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<document id="A88F12335CBCC8A5D5818F6DB181D088" ID-DOI="10.5070/P940561331" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913477" ID-ZooBank="1756B24A-813B-423F-896F-91B21FF58A79" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712169731283" checkinUser="felipe" docAuthor="Hickman, Carole S." docDate="2023" docId="C23987DDFFDF2916FC24FA9BEF3CBB58" docLanguage="en" docName="PaleoBios.40.5.1-51.pdf" docOrigin="PaleoBios 40 (5)" docSource="http://dx.doi.org/10.5070/p940561331" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.1:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="1" docTitle="Acesta oregonensis" docType="treatment" docVersion="2" lastPageNumber="41" masterDocId="3E00FFA5FFF8293CFF9FFFA4EE04BD6F" masterDocTitle="Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part II: The pteriomorphs" masterLastPageNumber="51" masterPageNumber="1" pageNumber="38" updateTime="1712170163329" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="19B6994D34C4550F31C0D3DCCF67ED82">Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part II: The pteriomorphs</mods:title>
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<treatment id="C23987DDFFDF2916FC24FA9BEF3CBB58" LSID="urn:lsid:plazi:treatment:C23987DDFFDF2916FC24FA9BEF3CBB58" httpUri="http://treatment.plazi.org/id/C23987DDFFDF2916FC24FA9BEF3CBB58" lastPageId="42" lastPageNumber="41" pageId="39" pageNumber="38">
<subSubSection id="028A6540FFDF291BFC24FA9BEB70B835" box="[955,1396,1343,1371]" pageId="39" pageNumber="38" type="nomenclature">
<paragraph id="4A2F36CBFFDF291BFC24FA9BEB70B835" blockId="39.[955,1396,1343,1406]" box="[955,1396,1343,1371]" pageId="39" pageNumber="38">
<heading id="116781A7FFDF291BFC24FA9BEB70B835" allCaps="true" box="[955,1396,1343,1371]" centered="true" fontSize="10" level="2" pageId="39" pageNumber="38" reason="2">
<taxonomicName id="8D904D48FFDF291BFC24FA9BEB70B835" authority="(CLARK, 1925)" baseAuthorityName="CLARK" baseAuthorityYear="1925" box="[955,1396,1343,1371]" class="Bivalvia" family="Limidae" genus="Acesta" kingdom="Animalia" order="Limida" pageId="39" pageNumber="41" phylum="Mollusca" rank="species" species="oregonensis">
<emphasis id="78E4EAD9FFDF291BFC24FA9BEAC9B834" box="[955,1229,1343,1371]" italics="true" pageId="39" pageNumber="38">ACESTA OREGONENSIS</emphasis>
(
<bibRefCitation id="2E014B3AFFDF291BFB43FAE6EB6EB835" author="Clark, B. L." box="[1244,1386,1346,1370]" pageId="39" pageNumber="38" pagination="69 - 136" refId="ref28111" refString="Clark, B. L. 1925. Pelecypoda from the marine Oligocene of western North America. University of California Bulletin of the Department of Geological Sciences 15 (4): 69 - 136, pls. 8 - 22." type="journal article" year="1925">CLARK, 1925</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDF291BFC62FAC2EB36B812" box="[1021,1330,1382,1406]" pageId="39" pageNumber="38" type="description">
<paragraph id="4A2F36CBFFDF291BFC62FAC2EB36B812" blockId="39.[955,1396,1343,1406]" box="[1021,1330,1382,1406]" pageId="39" pageNumber="38">
<figureCitation id="D2AB2A4EFFDF291BFC62FAC2EA8CB811" box="[1021,1160,1382,1406]" captionStart="Figure 25" captionStartId="40.[112,187,1537,1561]" captionTargetBox="[125,1498,192,1510]" captionTargetId="figure-144@40.[125,1498,192,1510]" captionTargetPageId="40" captionText="Figure 25. Holotype and paratype of Acesta oregonensis (Clark, 1925). A. Clark’s trimmed and retouched original figure of Keasey holotype, right valve, UCMP 30303. B. Unmodified color image of holotype in matrix. C. Clark’s original figure of the paratype, left valve, UCMP 30312. D. Unmodified color image of paratype in matrix. Scale bars=1 cm." figureDoi="http://doi.org/10.5281/zenodo.10913539" httpUri="https://zenodo.org/record/10913539/files/figure.png" pageId="39" pageNumber="38">FIGS. 25A–D</figureCitation>
;
<figureCitation id="D2AB2A4EFFDF291BFB0DFAC2EADBB812" box="[1170,1247,1382,1406]" captionStart="Figure 26" captionStartId="41.[110,185,1020,1044]" captionTargetBox="[140,1518,205,975]" captionTargetId="figure-403@41.[115,1522,192,998]" captionTargetPageId="41" captionText="Figure 26. Acesta oregonensis (Clark, 1925), Keasey hypotype UCMP 110740. A. Original shell, left valve exterior, fitted together from two siltstone blocks, matrix digitally removed. Scale bar=5 cm. B. Detail of posterior ear." figureDoi="http://doi.org/10.5281/zenodo.10913545" httpUri="https://zenodo.org/record/10913545/files/figure.png" pageId="39" pageNumber="38">26A, B</figureCitation>
;
<figureCitation id="D2AB2A4EFFDF291BFB76FAC2EB36B812" box="[1257,1330,1382,1406]" captionStart="Figure 27" captionStartId="42.[112,187,1039,1063]" captionTargetBox="[115,1514,192,1009]" captionTargetId="figure-339@42.[115,1514,192,1009]" captionTargetPageId="42" captionText="Figure 27. Acesta oregonensis (Clark, 1925). Hypotype, UCMP 110741. A. Right valve in matrix with adhering interior (aragonitic) and exterior (calcitic) shell layers. Scale bar=5 cm. B. Fragments of exfoliated aragonitic layers from A." figureDoi="http://doi.org/10.5281/zenodo.10913547" httpUri="https://zenodo.org/record/10913547/files/figure.png" pageId="39" pageNumber="38">27A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDF291BFCDFFA3CEDA3BB76" pageId="39" pageNumber="38" type="reference_group">
<paragraph id="4A2F36CBFFDF291BFCDFFA3CEDF8B8BE" blockId="39.[832,1514,1432,1937]" pageId="39" pageNumber="38">
<taxonomicName id="8D904D48FFDF291BFCDFFA3CEB76B8DB" authority="Clark (1925)" authorityName="Clark, Vokes" authorityYear="1925" box="[832,1394,1432,1462]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="39" pageNumber="38" phylum="Mollusca" rank="species" species="oregonensis" subGenus="Plagiostoma">
<emphasis id="78E4EAD9FFDF291BFCDFFA3CEACEB8D9" box="[832,1226,1432,1462]" italics="true" pageId="39" pageNumber="38">Lima (Plagiostoma) oregonensis</emphasis>
<bibRefCitation id="2E014B3AFFDF291BFB4EFA3DEB76B8DB" author="Clark, B. L." box="[1233,1394,1433,1460]" pageId="39" pageNumber="38" pagination="69 - 136" refId="ref28111" refString="Clark, B. L. 1925. Pelecypoda from the marine Oligocene of western North America. University of California Bulletin of the Department of Geological Sciences 15 (4): 69 - 136, pls. 8 - 22." type="journal article" year="1925">Clark (1925)</bibRefCitation>
</taxonomicName>
. p. 84; pl. 24, figs. 3, 4.
</paragraph>
<paragraph id="4A2F36CBFFDF291BFCDFFA44EDA3BB76" blockId="39.[832,1514,1432,1937]" pageId="39" pageNumber="38">
<taxonomicName id="8D904D48FFDF291BFCDFFA44EBE1B893" authority="Clark, Schenck (1936)" authorityName="Clark, Schenck" authorityYear="1936" box="[832,1509,1504,1534]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="39" pageNumber="38" phylum="Mollusca" rank="species" species="oregonensis" subGenus="Plagiostoma">
<emphasis id="78E4EAD9FFDF291BFCDFFA44EACEB891" box="[832,1226,1504,1534]" italics="true" pageId="39" pageNumber="38">Lima (Plagiostoma) oregonensis</emphasis>
Clark, Schenck (1936)
</taxonomicName>
. p. 62.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDF291BFCDFF98CEDD4BBC6" pageId="39" pageNumber="38" type="description">
<paragraph id="4A2F36CBFFDF291BFCDFF98CEA0FBB0E" blockId="39.[832,1514,1432,1937]" pageId="39" pageNumber="38">
<taxonomicName id="8D904D48FFDF291BFCDFF98CEDB8BB0E" authority="Clark, Woodring (1938)" authorityName="Clark, Woodring" authorityYear="1938" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="39" pageNumber="38" phylum="Mollusca" rank="species" species="oregonensis" subGenus="Plagiostoma">
<emphasis id="78E4EAD9FFDF291BFCDFF98CEACEBB29" box="[832,1226,1576,1606]" italics="true" pageId="39" pageNumber="38">Lima (Plagiostoma) oregonensis</emphasis>
Clark, Woodring (1938)
</taxonomicName>
. p. 48.
</paragraph>
<paragraph id="4A2F36CBFFDF291BFCDFF9D4EDD4BBC6" blockId="39.[832,1514,1432,1937]" pageId="39" pageNumber="38">
<taxonomicName id="8D904D48FFDF291BFCDFF9D4EB23BBE3" authority="Clark, Weaver (1943)" authorityName="Clark, Weaver" authorityYear="1943" box="[832,1319,1648,1678]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="39" pageNumber="38" phylum="Mollusca" rank="species" species="oregonensis">
<emphasis id="78E4EAD9FFDF291BFCDFF9D4EA11BBE1" box="[832,1045,1648,1678]" italics="true" pageId="39" pageNumber="38">Lima oregonensis</emphasis>
Clark, Weaver (1943)
</taxonomicName>
. p. 99; pl. 22, figs. 2, 6.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDF291BFCDFF91CEDEBBA72" pageId="39" pageNumber="38" type="reference_group">
<paragraph id="4A2F36CBFFDF291BFCDFF91CEB8BBBBB" blockId="39.[832,1514,1432,1937]" box="[832,1423,1720,1750]" pageId="39" pageNumber="38">
<taxonomicName id="8D904D48FFDF291BFCDFF91CEB34BBBB" authority="Clark, Durham (1944)" authorityName="Clark, Durham" authorityYear="1944" box="[832,1328,1720,1750]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="39" pageNumber="38" phylum="Mollusca" rank="species" species="oregonensis">
<emphasis id="78E4EAD9FFDF291BFCDFF91CEA11BBB9" box="[832,1045,1720,1750]" italics="true" pageId="39" pageNumber="38">Lima oregonensis</emphasis>
Clark,
<bibRefCitation id="2E014B3AFFDF291BFBF4F91DEB34BBBB" author="Durham, J. W." box="[1131,1328,1721,1748]" pageId="39" pageNumber="38" pagination="101 - 212" refId="ref29671" refString="Durham, J. W. 1944. Megafaunal zones of the Oligocene of northwestern Washington. University of California Publications, Department of Geological Sciences, Bulletin 27 (5): 101 - 212. pls. 13 - 18, figs. 1 - 7." type="journal article" year="1944">Durham (1944)</bibRefCitation>
</taxonomicName>
. p. 139.
</paragraph>
<paragraph id="4A2F36CBFFDF291BFCDFF947EDEBBA72" blockId="39.[832,1514,1432,1937]" pageId="39" pageNumber="38">
<taxonomicName id="8D904D48FFDF291BFCDFF947EAB0BB91" authority="Clark, Vokes" authorityName="Clark, Vokes" authorityYear="1925" box="[832,1204,1763,1793]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="39" pageNumber="38" phylum="Mollusca" rank="species" species="oregonensis">
<emphasis id="78E4EAD9FFDF291BFCDFF947EA11BA6E" box="[832,1045,1763,1793]" italics="true" pageId="39" pageNumber="38">Lima oregonensis</emphasis>
Clark, Vokes
</taxonomicName>
in Warren et al. (1945) (checklist).
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDF291BFCDFF88FEAAEBA0B" pageId="39" pageNumber="38" type="description">
<paragraph id="4A2F36CBFFDF291BFCDFF88FEAAEBA0B" blockId="39.[832,1514,1432,1937]" pageId="39" pageNumber="38">
<taxonomicName id="8D904D48FFDF291BFCDFF88FEAB0BA29" authority="Clark, Vokes" authorityName="Clark, Vokes" authorityYear="1925" box="[832,1204,1835,1865]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="39" pageNumber="38" phylum="Mollusca" rank="species" species="oregonensis">
<emphasis id="78E4EAD9FFDF291BFCDFF88FEA11BA26" box="[832,1045,1835,1865]" italics="true" pageId="39" pageNumber="38">Lima oregonensis</emphasis>
Clark, Vokes
</taxonomicName>
in Warren and Norbisrath (1946). p. 227.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFDF2914FCDFF8D7EEECBBF5" lastPageId="40" lastPageNumber="39" pageId="39" pageNumber="38" type="reference_group">
<paragraph id="4A2F36CBFFDF291BFCDFF8D7EBEEBAE0" blockId="39.[832,1514,1432,1937]" box="[832,1514,1907,1937]" pageId="39" pageNumber="38">
<taxonomicName id="8D904D48FFDF291BFCDFF8D7EBE1BAE0" authority="Clark, Hertlein (1952)" authorityName="Clark, Hertlein" authorityYear="1952" box="[832,1509,1907,1937]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="39" pageNumber="38" phylum="Mollusca" rank="species" species="oregonensis" subGenus="Plagiostoma">
<emphasis id="78E4EAD9FFDF291BFCDFF8D7EACEBAFE" box="[832,1226,1907,1937]" italics="true" pageId="39" pageNumber="38">Lima (Plagiostoma) oregonensis</emphasis>
Clark,
<bibRefCitation id="2E014B3AFFDF291BFA80F8D0EBE1BAE0" author="Hertlein, L. G." box="[1311,1509,1908,1935]" pageId="39" pageNumber="38" pagination="377 - 381" refId="ref30830" refString="Hertlein, L. G. 1952. Description of a new pelecypod of the genus Lima from deep water off Central California. California Academy of Sciences Proceedings 27 (12): 377 - 381." type="journal article" year="1952">Hertlein (1952)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="1EEF6643FFD02914FFEFF9A5EA17BB21" ID-DOI="http://doi.org/10.5281/zenodo.10913539" ID-Zenodo-Dep="10913539" httpUri="https://zenodo.org/record/10913539/files/figure.png" pageId="40" pageNumber="39" startId="40.[112,187,1537,1561]" targetBox="[125,1498,192,1510]" targetPageId="40" targetType="figure">
<paragraph id="4A2F36CBFFD02914FFEFF9A5EA17BB21" blockId="40.[112,1501,1537,1615]" pageId="40" pageNumber="39">
<emphasis id="78E4EAD9FFD02914FFEFF9A5EEDABB76" bold="true" box="[112,222,1537,1561]" pageId="40" pageNumber="39">Figure 25</emphasis>
. Holotype and paratype of
<taxonomicName id="8D904D48FFD02914FD9FF9A7ED59BB75" authority="(Clark, 1925)" baseAuthorityName="CLARK" baseAuthorityYear="1925" box="[512,861,1537,1563]" class="Bivalvia" family="Limidae" genus="Acesta" kingdom="Animalia" order="Limida" pageId="40" pageNumber="39" phylum="Mollusca" rank="species" species="oregonensis">
<emphasis id="78E4EAD9FFD02914FD9FF9A7ECCCBB74" box="[512,712,1539,1563]" italics="true" pageId="40" pageNumber="39">Acesta oregonensis</emphasis>
(
<bibRefCitation id="2E014B3AFFD02914FD48F9A5ED57BB76" author="Clark, B. L." box="[727,851,1537,1561]" pageId="40" pageNumber="39" pagination="69 - 136" refId="ref28111" refString="Clark, B. L. 1925. Pelecypoda from the marine Oligocene of western North America. University of California Bulletin of the Department of Geological Sciences 15 (4): 69 - 136, pls. 8 - 22." type="journal article" year="1925">Clark, 1925</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="78E4EAD9FFD02914FCF7F9A5ED7DBB76" bold="true" box="[872,889,1537,1561]" pageId="40" pageNumber="39">A</emphasis>
. Clark’s trimmed and retouched original figure of Keasey holotype, right valve, UCMP 30303.
<emphasis id="78E4EAD9FFD02914FE70F9B8EFFBBB5B" bold="true" box="[495,511,1564,1588]" pageId="40" pageNumber="39">B</emphasis>
. Unmodified color image of holotype in matrix.
<emphasis id="78E4EAD9FFD02914FB9FF9B8EA0BBB5B" bold="true" box="[1024,1039,1564,1588]" pageId="40" pageNumber="39">C</emphasis>
. Clark’s original figure of the paratype, left valve, UCMP 30312.
<emphasis id="78E4EAD9FFD02914FED4F993EF59BB20" bold="true" box="[331,349,1591,1615]" pageId="40" pageNumber="39">D</emphasis>
. Unmodified color image of paratype in matrix. Scale bars=1 cm.
</paragraph>
</caption>
<paragraph id="4A2F36CBFFD02914FF0BF925EEECBBF5" blockId="40.[112,800,1664,1945]" box="[148,232,1664,1691]" pageId="40" pageNumber="39">p. 379.</paragraph>
</subSubSection>
<subSubSection id="028A6540FFD02914FFEFF90DEEF4BA44" pageId="40" pageNumber="39" type="description">
<paragraph id="4A2F36CBFFD02914FFEFF90DEFECBB8D" blockId="40.[112,800,1664,1945]" pageId="40" pageNumber="39">
<taxonomicName id="8D904D48FFD02914FFEFF90DEC16BBA9" authority="(Clark)" baseAuthorityName="Clark" box="[112,530,1705,1735]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="40" pageNumber="39" phylum="Mollusca" rank="species" species="oregonensis" subGenus="Acesta">
<emphasis id="78E4EAD9FFD02914FFEFF90DEFB5BBA8" box="[112,433,1705,1735]" italics="true" pageId="40" pageNumber="39">Lima (Acesta) oregonensis</emphasis>
(Clark)
</taxonomicName>
,
<collectionCode id="2C81AE0EFFD02914FD82F90FEC35BBAA" box="[541,561,1707,1733]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="40" pageNumber="39">R</collectionCode>
.C. Moore and Vokes (1953). pp. 115, 118.
</paragraph>
<paragraph id="4A2F36CBFFD02914FFEFF955ECF3BA62" blockId="40.[112,800,1664,1945]" box="[112,759,1777,1807]" pageId="40" pageNumber="39">
<taxonomicName id="8D904D48FFD02914FFEFF955EC16BA61" authority="(Clark)" baseAuthorityName="Clark" box="[112,530,1777,1807]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="40" pageNumber="39" phylum="Mollusca" rank="species" species="oregonensis" subGenus="Acesta">
<emphasis id="78E4EAD9FFD02914FFEFF955EFB5BA60" box="[112,433,1777,1807]" italics="true" pageId="40" pageNumber="39">Lima (Acesta) oregonensis</emphasis>
(Clark)
</taxonomicName>
, Vokes (1963), pp.
</paragraph>
<paragraph id="4A2F36CBFFD02914FF04F8B4EEF4BA44" blockId="40.[112,800,1664,1945]" box="[155,240,1808,1835]" pageId="40" pageNumber="39">85–86.</paragraph>
</subSubSection>
<subSubSection id="028A6540FFD02915FF0FF89EEA7DBA78" lastPageId="41" lastPageNumber="40" pageId="40" pageNumber="39" type="discussion">
<paragraph id="4A2F36CBFFD02915FF0FF89EEC0EB842" blockId="40.[112,800,1664,1945]" lastBlockId="41.[112,801,1120,1959]" lastPageId="41" lastPageNumber="40" pageId="40" pageNumber="39">
<emphasis id="78E4EAD9FFD02914FF0FF89EEF21BA3B" bold="true" box="[144,293,1850,1876]" pageId="40" pageNumber="39">Discussion</emphasis>
—In the original description, Clark assigned this species to
<emphasis id="78E4EAD9FFD02914FE1CF8F9EC18BA16" box="[387,540,1885,1913]" italics="true" pageId="40" pageNumber="39">Plagiostoma</emphasis>
J. Sowerby (1814), a genus based on an Early Jurassic species with a thicker shell, a broad cardinal area, and a triangular resilifer located beneath the beaks. The diagnostic features of
<emphasis id="78E4EAD9FFD02914FCDFF969EDD3BB86" box="[832,983,1741,1769]" italics="true" pageId="40" pageNumber="39">Plagiostoma</emphasis>
are in marked contrast to the thin shell, narrow cardinal area and posteriorly oblique ligament pit of the Paleogene species, which here is assigned to
<taxonomicName id="8D904D48FFD02914FA04F8B6EBEEBA41" box="[1435,1514,1810,1838]" class="Bivalvia" family="Limidae" genus="Acesta" kingdom="Animalia" order="Limida" pageId="40" pageNumber="39" phylum="Mollusca" rank="genus">
<emphasis id="78E4EAD9FFD02914FA04F8B6EBEEBA41" box="[1435,1514,1810,1838]" italics="true" pageId="40" pageNumber="39">Acesta</emphasis>
</taxonomicName>
. Clark’s original illustrations are misleading photographs in which the
<typeStatus id="952B8869FFD02914FC43F8FEEA4FBA1B" box="[988,1099,1882,1908]" pageId="40" pageNumber="39" type="holotype">holotype</typeStatus>
(
<figureCitation id="D2AB2A4EFFD02914FBC7F8FEEAC5BA1A" box="[1112,1217,1882,1909]" captionStart="Figure 25" captionStartId="40.[112,187,1537,1561]" captionTargetBox="[125,1498,192,1510]" captionTargetId="figure-144@40.[125,1498,192,1510]" captionTargetPageId="40" captionText="Figure 25. Holotype and paratype of Acesta oregonensis (Clark, 1925). A. Clark’s trimmed and retouched original figure of Keasey holotype, right valve, UCMP 30303. B. Unmodified color image of holotype in matrix. C. Clark’s original figure of the paratype, left valve, UCMP 30312. D. Unmodified color image of paratype in matrix. Scale bars=1 cm." figureDoi="http://doi.org/10.5281/zenodo.10913539" httpUri="https://zenodo.org/record/10913539/files/figure.png" pageId="40" pageNumber="39">Fig. 25A</figureCitation>
) and
<typeStatus id="952B8869FFD02914FA9DF8FFEB76BA1A" box="[1282,1394,1883,1909]" pageId="40" pageNumber="39" type="paratype">paratype</typeStatus>
(
<figureCitation id="D2AB2A4EFFD02914FA1FF8FEEBECBA1A" box="[1408,1512,1882,1909]" captionStart="Figure 25" captionStartId="40.[112,187,1537,1561]" captionTargetBox="[125,1498,192,1510]" captionTargetId="figure-144@40.[125,1498,192,1510]" captionTargetPageId="40" captionText="Figure 25. Holotype and paratype of Acesta oregonensis (Clark, 1925). A. Clark’s trimmed and retouched original figure of Keasey holotype, right valve, UCMP 30303. B. Unmodified color image of holotype in matrix. C. Clark’s original figure of the paratype, left valve, UCMP 30312. D. Unmodified color image of paratype in matrix. Scale bars=1 cm." figureDoi="http://doi.org/10.5281/zenodo.10913539" httpUri="https://zenodo.org/record/10913539/files/figure.png" pageId="40" pageNumber="39">Fig. 25C</figureCitation>
) were trimmed to eliminate the matrix in which they are imbedded, and fail to show the distribution of adhering, poorly-preserved shell fragments. New photographs of the types (
<figureCitation id="D2AB2A4EFFD12915FF68FB0CEF90B9AC" box="[247,404,1192,1219]" captionStart="Figure 25" captionStartId="40.[112,187,1537,1561]" captionTargetBox="[125,1498,192,1510]" captionTargetId="figure-144@40.[125,1498,192,1510]" captionTargetPageId="40" captionText="Figure 25. Holotype and paratype of Acesta oregonensis (Clark, 1925). A. Clark’s trimmed and retouched original figure of Keasey holotype, right valve, UCMP 30303. B. Unmodified color image of holotype in matrix. C. Clark’s original figure of the paratype, left valve, UCMP 30312. D. Unmodified color image of paratype in matrix. Scale bars=1 cm." figureDoi="http://doi.org/10.5281/zenodo.10913539" httpUri="https://zenodo.org/record/10913539/files/figure.png" pageId="41" pageNumber="40">Figs. 25B, D</figureCitation>
) are provided here. Clark also reversed the orientation of the ears in his description, describing the larger posterior ear as anterior and the smaller anterior ear as posterior.
</paragraph>
<caption id="1EEF6643FFD12915FFF1FC58EA5EB941" ID-DOI="http://doi.org/10.5281/zenodo.10913545" ID-Zenodo-Dep="10913545" httpUri="https://zenodo.org/record/10913545/files/figure.png" pageId="41" pageNumber="40" startId="41.[110,185,1020,1044]" targetBox="[140,1518,205,975]" targetPageId="41" targetType="figure">
<paragraph id="4A2F36CBFFD12915FFF1FC58EA5EB941" blockId="41.[110,1481,1020,1071]" pageId="41" pageNumber="40">
<emphasis id="78E4EAD9FFD12915FFF1FC58EED9B97B" bold="true" box="[110,221,1020,1044]" pageId="41" pageNumber="40">Figure 26</emphasis>
.
<taxonomicName id="8D904D48FFD12915FF78FC59EC40B97B" authority="(Clark, 1925)" baseAuthorityName="CLARK" baseAuthorityYear="1925" box="[231,580,1020,1045]" class="Bivalvia" family="Limidae" genus="Acesta" kingdom="Animalia" order="Limida" pageId="41" pageNumber="40" phylum="Mollusca" rank="species" species="oregonensis">
<emphasis id="78E4EAD9FFD12915FF78FC59EFAAB97A" box="[231,430,1021,1045]" italics="true" pageId="41" pageNumber="40">Acesta oregonensis</emphasis>
(
<bibRefCitation id="2E014B3AFFD12915FE21FC58EC3EB97B" author="Clark, B. L." box="[446,570,1020,1044]" pageId="41" pageNumber="40" pagination="69 - 136" refId="ref28111" refString="Clark, B. L. 1925. Pelecypoda from the marine Oligocene of western North America. University of California Bulletin of the Department of Geological Sciences 15 (4): 69 - 136, pls. 8 - 22." type="journal article" year="1925">Clark, 1925</bibRefCitation>
)
</taxonomicName>
, Keasey hypotype UCMP 110740.
<emphasis id="78E4EAD9FFD12915FC2CFC58EDC0B97B" bold="true" box="[947,964,1020,1044]" pageId="41" pageNumber="40">A</emphasis>
. Original shell, left valve exterior, fitted together from two siltstone blocks, matrix digitally removed. Scale bar=5 cm.
<emphasis id="78E4EAD9FFD12915FCD1FBB3ED5AB940" bold="true" box="[846,862,1047,1071]" pageId="41" pageNumber="40">B</emphasis>
. Detail of posterior ear.
</paragraph>
</caption>
<paragraph id="4A2F36CBFFD12915FF0FFA92EC00BBBC" blockId="41.[112,801,1120,1959]" pageId="41" pageNumber="40">
Subsequent specimens collected by Hans Norbisrath,
<collectionCode id="2C81AE0EFFD12915FFEFFAFEEE81B81B" box="[112,133,1370,1396]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="41" pageNumber="40">R</collectionCode>
.M. Grivetti and Harold Vokes as well as specimens collected in preparation of this monograph are all from the upper member of the Keasey Formation, where original shell material is typically well preserved and the dominant (occurring at most localities) element in assemblages along with basal root tufts of hexactinellid sponges, less frequent specimens of
<taxonomicName id="8D904D48FFD12915FDCBF98FED15BB26" baseAuthorityName="Dickerson" baseAuthorityYear="1917" box="[596,785,1579,1609]" class="Bivalvia" family="Parallelodontidae" genus="Porterius" kingdom="Animalia" order="Arcida" pageId="41" pageNumber="40" phylum="Mollusca" rank="species" species="gabb">
<emphasis id="78E4EAD9FFD12915FDCBF98FED15BB26" box="[596,785,1579,1609]" italics="true" pageId="41" pageNumber="40">Porterius gabb</emphasis>
</taxonomicName>
i, and poorly-preserved terebratulid brachiopods. Failure to collect thin, delicate shell fragments undoubtedly accounts for under-representation of the species in both museum and private collections.
</paragraph>
<paragraph id="4A2F36CBFFD12915FF0FF978EB4FB867" blockId="41.[112,801,1120,1959]" lastBlockId="41.[832,1520,1120,1957]" pageId="41" pageNumber="40">
Careful examination of all available material is most informative in a composite characterization of the exterior ornamentation. Shells are smooth or with faint radial lines on the central part of the shell but pronounced on the anterior and posterior dorsal margins where numerous, finely incised radial lines are irregular in width and spacing (
<figureCitation id="D2AB2A4EFFD12915FC73FBC4EA59B914" box="[1004,1117,1120,1147]" captionStart="Figure 26" captionStartId="41.[110,185,1020,1044]" captionTargetBox="[140,1518,205,975]" captionTargetId="figure-403@41.[115,1522,192,998]" captionTargetPageId="41" captionText="Figure 26. Acesta oregonensis (Clark, 1925), Keasey hypotype UCMP 110740. A. Original shell, left valve exterior, fitted together from two siltstone blocks, matrix digitally removed. Scale bar=5 cm. B. Detail of posterior ear." figureDoi="http://doi.org/10.5281/zenodo.10913545" httpUri="https://zenodo.org/record/10913545/files/figure.png" pageId="41" pageNumber="40">Fig. 26A</figureCitation>
). A peculiar disjunct pattern of interrupted ribbing develops at intersections with commarginally bunched growth lines (
<figureCitation id="D2AB2A4EFFD12915FB75FB03EB52B9AD" box="[1258,1366,1191,1218]" captionStart="Figure 26" captionStartId="41.[110,185,1020,1044]" captionTargetBox="[140,1518,205,975]" captionTargetId="figure-403@41.[115,1522,192,998]" captionTargetPageId="41" captionText="Figure 26. Acesta oregonensis (Clark, 1925), Keasey hypotype UCMP 110740. A. Original shell, left valve exterior, fitted together from two siltstone blocks, matrix digitally removed. Scale bar=5 cm. B. Detail of posterior ear." figureDoi="http://doi.org/10.5281/zenodo.10913545" httpUri="https://zenodo.org/record/10913545/files/figure.png" pageId="41" pageNumber="40">Fig. 26B</figureCitation>
). The glossy, pale brown exterior calcitic layer is underlain by a white chalky inner aragonitic layer (
<figureCitation id="D2AB2A4EFFD12915FB2BFB49EB39B867" box="[1204,1341,1261,1288]" captionStart="Figure 27" captionStartId="42.[112,187,1039,1063]" captionTargetBox="[115,1514,192,1009]" captionTargetId="figure-339@42.[115,1514,192,1009]" captionTargetPageId="42" captionText="Figure 27. Acesta oregonensis (Clark, 1925). Hypotype, UCMP 110741. A. Right valve in matrix with adhering interior (aragonitic) and exterior (calcitic) shell layers. Scale bar=5 cm. B. Fragments of exfoliated aragonitic layers from A." figureDoi="http://doi.org/10.5281/zenodo.10913547" httpUri="https://zenodo.org/record/10913547/files/figure.png" pageId="41" pageNumber="40">Fig. 27A, B</figureCitation>
).
</paragraph>
<paragraph id="4A2F36CBFFD12915FCFFFAB4EA7DBA78" blockId="41.[832,1520,1120,1957]" pageId="41" pageNumber="40">
The Keasey species is most similar in shell shape and ornamentation to
<taxonomicName id="8D904D48FFD12915FBB9FA96EB7EB821" authority="(Dall, 1900)" baseAuthorityName="Dall" baseAuthorityYear="1900" box="[1062,1402,1330,1358]" class="Bivalvia" family="Limidae" genus="Acesta" kingdom="Animalia" order="Limida" pageId="41" pageNumber="40" phylum="Mollusca" rank="species" species="hamlini">
<emphasis id="78E4EAD9FFD12915FBB9FA96EADFB821" box="[1062,1243,1330,1358]" italics="true" pageId="41" pageNumber="40">Acesta hamlini</emphasis>
(
<bibRefCitation id="2E014B3AFFD12915FB71FA90EB6BB821" author="Dall, W. H." box="[1262,1391,1332,1358]" pageId="41" pageNumber="40" pagination="15 - 16" refId="ref28772" refString="Dall, W. H. 1900. A new species of Lima. Nautilus 14: 15 - 16." type="journal article" year="1900">Dall, 1900</bibRefCitation>
)
</taxonomicName>
from the Pliocene of the Los
<collectingRegion id="8854F829FFD12915FBB1FAF3EA94B81E" box="[1070,1168,1367,1393]" country="Philippines" name="Angeles" pageId="41" pageNumber="40">Angeles</collectingRegion>
Basin where it was carefully documented and figured by Woodring (1938).
<bibRefCitation id="2E014B3AFFD12915FA16FADEED9DB8D7" author="Durham, J. W." pageId="41" pageNumber="40" pagination="101 - 212" refId="ref29671" refString="Durham, J. W. 1944. Megafaunal zones of the Oligocene of northwestern Washington. University of California Publications, Department of Geological Sciences, Bulletin 27 (5): 101 - 212. pls. 13 - 18, figs. 1 - 7." type="journal article" year="1944">Durham (1944)</bibRefCitation>
described
<taxonomicName id="8D904D48FFD12915FBB4FA38EAEFB8D7" baseAuthorityName="Durham" baseAuthorityYear="1944" box="[1067,1259,1436,1464]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="41" pageNumber="40" phylum="Mollusca" rank="species" species="twinensis">
<emphasis id="78E4EAD9FFD12915FBB4FA38EAEFB8D7" box="[1067,1259,1436,1464]" italics="true" pageId="41" pageNumber="40">Lima twinensis</emphasis>
</taxonomicName>
and
<taxonomicName id="8D904D48FFD12915FAB0FA38EBF4B8D7" authorityName="Durham" authorityYear="1944" box="[1327,1520,1436,1464]" class="Bivalvia" family="Limidae" genus="Lima" kingdom="Animalia" order="Limida" pageId="41" pageNumber="40" phylum="Mollusca" rank="species" species="robertsae">
<emphasis id="78E4EAD9FFD12915FAB0FA38EBF4B8D7" box="[1327,1520,1436,1464]" italics="true" pageId="41" pageNumber="40">Lima robertsae</emphasis>
</taxonomicName>
from younger Oligocene horizons in the Twin River and Blakeley formations in northeastern
<collectingRegion id="8854F829FFD12915FA81FA40EBB3B891" box="[1310,1463,1508,1534]" country="United States of America" name="Washington" pageId="41" pageNumber="40">Washington</collectingRegion>
, but preservation is inadequate to permit detailed comparisons with the Keasey species other than to note that they are large-shelled species more appropriately assigned to
<taxonomicName id="8D904D48FFD12915FCFEF9CBEA89BBE4" authority="Vokes (1963)" authorityName="Vokes" authorityYear="1963" box="[865,1165,1647,1675]" class="Bivalvia" family="Limidae" genus="Plicacesta" kingdom="Animalia" order="Limida" pageId="41" pageNumber="40" phylum="Mollusca" rank="genus">
<emphasis id="78E4EAD9FFD12915FCFEF9CBEDDFBBE4" box="[865,987,1647,1675]" italics="true" pageId="41" pageNumber="40">Plicacesta</emphasis>
Vokes (1963)
</taxonomicName>
. Durham’s species are more similar to
<taxonomicName id="8D904D48FFD12915FC23F936EB84BBC1" authority="E. J. Moore (1984 a)" authorityName="E. J. Moore" authorityYear="1984" box="[956,1408,1682,1710]" class="Bivalvia" family="Limidae" genus="Plicacesta" kingdom="Animalia" order="Limida" pageId="41" pageNumber="40" phylum="Mollusca" rank="species" species="wilsoni">
<emphasis id="78E4EAD9FFD12915FC23F936EA97BBC1" box="[956,1171,1682,1710]" italics="true" pageId="41" pageNumber="40">Plicacesta wilsoni</emphasis>
E.J. Moore (1984a)
</taxonomicName>
from the uppermost exposed portion of Lincoln Creek Formation along with the Keasey-equivalent giant limid treated and figured separately below.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFD12916FCFFF884EF3CBB58" lastPageId="42" lastPageNumber="41" pageId="41" pageNumber="40" type="materials_examined">
<paragraph id="4A2F36CBFFD12915FCFFF884EACFBAEE" blockId="41.[832,1520,1120,1957]" pageId="41" pageNumber="40">
<emphasis id="78E4EAD9FFD12915FCFFF884EA5BBA55" bold="true" box="[864,1119,1824,1850]" pageId="41" pageNumber="40">Material examined</emphasis>
—11 partial to nearly-complete valves and numerous shell fragments suitable for investigation of shell microstructure.
</paragraph>
<paragraph id="4A2F36CBFFD12915FCFFF82EEBF4BACB" blockId="41.[832,1520,1120,1957]" box="[864,1520,1930,1957]" pageId="41" pageNumber="40">
<emphasis id="78E4EAD9FFD12915FCFFF82EEA4ABACB" bold="true" box="[864,1102,1930,1956]" pageId="41" pageNumber="40">Figured hypotypes</emphasis>
—
<collectionCode id="2C81AE0EFFD12915FBF6F82EEAB1BACB" box="[1129,1205,1930,1956]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="41" pageNumber="40" type="University or college">UCMP</collectionCode>
110740 (left valve), length
</paragraph>
<paragraph id="4A2F36CBFFD22916FF50FFCCEEDCB9B8" blockId="42.[207,1520,101,131]" lastBlockId="42.[112,800,1143,1592]" pageId="42" pageNumber="41">
HICKMAN—Paleogene pteriomorphs from the Keasey Fm.,
<collectingRegion id="8854F829FFD22916FC3EFFC2EDFABDEF" box="[929,1022,102,128]" country="United States of America" name="Oregon" pageId="42" pageNumber="41">Oregon</collectingRegion>
,
<collectingCountry id="3287765BFFD22916FB96FFC2EA39BDEF" box="[1033,1085,102,128]" name="United States of America" pageId="42" pageNumber="41">USA</collectingCountry>
<emphasis id="78E4EAD9FFD22916FB1BFFC1EB8EBDEE" box="[1156,1418,101,131]" italics="true" pageId="42" pageNumber="41">PaleoBios 40(5) 2023</emphasis>
41 7.3 cm, height 7.8 cm., Loc.
<collectionCode id="2C81AE0EFFD22916FE28FBD3EC07B9FE" box="[439,515,1143,1169]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">UCMP</collectionCode>
IP2540.
<collectionCode id="2C81AE0EFFD22916FDF1FBD3ECBEB9FE" box="[622,698,1143,1169]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">UCMP</collectionCode>
110741 (right valve), length 7.8 cm, height 7.7 mm, Loc.
<collectionCode id="2C81AE0EFFD22916FD4BFB3EED24B9DB" box="[724,800,1178,1204]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">UCMP</collectionCode>
110740.
</paragraph>
<caption id="1EEF6643FFD22916FFEFFBABEAB3B92E" ID-DOI="http://doi.org/10.5281/zenodo.10913547" ID-Zenodo-Dep="10913547" httpUri="https://zenodo.org/record/10913547/files/figure.png" pageId="42" pageNumber="41" startId="42.[112,187,1039,1063]" targetBox="[115,1514,192,1009]" targetPageId="42" targetType="figure">
<paragraph id="4A2F36CBFFD22916FFEFFBABEAB3B92E" blockId="42.[112,1504,1039,1090]" pageId="42" pageNumber="41">
<emphasis id="78E4EAD9FFD22916FFEFFBABEEDBB948" bold="true" box="[112,223,1039,1063]" pageId="42" pageNumber="41">Figure 27</emphasis>
.
<taxonomicName id="8D904D48FFD22916FF76FBB4EC42B948" authority="(Clark, 1925)" baseAuthorityName="CLARK" baseAuthorityYear="1925" box="[233,582,1039,1064]" class="Bivalvia" family="Limidae" genus="Acesta" kingdom="Animalia" order="Limida" pageId="42" pageNumber="41" phylum="Mollusca" rank="species" species="oregonensis">
<emphasis id="78E4EAD9FFD22916FF76FBB4EFB4B947" box="[233,432,1040,1064]" italics="true" pageId="42" pageNumber="41">Acesta oregonensis</emphasis>
(
<bibRefCitation id="2E014B3AFFD22916FE5FFBABEC38B948" author="Clark, B. L." box="[448,572,1039,1063]" pageId="42" pageNumber="41" pagination="69 - 136" refId="ref28111" refString="Clark, B. L. 1925. Pelecypoda from the marine Oligocene of western North America. University of California Bulletin of the Department of Geological Sciences 15 (4): 69 - 136, pls. 8 - 22." type="journal article" year="1925">Clark, 1925</bibRefCitation>
)
</taxonomicName>
. Hypotype, UCMP 110741.
<emphasis id="78E4EAD9FFD22916FCF4FBABED78B948" bold="true" box="[875,892,1039,1063]" pageId="42" pageNumber="41">A</emphasis>
. Right valve in matrix with adhering interior (aragonitic) and exterior (calcitic) shell layers. Scale bar=5 cm.
<emphasis id="78E4EAD9FFD22916FD0EFB8EECA5B92D" bold="true" box="[657,673,1066,1090]" pageId="42" pageNumber="41">B</emphasis>
. Fragments of exfoliated aragonitic layers from
<emphasis id="78E4EAD9FFD22916FB3DFB8DEAB7B92E" bold="true" box="[1186,1203,1065,1089]" pageId="42" pageNumber="41">A</emphasis>
.
</paragraph>
</caption>
<paragraph id="4A2F36CBFFD22916FF0FFB44EC7BB871" blockId="42.[112,800,1143,1592]" pageId="42" pageNumber="41">
<emphasis id="78E4EAD9FFD22916FF0FFB44EFA0B994" bold="true" box="[144,420,1248,1275]" pageId="42" pageNumber="41">Unfigured hypotype</emphasis>
—
<collectionCode id="2C81AE0EFFD22916FE5FFB45EC0AB994" box="[448,526,1249,1275]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">UCMP</collectionCode>
110742 (incomplete righty valve in matrix), Loc.
<collectionCode id="2C81AE0EFFD22916FE54FAA0EC13B871" box="[459,535,1284,1310]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">UCMP</collectionCode>
IP2540.
</paragraph>
<paragraph id="4A2F36CBFFD22916FF0FFA83EF0AB89E" blockId="42.[112,800,1143,1592]" pageId="42" pageNumber="41">
<emphasis id="78E4EAD9FFD22916FF0FFA83EF0AB82E" bold="true" box="[144,270,1319,1345]" pageId="42" pageNumber="41">Localities</emphasis>
—
<materialsCitation id="FAF83C96FFD22916FEB6FA83ECD5B82D" box="[297,721,1319,1346]" collectionCode="USGS" pageId="42" pageNumber="41" specimenCode="USGS 15269, 15315, 15518, 15601" specimenCount="1">
<specimenCode id="1A369EB0FFD22916FEB6FA83EFC5B82D" box="[297,449,1319,1346]" collectionCode="USGS" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="42" pageNumber="41">USGS 15269</specimenCode>
,
<specimenCode id="1A369EB0FFD22916FE56FA83EC1EB82D" box="[457,538,1319,1346]" collectionCode="USGS" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="42" pageNumber="41">15315</specimenCode>
,
<specimenCode id="1A369EB0FFD22916FDBDFA83EC77B82D" box="[546,627,1319,1346]" collectionCode="USGS" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="42" pageNumber="41">15518</specimenCode>
,
<specimenCode id="1A369EB0FFD22916FDE3FA83ECC8B82D" box="[636,716,1319,1346]" collectionCode="USGS" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="42" pageNumber="41">15601</specimenCode>
,
</materialsCitation>
<materialsCitation id="FAF83C96FFD22916FD4AFA83EFC1B80A" collectionCode="UCMP" pageId="42" pageNumber="41" specimenCode="UCMP 4118" specimenCount="1">
<specimenCode id="1A369EB0FFD22916FD4AFA83EEB5B80B" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">UCMP 4118</specimenCode>
(Clark’s
<typeStatus id="952B8869FFD22916FE80FAEFEF52B80A" box="[287,342,1355,1381]" pageId="42" pageNumber="41">type</typeStatus>
locality)
</materialsCitation>
,
<materialsCitation id="FAF83C96FFD22916FE4EFAEEED18B80A" box="[465,796,1354,1381]" collectionCode="UCMP" pageId="42" pageNumber="41" specimenCode="IP2540, 2549, 2550" specimenCount="1">
<collectionCode id="2C81AE0EFFD22916FE4EFAEEEC19B80B" box="[465,541,1354,1380]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">UCMP</collectionCode>
<specimenCode id="1A369EB0FFD22916FDBCFAEEEC84B80A" box="[547,640,1354,1381]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">IP2540</specimenCode>
,
<specimenCode id="1A369EB0FFD22916FD14FAEEECC9B80A" box="[651,717,1354,1381]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">2549</specimenCode>
,
<specimenCode id="1A369EB0FFD22916FD46FAEEED18B80A" box="[729,796,1354,1381]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="42" pageNumber="41" type="University or college">2550</specimenCode>
</materialsCitation>
. The
<typeStatus id="952B8869FFD22916FF37FACAEEDBB8E7" box="[168,223,1390,1416]" pageId="42" pageNumber="41">type</typeStatus>
locality is an abandoned railroad tunnel with outcrops that are no longer accessible, but the beds fall within the stratigraphic interval represented by the upper member.
</paragraph>
<paragraph id="4A2F36CBFFD22916FF0FFA5EEF3CBB58" blockId="42.[112,800,1143,1592]" pageId="42" pageNumber="41">
<emphasis id="78E4EAD9FFD22916FF0FFA5EEF8DBB7B" bold="true" box="[144,393,1530,1556]" pageId="42" pageNumber="41">Stratigraphic range</emphasis>
—Restricted to the upper member of the Keasey Formation.
</paragraph>
</subSubSection>
</treatment>
</document>
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