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<document id="F7BF15B478EC647644C34C434DA46EF1" ID-CLB-Dataset="24637" ID-DOI="10.11646/zootaxa.4711.1.1" ID-GBIF-Dataset="b37adfc9-f522-4496-beec-6fb0231e325b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3573366" ID-ZooBank="604227AA-58EB-408C-8794-6E30192C3F74" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576220502254" checkinUser="plazi" docAuthor="Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. &amp; Maleki-Ravasan, Naseh" docDate="2019" docId="B056F818FF862552FF1BAD01A9A6CD91" docLanguage="en" docName="zootaxa.4711.1.1.pdf" docOrigin="Zootaxa 4711 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaselia khaghaniniai Khameneh &amp; Khaghaninia &amp; Disney &amp; Maleki-Ravasan 2019, n. sp." docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="4C6F8060FF9F2548FF8CAB6BAC37CF3A" masterDocTitle="Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran" masterLastPageNumber="50" masterPageNumber="1" pageNumber="26" updateTime="1698773750373" updateUser="ExternalLinkService">
<mods:mods id="B7D471C1CC9C60686B1D8EADEF7A2C1C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="639CE06F51FCB305C8016A88DFE8CE8F">Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran</mods:title>
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<mods:name id="68E9D9B2E1AF95A610DC3A1B87B897E9" type="personal">
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<mods:namePart id="C8E6113EF576273D67BA54A7A7CC2684">Khameneh, Roya Namaki</mods:namePart>
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<mods:namePart id="951B19C2A90D3472F43B2890623923DD">Khaghaninia, Samad</mods:namePart>
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<mods:namePart id="5FD33DF8F9FA53A1657893A7214D2A83">Disney, R. Henry L.</mods:namePart>
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<mods:namePart id="556859B1930DECAB393B98ADCAF4C64B">Maleki-Ravasan, Naseh</mods:namePart>
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<mods:title id="CFD0D15C721650747F9F7BB5FAF0E531">Zootaxa</mods:title>
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<mods:date id="B55C68475E4343DA91CC24DB7903C845">2019</mods:date>
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<mods:number id="647953E31A526C0AAC8D8F0EB1E65AA1">2019-12-13</mods:number>
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<treatment id="B056F818FF862552FF1BAD01A9A6CD91" ID-DOI="http://doi.org/10.5281/zenodo.5933512" ID-GBIF-Taxon="160702432" ID-Zenodo-Dep="5933512" LSID="urn:lsid:plazi:treatment:B056F818FF862552FF1BAD01A9A6CD91" httpUri="http://treatment.plazi.org/id/B056F818FF862552FF1BAD01A9A6CD91" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<subSubSection id="70E51A85FF862551FF1BAD01AED9C9BE" box="[151,750,1642,1669]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="3840490EFF862551FF1BAD01AED9C9BE" blockId="25.[151,750,1642,1704]" box="[151,750,1642,1669]" pageId="25" pageNumber="26">
<heading id="6308FE62FF862551FF1BAD01AED9C9BE" bold="true" box="[151,750,1642,1669]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="0A8B951CFF862551FF1BAD01AED9C9BE" bold="true" box="[151,750,1642,1669]" pageId="25" pageNumber="26">
<taxonomicName id="FFFF328DFF862551FF1BAD01AE93C9BE" ID-CoL="9J24C" authority="Namaki &amp; Disney" authorityName="Khameneh &amp; Khaghaninia &amp; Disney &amp; Maleki-Ravasan" authorityYear="2019" box="[151,676,1642,1669]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="khaghaniniai" status="sp. nov.">
<emphasis id="0A8B951CFF862551FF1BAD01AD8AC9BF" bold="true" box="[151,445,1642,1669]" italics="true" pageId="25" pageNumber="26">Megaselia khaghaniniai</emphasis>
Namaki &amp; Disney
</taxonomicName>
<taxonomicNameLabel id="11B82867FF862551FD27AD01AED9C9BE" box="[683,750,1642,1668]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="70E51A85FF862551FF1BADE5AD74C992" box="[151,323,1677,1704]" pageId="25" pageNumber="26" type="description">
<paragraph id="3840490EFF862551FF1BADE5AD74C992" blockId="25.[151,750,1642,1704]" box="[151,323,1677,1704]" pageId="25" pageNumber="26">
(
<figureCitation id="A0C4558BFF862551FF13ADE5AD0CC992" box="[159,315,1677,1704]" captionStart-0="FIGURES 165172" captionStart-1="FIGURES 173176" captionStartId-0="28.[151,264,1945,1970]" captionStartId-1="29.[151,264,924,949]" captionTargetBox-0="[174,1416,203,1904]" captionTargetBox-1="[174,1423,187,886]" captionTargetId-0="figure@28.[156,1432,181,1920]" captionTargetId-1="figure@29.[151,1436,181,898]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionText-0="FIGURES 165172. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 165, whole fly; 166, frons; 167, detail of frons; 168, postpedicels, palps and proboscis; 169, outer half of abdomen; 170, left face of epandrium; 171, epandrium and anal tube; 172, hypandrium and penis complex. Scale bars 20 μm." captionText-1="FIGURES 173176. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 173, right face of hypopygium; 174, front tarsus; 175, hind femur and tibia; 176, wing. Scale bars 20 μm." figureDoi-0="http://doi.org/10.5281/zenodo.3573414" figureDoi-1="http://doi.org/10.5281/zenodo.3573416" httpUri-0="https://zenodo.org/record/3573414/files/figure.png" httpUri-1="https://zenodo.org/record/3573416/files/figure.png" pageId="25" pageNumber="26">Figs 165176</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="70E51A85FF862551FF1BADBEA828C82E" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="3840490EFF862551FF1BADBEA828C82E" blockId="25.[151,1437,1749,2029]" pageId="25" pageNumber="26">
<emphasis id="0A8B951CFF862551FF1BADBEADB6C9CA" bold="true" box="[151,385,1749,1776]" pageId="25" pageNumber="26">Material examined.</emphasis>
<materialsCitation id="88974353FF862551FE05ADBDA828C82E" ID-GBIF-Occurrence="2468988936" collectingDate="2012-05-22" collectorName="S. Khaghaninia" country="Azerbaijan" county="Pere region" elevation="1405" latitude="38.695316" location="West" longLatPrecision="1" longitude="44.900684" municipality="Khoy" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<typeStatus id="E744F7ACFF862551FE05ADBDADC6C9CA" box="[393,497,1750,1776]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
male,
<location id="3D201FD5FF862551FDB1ADBCAE43C9CB" LSID="urn:lsid:plazi:treatment:B056F818FF862552FF1BAD01A9A6CD91:3D201FD5FF862551FDB1ADBCAE43C9CB" box="[573,628,1751,1777]" country="Azerbaijan" county="Pere region" latitude="38.695316" longLatPrecision="1" longitude="44.900684" municipality="Khoy" name="West" pageId="25" pageNumber="26">West</location>
<collectingCountry id="40E8099EFF862551FDF7ADBDAEC1C9CA" box="[635,758,1750,1776]" name="Azerbaijan" pageId="25" pageNumber="26">Azerbaijan</collectingCountry>
province,
<collectingMunicipality id="D824D374FF862551FCE2ADBDAF9BC9CA" box="[878,940,1750,1776]" pageId="25" pageNumber="26">Khoy</collectingMunicipality>
city,
<collectingCounty id="D1213182FF862551FC67ADBCA859C9CA" box="[1003,1134,1750,1777]" pageId="25" pageNumber="26">Pere region</collectingCounty>
,
<geoCoordinate id="5DCB2FC9FF862551FBF7ADBDA922C9CB" box="[1147,1301,1750,1777]" degrees="38" direction="north" minutes="41.719" orientation="latitude" pageId="25" pageNumber="26" precision="1" value="38.695316">38°41.719 N</geoCoordinate>
,
<geoCoordinate id="5DCB2FC9FF862551FAADADBEAC9DC82F" degrees="44" direction="east" minutes="54.041" orientation="longitude" pageId="25" pageNumber="26" precision="1" value="44.900684">44°54.041 E</geoCoordinate>
,
<quantity id="FF07E4EBFF862551FF3AAD92AD3AC82F" box="[182,269,1785,1813]" metricMagnitude="3" metricUnit="m" metricValue="1.405" pageId="25" pageNumber="26" unit="m" value="1405.0">
<elevation id="B3D2AE3DFF862551FF3AAD92AD3AC82F" box="[182,269,1785,1813]" metricMagnitude="3" metricUnit="m" metricValue="1.405" pageId="25" pageNumber="26" unit="m" value="1405.0">1405 m</elevation>
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, montane grassland,
<date id="4C416FCEFF862551FE74AD91AE5AC82E" box="[504,621,1786,1812]" pageId="25" pageNumber="26" value="2012-05-22">
<collectingDate id="5C059626FF862551FE74AD91AE5AC82E" box="[504,621,1786,1812]" pageId="25" pageNumber="26" value="2012-05-22">22.V.2012</collectingDate>
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,
<collectorName id="950A2CD8FF862551FDF5AD91AF12C82E" box="[633,805,1786,1812]" pageId="25" pageNumber="26">S. Khaghaninia</collectorName>
(57, CUMZ—13-99).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="70E51A85FF862552FF4BAC76AD98CDBD" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="3840490EFF862552FF4BAC76A8F8CECD" blockId="25.[151,1437,1749,2029]" lastBlockId="26.[151,1436,153,683]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<emphasis id="0A8B951CFF862551FF4BAC76AD93C802" bold="true" box="[199,420,1821,1848]" pageId="25" pageNumber="26">Description. Male.</emphasis>
Whole fly as
<figureCitation id="A0C4558BFF862551FDCFAC75AE93C802" box="[579,676,1821,1848]" captionStart="FIGURES 165172" captionStartId="28.[151,264,1945,1970]" captionTargetBox="[174,1416,203,1904]" captionTargetId="figure@28.[156,1432,181,1920]" captionTargetPageId="28" captionText="FIGURES 165172. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 165, whole fly; 166, frons; 167, detail of frons; 168, postpedicels, palps and proboscis; 169, outer half of abdomen; 170, left face of epandrium; 171, epandrium and anal tube; 172, hypandrium and penis complex. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3573414" httpUri="https://zenodo.org/record/3573414/files/figure.png" pageId="25" pageNumber="26">Fig. 165</figureCitation>
. Frons as
<figureCitation id="A0C4558BFF862551FC9AAC75AF41C802" box="[790,886,1822,1848]" captionStart="FIGURES 165172" captionStartId="28.[151,264,1945,1970]" captionTargetBox="[174,1416,203,1904]" captionTargetId="figure@28.[156,1432,181,1920]" captionTargetPageId="28" captionText="FIGURES 165172. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 165, whole fly; 166, frons; 167, detail of frons; 168, postpedicels, palps and proboscis; 169, outer half of abdomen; 170, left face of epandrium; 171, epandrium and anal tube; 172, hypandrium and penis complex. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3573414" httpUri="https://zenodo.org/record/3573414/files/figure.png" pageId="25" pageNumber="26">Fig. 166</figureCitation>
, with dense but very fine microtrichia (
<figureCitation id="A0C4558BFF862551FAB8AC75A9A3C803" box="[1332,1428,1822,1849]" captionStart="FIGURES 165172" captionStartId="28.[151,264,1945,1970]" captionTargetBox="[174,1416,203,1904]" captionTargetId="figure@28.[156,1432,181,1920]" captionTargetPageId="28" captionText="FIGURES 165172. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 165, whole fly; 166, frons; 167, detail of frons; 168, postpedicels, palps and proboscis; 169, outer half of abdomen; 170, left face of epandrium; 171, epandrium and anal tube; 172, hypandrium and penis complex. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3573414" httpUri="https://zenodo.org/record/3573414/files/figure.png" pageId="25" pageNumber="26">Fig. 167</figureCitation>
) and notable for the lower supra-antennal bristles being as long as the upper pair, which are further then the pre-ocellar bristles, and the anterolaterals being well above the antials which are situated close to the eye margins. Cheek with a single bristle and jowl likewise but it is longer and more robust. Postpedicels brown, without SPS vesicles, the yellow palps and proboscis as
<figureCitation id="A0C4558BFF862551FD90ACC5AE4AC8F2" box="[540,637,1966,1992]" captionStart="FIGURES 165172" captionStartId="28.[151,264,1945,1970]" captionTargetBox="[174,1416,203,1904]" captionTargetId="figure@28.[156,1432,181,1920]" captionTargetPageId="28" captionText="FIGURES 165172. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 165, whole fly; 166, frons; 167, detail of frons; 168, postpedicels, palps and proboscis; 169, outer half of abdomen; 170, left face of epandrium; 171, epandrium and anal tube; 172, hypandrium and penis complex. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3573414" httpUri="https://zenodo.org/record/3573414/files/figure.png" pageId="25" pageNumber="26">Fig. 168</figureCitation>
, the labella with only a few short hairs/spinules below. Thorax brown. With 3 notopleural bristles and no cleft in front of these. Mesopleuron bare. Scutellum with an anterior pair of small hairs and a posterior pair of bristles. Abdominal tergites brown with small hairs hairs, but 12 longer ones at sides of T2 and at rear of T6; and venter paler brown with hairs on segments 36. Hypopygium as
<figureCitation id="A0C4558BFF852552FB14ABD6A904CFED" box="[1176,1331,189,215]" captionStart-0="FIGURES 165172" captionStart-1="FIGURES 173176" captionStartId-0="28.[151,264,1945,1970]" captionStartId-1="29.[151,264,924,949]" captionTargetBox-0="[174,1416,203,1904]" captionTargetBox-1="[174,1423,187,886]" captionTargetId-0="figure@28.[156,1432,181,1920]" captionTargetId-1="figure@29.[151,1436,181,898]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionText-0="FIGURES 165172. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 165, whole fly; 166, frons; 167, detail of frons; 168, postpedicels, palps and proboscis; 169, outer half of abdomen; 170, left face of epandrium; 171, epandrium and anal tube; 172, hypandrium and penis complex. Scale bars 20 μm." captionText-1="FIGURES 173176. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 173, right face of hypopygium; 174, front tarsus; 175, hind femur and tibia; 176, wing. Scale bars 20 μm." figureDoi-0="http://doi.org/10.5281/zenodo.3573414" figureDoi-1="http://doi.org/10.5281/zenodo.3573416" httpUri-0="https://zenodo.org/record/3573414/files/figure.png" httpUri-1="https://zenodo.org/record/3573416/files/figure.png" pageId="26" pageNumber="27">Figs 169173</figureCitation>
, the hairs on epandrium being smaller than those at rear of tergite 6, on cerci and proctor apart from one posterolaterally that is more robust than rest and longer than the hairs of procter. The hypandrial lobe of the right side is vestigial. Legs yellow apart from dark tip to hind femur. Fore tarsus slender with postero-dorsal hair palisades on segments 15 (
<figureCitation id="A0C4558BFF852552FF2CAA26AD36CE5D" box="[160,257,333,359]" captionStart="FIGURES 173176" captionStartId="29.[151,264,924,949]" captionTargetBox="[174,1423,187,886]" captionTargetId="figure@29.[151,1436,181,898]" captionTargetPageId="29" captionText="FIGURES 173176. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 173, right face of hypopygium; 174, front tarsus; 175, hind femur and tibia; 176, wing. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3573416" httpUri="https://zenodo.org/record/3573416/files/figure.png" pageId="26" pageNumber="27">Fig. 174</figureCitation>
). Dorsal hair palisade of mid tibia extends about 0.58 times its length. Hairs below basal half of hind femur longer than those of the anteroventral row (
<figureCitation id="A0C4558BFF852552FD0CAA1AAED4CEB1" box="[640,739,368,395]" captionStart="FIGURES 173176" captionStartId="29.[151,264,924,949]" captionTargetBox="[174,1423,187,886]" captionTargetId="figure@29.[151,1436,181,898]" captionTargetPageId="29" captionText="FIGURES 173176. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 173, right face of hypopygium; 174, front tarsus; 175, hind femur and tibia; 176, wing. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3573416" httpUri="https://zenodo.org/record/3573416/files/figure.png" pageId="26" pageNumber="27">Fig. 175</figureCitation>
). Hind tibia with 10 differentiated posterodorsal hairs with all but the basal 2 robust (
<figureCitation id="A0C4558BFF852552FE18AAFEADC3CE95" box="[404,500,404,431]" captionStart="FIGURES 173176" captionStartId="29.[151,264,924,949]" captionTargetBox="[174,1423,187,886]" captionTargetId="figure@29.[151,1436,181,898]" captionTargetPageId="29" captionText="FIGURES 173176. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 173, right face of hypopygium; 174, front tarsus; 175, hind femur and tibia; 176, wing. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3573416" httpUri="https://zenodo.org/record/3573416/files/figure.png" pageId="26" pageNumber="27">Fig. 175</figureCitation>
), without anterodorsals, and spinules of apical combs simple. Wings (
<figureCitation id="A0C4558BFF852552FB71AAFEA96BCE8A" box="[1277,1372,405,432]" captionStart="FIGURES 173176" captionStartId="29.[151,264,924,949]" captionTargetBox="[174,1423,187,886]" captionTargetId="figure@29.[151,1436,181,898]" captionTargetPageId="29" captionText="FIGURES 173176. Megaselia khaghaniniai Namaki &amp; Disney n. sp. male. 173, right face of hypopygium; 174, front tarsus; 175, hind femur and tibia; 176, wing. Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.3573416" httpUri="https://zenodo.org/record/3573416/files/figure.png" pageId="26" pageNumber="27">Fig. 176</figureCitation>
)
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long. Costal index 0.49. Costal ratios 3.34: 2.31: 1. Costal cilia (of section 3)
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long. No vein 3 hair. 2 unequal axillary bristles, the outermost being
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long. Sc not reaching R1. Haltere brown.
</paragraph>
<paragraph id="3840490EFF852552FF4BA96BAD98CDBD" blockId="26.[151,1436,153,683]" pageId="26" pageNumber="27">
<emphasis id="0A8B951CFF852552FF4BA96BAD6ACD21" bold="true" box="[199,349,512,539]" pageId="26" pageNumber="27">Recognition.</emphasis>
In the key to the
<taxonomicName id="FFFF328DFF852552FD93A96AAEA3CD21" authorityName="Rondani" authorityYear="1856" box="[543,660,513,539]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="0A8B951CFF852552FD93A96AAEA3CD21" box="[543,660,513,539]" italics="true" pageId="26" pageNumber="27">Megaselia</emphasis>
</taxonomicName>
males of the British Isles (
<bibRefCitation id="5C6E34FFFF852552FC32A96AA878CD21" author="Disney, R. H. L." box="[958,1103,513,539]" pageId="26" pageNumber="27" pagination="1 - 155" refId="ref21228" refString="Disney, R. H. L. (1989) Scuttle Flies-Diptera Phoridae Genus Megaselia. Handbooks for the Identification of British Insects, 10 (8), 1 - 155." type="journal article" year="1989">Disney 1989</bibRefCitation>
) it runs to couplet 203, where each lead only applies in part. In
<bibRefCitation id="5C6E34FFFF852552FD86A94EAED1CD7A" author="Borgmeier, T." box="[522,742,549,576]" pageId="26" pageNumber="27" pagination="1 - 160" refId="ref20965" refString="Borgmeier, T. (1966) Revision of the North American Phorid flies. Part III. The species of the genus Megaselia, subgenus Megaselia (Diptera, Phoridae). Studia Entomologica, Petropolis, 8, 1 - 160. [1965]" type="journal article" year="1966">Borgmeiers (1966)</bibRefCitation>
key to Nearctic Group VII it runs to couplet 37 lead
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1
<taxonomicName id="FFFF328DFF852552FAC6A94DA95ECD05" authority="(Coquillett)" authorityName="Coquillett" baseAuthorityName="Coquillett" baseAuthorityYear="1895" box="[1354,1385,550,575]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fungicola">
<emphasis id="0A8B951CFF852552FAC6A94DA95ECD05" box="[1354,1385,550,575]" italics="true" pageId="26" pageNumber="27">M.</emphasis>
</taxonomicName>
</specimenCount>
fungicola (Coquillett) but its lower supra-antennal bristles are much shorter and finer than the upper pair and its wing is less than
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long.
</paragraph>
</subSubSection>
<subSubSection id="70E51A85FF852552FF4BA9FAA9A6CD91" box="[199,1425,657,683]" pageId="26" pageNumber="27" type="etymology">
<paragraph id="3840490EFF852552FF4BA9FAA9A6CD91" blockId="26.[151,1436,153,683]" box="[199,1425,657,683]" pageId="26" pageNumber="27">
<emphasis id="0A8B951CFF852552FF4BA9FAAD79CD91" bold="true" box="[199,334,657,683]" pageId="26" pageNumber="27">Etymology.</emphasis>
Named after professor Samad Khaghaninia, who collected the specimens described in this paper.
</paragraph>
</subSubSection>
</treatment>
</document>