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<document id="7593026D0582793F2E5828BCD0440772" ID-DOI="10.24199/j.mmv.2007.64.8" ID-ISSN="1447-2554" ID-Zenodo-Dep="12211073" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718992553403" checkinUser="felipe" docAuthor="Staples, David A." docDate="2007" docId="D928B2224E2AFFE889C99034FBC4FA4F" docLanguage="en" docName="MemMusVictoria.64.79-94.pdf" docOrigin="Memoirs of Museum Victoria 64" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-64-2007/pages-79-94/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Colossendeis spicula Child 1994" docType="treatment" docVersion="2" lastPageNumber="90" masterDocId="2511CA5A4E21FFE3895C9502FFC6FF95" masterDocTitle="A new species of Colossendeis (Pycnogonida: Colossendeidae) together with records from Australian and New Zealand waters" masterLastPageNumber="94" masterPageNumber="79" pageNumber="90" updateTime="1719004432111" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0EC256746F337AE3B300B507725483BF">A new species of Colossendeis (Pycnogonida: Colossendeidae) together with records from Australian and New Zealand waters</mods:title>
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<mods:namePart id="8A4298A215393C71D8C3D17E7BA2DB36">Staples, David A.</mods:namePart>
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<mods:title id="C3F3CE7831A715E9C3E2BAC4F7FB9419">Memoirs of Museum Victoria</mods:title>
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<mods:date id="663C640EAC02F39683A7AB4B194C41DD">2007</mods:date>
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<mods:number id="1FBD54954DA594A8AFA300F52B0B3FA8">2007-12-31</mods:number>
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<mods:number id="9EE3517E53CE1BCF9E4037AA82F3FAFF">64</mods:number>
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<mods:url id="BC29E6966A9D39936FE8FD90B927C961">https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-64-2007/pages-79-94/</mods:url>
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<mods:identifier id="AF9CC0BC18925EC512174FA420EB488D" type="DOI">10.24199/j.mmv.2007.64.8</mods:identifier>
<mods:identifier id="C3C75A279FB4E7B3C1A182AC4090A651" type="ISSN">1447-2554</mods:identifier>
<mods:identifier id="6D9C3140EF131BA030CBE7B844B8676B" type="Zenodo-Dep">12211073</mods:identifier>
</mods:mods>
<treatment id="D928B2224E2AFFE889C99034FBC4FA4F" LSID="urn:lsid:plazi:treatment:D928B2224E2AFFE889C99034FBC4FA4F" httpUri="http://treatment.plazi.org/id/D928B2224E2AFFE889C99034FBC4FA4F" lastPageNumber="90" pageId="11" pageNumber="90">
<subSubSection id="199B50BF4E2AFFE889C99034FE2DFAD9" box="[149,491,1334,1357]" pageId="11" pageNumber="90" type="nomenclature">
<paragraph id="513E03344E2AFFE889C99034FE2DFAD9" blockId="11.[149,491,1334,1357]" box="[149,491,1334,1357]" pageId="11" pageNumber="90">
<heading id="0A76B4584E2AFFE889C99034FE2DFAD9" box="[149,491,1334,1357]" fontSize="9" level="2" pageId="11" pageNumber="90" reason="5">
<taxonomicName id="968178B74E2AFFE889C99034FE2DFAD9" ID-CoL="5ZMGG" authority="Child, 1994" authorityName="Child" authorityYear="1994" box="[149,491,1334,1357]" class="Pycnogonida" family="Colossendeidae" genus="Colossendeis" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="90" phylum="Arthropoda" rank="species" species="spicula">
<emphasis id="63F5DF264E2AFFE889C99034FEA9FAD9" bold="true" box="[149,367,1334,1356]" italics="true" pageId="11" pageNumber="90">Colossendeis spicula</emphasis>
<bibRefCitation id="35107EC54E2AFFE888299035FE2DFAD9" author="Child, C. A." box="[373,491,1335,1357]" pageId="11" pageNumber="90" pagination="1 - 23" refId="ref9937" refString="Child, C. A. 1994. Deep-Sea Pycnogonida from the temperate west coast of the United States. Smithsonian Contributions to Zoology 556: 1 - 23." type="journal article" year="1994">Child, 1994</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="199B50BF4E2AFFE889C99061FEDEFAED" box="[149,280,1379,1401]" pageId="11" pageNumber="90" type="description">
<paragraph id="513E03344E2AFFE889C99061FEDEFAED" blockId="11.[149,280,1379,1401]" box="[149,280,1379,1401]" pageId="11" pageNumber="90">
<figureCitation id="C9BA1FB14E2AFFE889C99061FEDEFAED" box="[149,280,1379,1401]" captionStart="Figure 4" captionStartId="5.[147,207,1716,1735]" captionTargetBox="[156,1444,246,1659]" captionTargetId="figure-45@5.[134,1450,227,1702]" captionTargetPageId="5" captionText="Figure 4. Hedgpethia dampieri Child, 1975 (NMV J48903): A, Trunk, lateral view; B, ocular tubercle; C, oviger segment 10; holotype (WAM 70–3953) D, oviger spine. Colossendeis spicula (NMV J54504): E, palp; F, leg 2; G, tarsus and propodus; H, oviger." figureDoi="http://doi.org/10.5281/zenodo.12211082" httpUri="https://zenodo.org/record/12211082/files/figure.png" pageId="11" pageNumber="90">Figure 4E–H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="199B50BF4E2AFFE889E6908CFD4EFA34" box="[186,648,1422,1441]" pageId="11" pageNumber="90" type="reference_group">
<paragraph id="513E03344E2AFFE889E6908CFD4EFA34" blockId="11.[186,648,1422,1441]" box="[186,648,1422,1441]" pageId="11" pageNumber="90">
<treatmentCitationGroup id="7191241A4E2AFFE889E6908CFD4EFA34" box="[186,648,1422,1441]" pageId="11" pageNumber="90">
<treatmentCitation id="D02025254E2AFFE889E6908CFDC9FA34" author="Child, C. A." box="[186,527,1422,1441]" page="14" pageId="11" pageNumber="90" year="1994">
<taxonomicName id="968178B74E2AFFE889E6908CFDC9FA34" ID-CoL="5ZMGG" authority="Child, 1994: 14" authorityName="Child" authorityPageNumber="14" authorityYear="1994" box="[186,527,1422,1441]" class="Pycnogonida" family="Colossendeidae" genus="Colossendeis" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="90" phylum="Arthropoda" rank="species" species="spicula">
<emphasis id="63F5DF264E2AFFE889E6908CFEBDFA34" box="[186,379,1422,1441]" italics="true" pageId="11" pageNumber="90">Colossendeis spicula</emphasis>
<bibRefCitation id="35107EC54E2AFFE888DD908CFDC9FA34" author="Child, C. A." box="[385,527,1422,1441]" pageId="11" pageNumber="90" pagination="1 - 23" refId="ref9937" refString="Child, C. A. 1994. Deep-Sea Pycnogonida from the temperate west coast of the United States. Smithsonian Contributions to Zoology 556: 1 - 23." type="journal article" year="1994">Child, 1994: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<figureCitation id="C9BA1FB14E2AFFE88B4B908CFD8DFA34" box="[535,587,1422,1441]" captionStart="Figure 6" captionStartId="7.[147,206,1852,1871]" captionTargetBox="[168,1406,195,1846]" captionTargetId="figure-5@7.[167,1429,194,1990]" captionTargetPageId="7" captionText="Figure 6. Colossendeis mycterismos sp. nov. holotype (NMV J48824): A, B, dorsal and lateral views of trunk; C, oviger; D, oviger segments 7–10; E, palp; F, leg 3; G, tarsus and propodus." figureDoi="http://doi.org/10.5281/zenodo.12211086" httpUri="https://zenodo.org/record/12211086/files/figure.png" pageId="11" pageNumber="90">Fig. 6</figureCitation>
(A–B)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="199B50BF4E2AFFE889C990BAFBC4FA4F" pageId="11" pageNumber="90" type="materials_examined">
<paragraph id="513E03344E2AFFE889C990BAFE59F98F" blockId="11.[149,780,1464,1562]" pageId="11" pageNumber="90">
<emphasis id="63F5DF264E2AFFE889C990BAFE89FA5E" box="[149,335,1464,1483]" italics="true" pageId="11" pageNumber="90">Material examined.</emphasis>
<materialsCitation id="E1E909694E2AFFE8880590BAFF2DF995" collectionCode="RV" collectorName="Beam" elevation="1986" latitude="-38.509666" location="Point Hicks" longLatPrecision="11" longitude="149.35834" pageId="11" pageNumber="90" specimenCount="1" stateProvince="Victoria">
<collectingRegion id="9345CDD64E2AFFE8880590BAFE60FA5E" box="[345,422,1464,1483]" country="Australia" name="Victoria" pageId="11" pageNumber="90">Victoria</collectingRegion>
.
<quantity id="9679AED14E2AFFE888EF90BAFE37FA5E" box="[435,497,1464,1483]" metricMagnitude="4" metricUnit="m" metricValue="8.5" pageId="11" pageNumber="90" unit="km" value="85.0">85 km</quantity>
S of
<location id="545E55EF4E2AFFE88B7390BAFD58FA5E" LSID="urn:lsid:plazi:treatment:D928B2224E2AFFE889C99034FBC4FA4F:545E55EF4E2AFFE88B7390BAFD58FA5E" box="[559,670,1464,1483]" latitude="-38.509666" longLatPrecision="11" longitude="149.35834" name="Point Hicks" pageId="11" pageNumber="90" stateProvince="Victoria">Point Hicks</location>
<geoCoordinate id="34B565F34E2AFFE88BF490BAFCCEFA5E" box="[680,776,1464,1483]" degrees="38" direction="south" minutes="31.41" orientation="latitude" pageId="11" pageNumber="90" precision="9" value="-38.5235">38°31.41'S</geoCoordinate>
,
<geoCoordinate id="34B565F34E2AFFE889C990D0FEC6FA70" box="[149,256,1490,1509]" degrees="149" direction="east" minutes="21.10" orientation="longitude" pageId="11" pageNumber="90" precision="9" value="149.35167">149°21.10'E</geoCoordinate>
,
<quantity id="9679AED14E2AFFE8885190D0FE94FA73" box="[269,338,1490,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.986" pageId="11" pageNumber="90" unit="m" value="1986.0">
<elevation id="DAACE4074E2AFFE8885190D0FE94FA73" box="[269,338,1490,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.986" pageId="11" pageNumber="90" unit="m" value="1986.0">1986 m</elevation>
</quantity>
, to
<geoCoordinate id="34B565F34E2AFFE8882B90D0FE1FFA70" box="[375,473,1490,1509]" degrees="38" direction="south" minutes="30.58" orientation="latitude" pageId="11" pageNumber="90" precision="9" value="-38.509666">38°30.58'S</geoCoordinate>
,
<geoCoordinate id="34B565F34E2AFFE888BA90D0FD92FA70" box="[486,596,1490,1509]" degrees="149" direction="east" minutes="21.50" orientation="longitude" pageId="11" pageNumber="90" precision="9" value="149.35834">149°21.50'E</geoCoordinate>
,
<quantity id="9679AED14E2AFFE88B3D90D0FD61FA73" box="[609,679,1490,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="11" pageNumber="90" unit="m" value="1360.0">
<elevation id="DAACE4074E2AFFE88B3D90D0FD61FA73" box="[609,679,1490,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="11" pageNumber="90" unit="m" value="1360.0">1360 m</elevation>
</quantity>
, 3.5
<collectorName id="FC7466E24E2AFFE88B8A90D1FCCFFA73" box="[726,777,1491,1510]" pageId="11" pageNumber="90">Beam</collectorName>
trawl,
<collectionCode id="37909BF14E2AFFE8899290EFFF2DF995" box="[206,235,1517,1536]" country="Italy" name="Collection of Leptospira Strains" pageId="11" pageNumber="90">RV</collectionCode>
</materialsCitation>
<materialsCitation id="E1E909694E2AFFE889AD90EFFE5DF98F" collectingDate="1988-10-26" collectionCode="NMV" collectorName="Franklin, G. C. B" pageId="11" pageNumber="90" specimenCode="SLOPE 72, J54504" specimenCount="3" stateProvince="Victoria">
<collectorName id="FC7466E24E2AFFE889AD90EFFE4EF995" box="[241,392,1517,1536]" pageId="11" pageNumber="90">
<emphasis id="63F5DF264E2AFFE889AD90EFFE87F995" box="[241,321,1517,1536]" italics="true" pageId="11" pageNumber="90">Franklin</emphasis>
, G.C.B
</collectorName>
. Poore
<emphasis id="63F5DF264E2AFFE8889790EFFE3CF995" box="[459,506,1517,1536]" italics="true" pageId="11" pageNumber="90">et al.</emphasis>
,
<date id="253F25F44E2AFFE88B5890EFFDB5F995" box="[516,627,1517,1536]" pageId="11" pageNumber="90" value="1988-10-26">
<collectingDate id="357BDC1C4E2AFFE88B5890EFFDB5F995" box="[516,627,1517,1536]" pageId="11" pageNumber="90" value="1988-10-26">26 Oct 1988</collectingDate>
</date>
(stn
<specimenCode id="0127AB4F4E2AFFE88BC390EFFCC4F995" box="[671,770,1517,1536]" collectionCode="SLOPE" pageId="11" pageNumber="90">SLOPE 72</specimenCode>
),
<collectionCode id="37909BF14E2AFFE889C99305FF0DF98F" box="[149,203,1543,1562]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="11" pageNumber="90" type="Museum">NMV</collectionCode>
<specimenCode id="0127AB4F4E2AFFE8898C9305FED6F98F" box="[208,272,1543,1562]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="11" pageNumber="90" type="Museum">J54504</specimenCode>
(
<specimenCount id="4787C8BD4E2AFFE8887F9305FE50F98F" box="[291,406,1543,1562]" count="3" pageId="11" pageNumber="90" type="generic">3 specimens</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="513E03344E2AFFE889C99330FB5FFE18" blockId="11.[149,780,1585,1898]" lastBlockId="11.[817,1449,229,398]" pageId="11" pageNumber="90">
<emphasis id="63F5DF264E2AFFE889C99330FEC4F9D2" box="[149,258,1586,1607]" italics="true" pageId="11" pageNumber="90">Diagnosis.</emphasis>
Leg span to about
<quantity id="9679AED14E2AFFE888E59333FDCBF9D2" box="[441,525,1585,1607]" metricMagnitude="-1" metricUnit="m" metricValue="2.85" pageId="11" pageNumber="90" unit="mm" value="285.0">285 mm</quantity>
. Lateral processes shorter than their diameters, fringed distally with a few scattered, short, spines; 2nd and 3rd lateral processes separated by about one-third basal widths, 1st pair not conspicuously raised. Anterior margin of cephalon with pair of small but prominent apophyses over base of palp insertions. Ocular tubercle low, two eyes, unpigmented. Proximal quarter of proboscis narrow, inflated in mid-region, tapering distally to a rounded tip, width marginally less than basal diameter. Palps conspicuously spiny, segment 3, 1.4 times segment 5; segment 7 greater than twice length of segment 6. Abdomen articulated, clavate, little less than 30% of trunk length. Oviger 10-segmented, terminal claw gently curved and articulated with segment 10. Legs about 9 times length of trunk; femur marginally longer than tibia 1; tibia 1, 1.3 times tibia 2. Tarsus 1.8 times propodus, claw 36% of propodus. Spines in rows, straight or curved slightly, longer spines equal to, or longer than width of segment.
</paragraph>
<paragraph id="513E03344E2AFFE88A6D94A6FB58FE42" blockId="11.[817,1447,420,471]" pageId="11" pageNumber="90">
<emphasis id="63F5DF264E2AFFE88A6D94A6FC70FE2C" box="[817,950,420,441]" italics="true" pageId="11" pageNumber="90">Distribution.</emphasis>
West and east Pacific (
<collectingRegion id="9345CDD64E2AFFE88DF194A6FB38FE2C" box="[1197,1278,420,441]" country="New Zealand" name="Tasman" pageId="11" pageNumber="90">Tasman</collectingRegion>
Sea and off the
<collectingRegion id="9345CDD64E2AFFE88A6D94C0FCBAFE42" box="[817,892,450,471]" country="United States of America" name="Oregon" pageId="11" pageNumber="90">Oregon</collectingRegion>
coast), Depth
<quantity id="9679AED14E2AFFE88D5194C3FB5CFE42" box="[1037,1178,449,471]" metricMagnitude="3" metricUnit="m" metricValue="2.096" metricValueMax="2.832" metricValueMin="1.3599999999999999" pageId="11" pageNumber="90" unit="m" value="2096.0" valueMax="2832.0" valueMin="1360.0">1360–2832 m</quantity>
.
</paragraph>
<paragraph id="513E03344E2AFFE88A6D94EDFB1FFD96" blockId="11.[817,1241,494,516]" box="[817,1241,494,516]" pageId="11" pageNumber="90">
<materialsCitation id="E1E909694E2AFFE88A6D94EDFB13FD96" box="[817,1237,494,516]" country="United States of America" location="Off" pageId="11" pageNumber="90" specimenCount="1" stateProvince="Oregon" typeStatus="holotype">
<emphasis id="63F5DF264E2AFFE88A6D94EDFC7DFD96" box="[817,955,494,516]" italics="true" pageId="11" pageNumber="90">
<typeStatus id="8E3ABD964E2AFFE88A6D94EDFCA4FD91" box="[817,866,495,516]" pageId="11" pageNumber="90">Type</typeStatus>
locality.
</emphasis>
<location id="545E55EF4E2AFFE88A9C94ECFC25FD91" LSID="urn:lsid:plazi:treatment:D928B2224E2AFFE889C99034FBC4FA4F:545E55EF4E2AFFE88A9C94ECFC25FD91" box="[960,995,494,516]" country="United States of America" name="Off" pageId="11" pageNumber="90" stateProvince="Oregon">Off</location>
the
<collectingRegion id="9345CDD64E2AFFE88D5394ECFB9CFD96" box="[1039,1114,494,515]" country="United States of America" name="Oregon" pageId="11" pageNumber="90">Oregon</collectingRegion>
coast,
<collectingCountry id="299643A44E2AFFE88DFD94ECFB13FD96" box="[1185,1237,494,515]" name="United States of America" pageId="11" pageNumber="90">USA</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="513E03344E2AFFE88A6D9718FB33FA98" blockId="11.[817,1448,538,1498]" pageId="11" pageNumber="90">
<emphasis id="63F5DF264E2AFFE88A6D9718FC57FDBA" box="[817,913,538,559]" italics="true" pageId="11" pageNumber="90">Remarks.</emphasis>
This collection consists of one large and two smaller specimens about half the size of the larger specimen. Although having fewer spines, they are presumed to be sub-adults of this species. The ocular tubercles of the smaller specimens terminate in an apical cone which was not present in the larger form. The larger specimen is of the same size as the
<typeStatus id="8E3ABD964E2AFFE88D8097AFFAF2FD56" box="[1244,1332,685,707]" pageId="11" pageNumber="90" type="holotype">holotype</typeStatus>
and agrees in almost all respects with Child’s description. Differences are principally confined to spination of oviger segments 4–6 which carry numerous long, sharp, simple spines not present on the
<typeStatus id="8E3ABD964E2AFFE88A6D9620FC4AFCAD" box="[817,908,802,824]" pageId="11" pageNumber="90" type="holotype">holotype</typeStatus>
. Another minor difference is found in the proportions of the femur and tibia 2. Internal ganglia and smooth cuticle in the area where two eyes are usually positioned, suggests that the apparent absence of eyes in the
<typeStatus id="8E3ABD964E2AFFE88DCC9678FB2EFC05" box="[1168,1256,890,912]" pageId="11" pageNumber="90" type="holotype">holotype</typeStatus>
may be a legacy of preservation. Well-defined lateral sensory organs were present on the dorsolateral surface of the ocular tubercle. Palp and oviger gland openings are not evident.
<bibRefCitation id="35107EC54E2AFFE88D9296D0FA8DFC72" author="Child, C. A." box="[1230,1355,978,1000]" pageId="11" pageNumber="90" pagination="1 - 23" refId="ref9937" refString="Child, C. A. 1994. Deep-Sea Pycnogonida from the temperate west coast of the United States. Smithsonian Contributions to Zoology 556: 1 - 23." type="journal article" year="1994">Child (1994</bibRefCitation>
, fig. 4A) clearly figures a 10-segmented palp; however his description overlooks the small 1st segment. His reference to the 2nd palp segment being longest should in fact be the 3rd segment; the 4th segment should be the 5th and so on. Spines on all longer segments are straight and often longer than the corresponding segment. Child describes the three distal segments as being progressively shorter but in this specimen the 10th segment is 1.2 times longer than the 9th, otherwise all segments agree with the
<typeStatus id="8E3ABD964E2AFFE88AC891D8FC2AFB65" box="[916,1004,1242,1264]" pageId="11" pageNumber="90" type="holotype">holotype</typeStatus>
in their proportions. Coxal pellicula and genital pores are not evident in this material.
</paragraph>
<paragraph id="513E03344E2AFFE88A0B9017FBC4FA4F" blockId="11.[817,1448,538,1498]" pageId="11" pageNumber="90">
Numerous gastropod egg capsules were attached to the dorsal surface of the trunk of the larger specimen and ventral surface of one or more legs of all specimens. Most of these capsules were vacated. This species has also much in common with
<taxonomicName id="968178B74E2AFFE88A389088FB85FA0A" authority="Child, 1995" authorityName="Child" authorityYear="1995" box="[868,1091,1417,1440]" class="Pycnogonida" family="Colossendeidae" genus="Colossendeis" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="90" phylum="Arthropoda" rank="species" species="ensifer">
<emphasis id="63F5DF264E2AFFE88A389088FC01FA0A" box="[868,967,1418,1439]" italics="true" pageId="11" pageNumber="90">C. ensifer</emphasis>
<bibRefCitation id="35107EC54E2AFFE88A909088FB85FA0A" author="Child, C. A." box="[972,1091,1417,1440]" pageId="11" pageNumber="90" pagination="69 - 111" refId="ref9967" refString="Child, C. A. 1995. Antarctic and Subantarctic Pycnogonida IV. The families Colossendeidae and Rhynchothoracidae. Pp. 69 - 111 in: Cairns, S. (ed.), Biology of the Antarctic Seas XXIV Antarctic Research Series, 69. American Geophysical Union: Washington D. C." type="book chapter" year="1995">Child, 1995</bibRefCitation>
</taxonomicName>
but that species is distinguished by the presence of strong, hooked spines on the palps and on the longer leg segments.
</paragraph>
</subSubSection>
</treatment>
</document>
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