I. karuk</emphasis>
</taxonomicName>
emergence period, on the Mad River, followed the peak emergence of
<taxonomicName authority="Frison" authorityName="Frison" authorityYear="1937" box="[997,1153,1621,1645]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pinta">
<emphasis box="[997,1074,1621,1644]" italics="true" pageId="7" pageNumber="8">I. pinta</emphasis>
Frison
</taxonomicName>
and preceded the peak
<taxonomicName authority="Banks" authorityName="Banks" authorityYear="1920" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mormona">
<emphasis italics="true" pageId="7" pageNumber="8">I. mormona</emphasis>
Banks
</taxonomicName>
emergence period. The riverine habitat made benthic collection of nymphs problematic. Several nymphs were collected, however, and three reared nymphs turned out to be three separate
<taxonomicName box="[1015,1193,1798,1822]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[1015,1099,1798,1822]" italics="true" pageId="7" pageNumber="8">Isoperla</emphasis>
species
</taxonomicName>
(
<emphasis box="[1212,1443,1798,1822]" italics="true" pageId="7" pageNumber="8">
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[1212,1301,1798,1822]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="karuk">I. karuk</taxonomicName>
,
<taxonomicName authorityName="Banks" authorityYear="1920" box="[1315,1443,1799,1822]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mormona">I. mormona</taxonomicName>
</emphasis>
and
<taxonomicName baseAuthorityName="Szczytko &amp; Stewart" baseAuthorityYear="1979" box="[888,1029,1834,1857]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marmorata">
<emphasis box="[888,1029,1834,1857]" italics="true" pageId="7" pageNumber="8">I. marmorata</emphasis>
</taxonomicName>
?). Field caught females and males were maintained together in order to obtain mature ova. Females were alive after 43 days and appeared to feed on, or at least consume the juice of apple slices.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4759000" ID-Zenodo-Dep="4759000" box="[189,860,873,897]" httpUri="https://zenodo.org/record/4759000/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,269,873,897]" targetBox="[249,1383,203,817]" targetPageId="8">
<paragraph blockId="8.[189,860,873,897]" box="[189,860,873,897]" pageId="8" pageNumber="9">
Figure. 33.
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[319,474,873,897]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="laucki">
<emphasis box="[319,474,873,897]" italics="true" pageId="8" pageNumber="9">Isoperla laucki</emphasis>
</taxonomicName>
. Mature nymph, dorsal, habitus.
</paragraph>
</caption>
<paragraph blockId="8.[189,798,980,1394]" pageId="8" pageNumber="9">
Since this species is difficult to separate from
<taxonomicName class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fulva">
<emphasis italics="true" pageId="8" pageNumber="9">I. fulva</emphasis>
</taxonomicName>
in both the male and female adult stages without extruding the aedeagus or studying the eggs, the identity of specimens previously listed as
<taxonomicName box="[721,798,1157,1181]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fulva">
<emphasis box="[721,798,1157,1181]" italics="true" pageId="8" pageNumber="9">I. fulva</emphasis>
</taxonomicName>
from California is questionable.
<bibRefCitation author="Jewett, S. G., Jr." box="[572,724,1193,1217]" pageId="8" pageNumber="9" pagination="125 - 177" refId="ref4081" refString="Jewett, S. G., Jr. 1960. The stoneflies (Plecoptera) of California. Bulletin of the California Insect Survey, 6: 125 - 177." type="journal article" year="1960">Jewett (1960)</bibRefCitation>
in his study of the California stonefly fauna listed two records for
<emphasis box="[322,403,1264,1288]" italics="true" pageId="8" pageNumber="9">
<taxonomicName box="[322,399,1264,1288]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="fulva">I. fulva</taxonomicName>
:
</emphasis>
one in El Dorado County and one in Modoc County. Later,
<bibRefCitation author="Szczytko, S. W. &amp; K. W. Stewart" box="[488,797,1299,1323]" pageId="8" pageNumber="9" pagination="1 - 120" refId="ref4166" refString="Szczytko, S. W. and K. W. Stewart. 1979. The genus Isoperla (Plecoptera) of Western North America; holomorphology and systematics, and a new stonefly genus Cascadoperla. Memoirs of the Entomological Society of America, 32: 1 - 120." type="journal article" year="1979">Szczytko &amp; Stewart (1979)</bibRefCitation>
listed two additional records: Plumas County and Modoc County (Oregon).
</paragraph>
</subSubSection>
</treatment>
</document>