treatments-xml/data/A1/85/AF/A185AFE24AA551278669452B0925B575.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.96.48312" ID-GBIF-Dataset="7de3ea60-ba35-4831-ae1d-048dc362f8b0" ID-GBIF-Taxon="161939046" ID-Pensoft-Pub="1860-0743-1-25" ID-Pensoft-UUID="4EDA9B5B777551B7BBB0152DD188E5EF" ID-ZooBank="F2C8585A1268443693348B64AE20F6EE" ModsDocID="1860-0743-1-25" checkinTime="1580744411276" checkinUser="pensoft" docAuthor="Sands, Arthur F, Gloeer, Peter, Guerlek, Mustafa E, Albrecht, Christian &amp; Neubauer, Thomas A" docDate="2020" docId="A185AFE24AA551278669452B0925B575" docLanguage="en" docName="ZoosystEvol 96(1): 25-66" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.48312" docTitle="Theodoxus jordani" docType="treatment" docVersion="6" id="4EDA9B5B777551B7BBB0152DD188E5EF" lastPageNumber="25" masterDocId="4EDA9B5B777551B7BBB0152DD188E5EF" masterDocTitle="A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species" masterLastPageNumber="66" masterPageNumber="25" pageId="0" pageNumber="25" updateTime="1635972787372" updateUser="ExternalLinkService">
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<mods:title>A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sands, Arthur F</mods:namePart>
<mods:affiliation>Justus Liebig Univrsity Giessen, Giessen, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gloeer, Peter</mods:namePart>
<mods:affiliation>Biodiversity Research Laboratory, Hetlingen, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Guerlek, Mustafa E</mods:namePart>
<mods:affiliation>Mehmet Akif Ersoy niversity, Burdur, Turkey</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Albrecht, Christian</mods:namePart>
<mods:affiliation>Justus Liebig University Giessen, Giessen, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Neubauer, Thomas A</mods:namePart>
<mods:affiliation>Justus Liebig University Giessen, Giessen, Germany &amp; Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
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<mods:date>2020</mods:date>
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<mods:number>96</mods:number>
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<treatment ID-GBIF-Taxon="161939046" LSID="urn:lsid:plazi:treatment:A185AFE24AA551278669452B0925B575" httpUri="http://treatment.plazi.org/id/A185AFE24AA551278669452B0925B575" lastPageNumber="25" pageId="0" pageNumber="25">
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<taxonomicName LSID="A185AFE2-4AA5-5127-8669-452B0925B575" authority="(G. B. Sowerby I, 1836)" baseAuthorityName="G. B. Sowerby I" baseAuthorityYear="1836" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">Theodoxus jordani (G.B. Sowerby I, 1836)</taxonomicName>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure15" httpUri="https://binary.pensoft.net/fig/376464" pageId="0" pageNumber="25">Figures 15A-V</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Theodoxus jordani (G. B. Sowerby I, 1836). A, B. Syntype of N. euphratica from the Euphrates River at Samawah, Iraq (ZMZ 528916); C - E. Syntype of N. michonii from Syria (MHNG-MOLL- 111706); F - H. Syntype of N. karasuna collected from the Karasu River, south-eastern Turkey (ZMZ 528937); I, J; K, L. Two syntypes of N. ponsoti from Lake Muzayrīb, Syria (MNHN-IM- 2000 - 32754); M. Syntype (labelled as paratype) of N. ponsoti from Lake Muzayrīb (MT 820, RBINS); N. Syntype of N. jordani var. turris collected in the Sea of Galilee (ZMZ 528930); O, P. Syntype of N. bellardii from the Litani River, Beqaa valley, Lebanon (ZMZ 528918). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure16" httpUri="https://binary.pensoft.net/fig/376465" pageId="0" pageNumber="25">, 16A-P</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure17" httpUri="https://binary.pensoft.net/fig/376466" pageId="0" pageNumber="25">, 17A-J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="reference_group">
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Jordani G.B. Sowerby 
<date value="1836-01">I 1836</date>
: 4, pl. 99, fig. 49; G.B. Sowerby 
<date value="1849-02">II 1849</date>
: 531, pl. 115, figs 213-215; 
<bibRefCitation author="Bourguignat, JR" journalOrPublisher="Baudry, Paris" pageId="0" pageNumber="25" publicationUrl="http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item" refId="B31" refString="Bourguignat, JR, 1853. Catalogue raisonne des mollusques terrestres et fluviatiles recueillis par M. F. de Saulcy pendant son voyage en Orient. Gide et J. Baudry, Paris, http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item" title="Catalogue raisonne des mollusques terrestres et fluviatiles recueillis par M. F. de Saulcy pendant son voyage en Orient. Gide et J." url="http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item" year="1853">Bourguignat 1853</bibRefCitation>
: 69; 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 84-86, pl. 2, figs 14-16; 
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 2-3, pl. 211, fig. 1319 (as &quot; 
<taxonomicName family="Neritidae" genus="N." higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="N. jordanica" order="Cycloneritida" pageId="0" pageNumber="25" rank="species" species="jordanica">N. jordanica</taxonomicName>
&quot; in figure caption).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Michonii 
<bibRefCitation author="Bourguignat, JR" journalOrPublisher="Bailliere, Lutetia" pageId="0" pageNumber="25" publicationUrl="https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image" refId="B30" refString="Bourguignat, JR, 1852. Testacea novissima quae Cl. de Saulcy in itinere per Orientem annis 1850 et 1851, collegit. J.-B. Bailliere, Lutetia, https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image" title="Testacea novissima quae Cl. de Saulcy in itinere per Orientem annis 1850 et 1851, collegit. J. - B." url="https://gallica.bnf.fr/ark:/12148/bpt6k99037b.image" year="1852">Bourguignat 1852</bibRefCitation>
: 25; 
<bibRefCitation author="Bourguignat, JR" journalOrPublisher="Baudry, Paris" pageId="0" pageNumber="25" publicationUrl="http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item" refId="B31" refString="Bourguignat, JR, 1853. Catalogue raisonne des mollusques terrestres et fluviatiles recueillis par M. F. de Saulcy pendant son voyage en Orient. Gide et J. Baudry, Paris, http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item" title="Catalogue raisonne des mollusques terrestres et fluviatiles recueillis par M. F. de Saulcy pendant son voyage en Orient. Gide et J." url="http://gallica.bnf.fr/ark:/12148/bpt6k99038p/f1.item" year="1853">Bourguignat 1853</bibRefCitation>
: 70, pl. 2, figs 48-51.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Bellardii 
<bibRefCitation author="Mousson, A" journalOrPublisher="Zurcher et Furrer, Zuerich" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/47588377" refId="B109" refString="Mousson, A, 1854. Coquilles terrestres et fluviatiles recueillies par M. le prof Bellardi dans un voyage en Orient. Zurcher et Furrer, Zuerich, https://biodiversitylibrary.org/page/47588377" title="Coquilles terrestres et fluviatiles recueillies par M. le prof Bellardi dans un voyage en Orient." url="https://biodiversitylibrary.org/page/47588377" year="1854">Mousson 1854</bibRefCitation>
: 52-53.
</paragraph>
<paragraph pageId="0" pageNumber="25">
? 
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Africana 
<bibRefCitation author="Reeve, LA" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/9141496" refId="B134" refString="Reeve, LA, 1856. Monograph of the genus Neritina. In: Reeve LA (Ed.) Conchologia iconica: or, Illustrations of the shells of molluscous animals. Vol. 4. Privately published, London. [1-37 pls. and unpaginated index] https://biodiversitylibrary.org/page/9141496" title="Monograph of the genus Neritina. In: Reeve LA (Ed.) Conchologia iconica: or, Illustrations of the shells of molluscous animals. Vol. 4. Privately published, London. [1 - 37 pls. and unpaginated index]" url="https://biodiversitylibrary.org/page/9141496" year="1856">Reeve 1856</bibRefCitation>
: pl. 30, fig. 138a, b.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Nilotica 
<bibRefCitation author="Reeve, LA" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/9141496" refId="B134" refString="Reeve, LA, 1856. Monograph of the genus Neritina. In: Reeve LA (Ed.) Conchologia iconica: or, Illustrations of the shells of molluscous animals. Vol. 4. Privately published, London. [1-37 pls. and unpaginated index] https://biodiversitylibrary.org/page/9141496" title="Monograph of the genus Neritina. In: Reeve LA (Ed.) Conchologia iconica: or, Illustrations of the shells of molluscous animals. Vol. 4. Privately published, London. [1 - 37 pls. and unpaginated index]" url="https://biodiversitylibrary.org/page/9141496" year="1856">Reeve 1856</bibRefCitation>
: pl. 34, fig. 157a, b; 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 82-83, pl. 2, figs 17-19, pl. 13, figs 14-16; 
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 2, pl. 211, fig. 1317.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Sowerby" authorityYear="1832" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina Jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Jordani">Neritina Jordani</taxonomicName>
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="var. turris" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="Jordani" variety="turris">var. turris</taxonomicName>
<bibRefCitation author="Mousson, A" journalOrPublisher="Zurcher et Furrer, Zuerich" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/8742661" refId="B110" refString="Mousson, A, 1861. Coquilles terrestres et fluviatiles recueillies par M. le Prof. J. R. Roth dans son dernier voyage en Orient. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zuerich 6: 1-34, 124-156. https://biodiversitylibrary.org/page/8742661" title="Coquilles terrestres et fluviatiles recueillies par M. le Prof. J. R. Roth dans son dernier voyage en Orient. Vierteljahrsschrift der Naturforschenden Gesellschaft in Zuerich 6: 1 - 34, 124 - 156." url="https://biodiversitylibrary.org/page/8742661" year="1861">Mousson 1861</bibRefCitation>
: 151-152; 
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 3, pl. 211, fig. 1320.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Karasuna 
<bibRefCitation author="Mousson, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="25" pagination="5 - 60" publicationUrl="https://biodiversitylibrary.org/page/15661760" refId="B111" refString="Mousson, A, 1874. Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient. Journal de Conchyliologie 22 (1): 5 - 60, https://biodiversitylibrary.org/page/15661760" title="Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient." url="https://biodiversitylibrary.org/page/15661760" volume="22" year="1874">Mousson 1874</bibRefCitation>
: 34-35; 
<bibRefCitation author="Blanckenhorn, M" journalOrPublisher="Palaeontographica" pageId="0" pageNumber="25" pagination="71 - 144" publicationUrl="https://biodiversitylibrary.org/page/33130542" refId="B25" refString="Blanckenhorn, M, 1897. Zur Kenntnis der Suesswasserablagerungen und Mollusken Syriens. Palaeontographica 44: 71 - 144, https://biodiversitylibrary.org/page/33130542" title="Zur Kenntnis der Suesswasserablagerungen und Mollusken Syriens." url="https://biodiversitylibrary.org/page/33130542" volume="44" year="1897">Blanckenhorn 1897</bibRefCitation>
: 101-102, pl. 8, figs 6-8.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina meridionalis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="meridionalis">Neritina meridionalis</taxonomicName>
var. 
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="var. Mesopotamica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="meridionalis" variety="Mesopotamica">Mesopotamica</taxonomicName>
<bibRefCitation author="Mousson, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="25" pagination="5 - 60" publicationUrl="https://biodiversitylibrary.org/page/15661760" refId="B111" refString="Mousson, A, 1874. Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient. Journal de Conchyliologie 22 (1): 5 - 60, https://biodiversitylibrary.org/page/15661760" title="Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient." url="https://biodiversitylibrary.org/page/15661760" volume="22" year="1874">Mousson 1874</bibRefCitation>
: 35.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Mousson" authorityYear="1874" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina Euphratica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Euphratica">Neritina Euphratica</taxonomicName>
<bibRefCitation author="Mousson, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="25" pagination="5 - 60" publicationUrl="https://biodiversitylibrary.org/page/15661760" refId="B111" refString="Mousson, A, 1874. Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient. Journal de Conchyliologie 22 (1): 5 - 60, https://biodiversitylibrary.org/page/15661760" title="Coquilles terrestres et fluviatiles recueillies par M. le Dr Alex. Schlaefli en Orient." url="https://biodiversitylibrary.org/page/15661760" volume="22" year="1874">Mousson 1874</bibRefCitation>
: 49; 
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 2, pl. 211, fig. 1318.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Recluz" authorityYear="1841" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina Anatolica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Anatolica">Neritina Anatolica</taxonomicName>
var. 
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="var. Mesopotamica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="Anatolica" variety="Mesopotamica">Mesopotamica</taxonomicName>
: 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Novitates Conchologicae, Supplement" pageId="0" pageNumber="25" pagination="1 - 127" publicationUrl="https://biodiversitylibrary.org/page/12980992" refId="B98" refString="Martens, Ev, 1874. Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates Conchologicae, Supplement 5: 1 - 127, https://biodiversitylibrary.org/page/12980992" title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht." url="https://biodiversitylibrary.org/page/12980992" volume="5" year="1874">Martens 1874</bibRefCitation>
: 33-34, pl. 5, fig. 42.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Martens" authorityYear="1874" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina cinctella" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="cinctella">Neritina cinctella</taxonomicName>
<bibRefCitation author="Martens, Ev" journalOrPublisher="Novitates Conchologicae, Supplement" pageId="0" pageNumber="25" pagination="1 - 127" publicationUrl="https://biodiversitylibrary.org/page/12980992" refId="B98" refString="Martens, Ev, 1874. Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates Conchologicae, Supplement 5: 1 - 127, https://biodiversitylibrary.org/page/12980992" title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht." url="https://biodiversitylibrary.org/page/12980992" volume="5" year="1874">Martens 1874</bibRefCitation>
: 34, pl. 5, fig. 43; 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 91, pl. 13, figs 22-24; 
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 4-5, pl. 211, fig. 1326.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Macrii [sic]: 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 88-90, pl. 4, figs 11-13, pl. 13, figs 27-29 (
<emphasis bold="true" pageId="0" pageNumber="25">non</emphasis>
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina macri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="macri">Neritina macri</taxonomicName>
G.B. Sowerby 
<collectionCode>II</collectionCode>
, 1849; partim, non pl. 13, fig. 13).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina Mesopotamica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Mesopotamica">Neritina Mesopotamica</taxonomicName>
: 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
: 90, pl. 13, figs 20, 21; 
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- &amp; Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
: 4, pl. 211, fig. 1325.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
(
<taxonomicName authority="Bourguignat, 1877" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Neritidae" genus="Theodoxia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theodoxia" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Theodoxia</taxonomicName>
) Jordani 
<taxonomicName authority="Bourguignat, 1877" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Neritidae" genus="Theodoxia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="var. aberrans" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="Mesopotamica" variety="aberrans">var. aberrans</taxonomicName>
<bibRefCitation author="Dautzenberg, P" journalOrPublisher="Revue biologique du nord de la France" pageId="0" pageNumber="25" pagination="329 - 353" publicationUrl="https://biodiversitylibrary.org/page/10877573" refId="B42" refString="Dautzenberg, P, 1894. Liste des mollusques terrestres et fluviatiles recueillis par M. Th. Barrois en Palestine et en Syrie. Revue biologique du nord de la France 6: 329 - 353, https://biodiversitylibrary.org/page/10877573" title="Liste des mollusques terrestres et fluviatiles recueillis par M. Th. Barrois en Palestine et en Syrie." url="https://biodiversitylibrary.org/page/10877573" volume="6" year="1894">Dautzenberg 1894</bibRefCitation>
: 351.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Orontis 
<bibRefCitation author="Blanckenhorn, M" journalOrPublisher="Palaeontographica" pageId="0" pageNumber="25" pagination="71 - 144" publicationUrl="https://biodiversitylibrary.org/page/33130542" refId="B25" refString="Blanckenhorn, M, 1897. Zur Kenntnis der Suesswasserablagerungen und Mollusken Syriens. Palaeontographica 44: 71 - 144, https://biodiversitylibrary.org/page/33130542" title="Zur Kenntnis der Suesswasserablagerungen und Mollusken Syriens." url="https://biodiversitylibrary.org/page/33130542" volume="44" year="1897">Blanckenhorn 1897</bibRefCitation>
: 101, pl. 8, figs 3-5.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authority="Bourguignat, 1877" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Neritidae" genus="Theodoxia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theodoxia" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Theodoxia</taxonomicName>
Macrii [sic]: 
<bibRefCitation author="Germain, L" journalOrPublisher="Bailliere et fils, Paris" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/13187112" refId="B59" refString="Germain, L, 1921. Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J.-B. Bailliere et fils, Paris, https://biodiversitylibrary.org/page/13187112" title="Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J. - B." url="https://biodiversitylibrary.org/page/13187112" year="1921">Germain 1921</bibRefCitation>
: 516-518 (
<emphasis bold="true" pageId="0" pageNumber="25">non</emphasis>
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina macri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="macri">Neritina macri</taxonomicName>
G.B. Sowerby 
<collectionCode>II</collectionCode>
, 1849).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authority="Bourguignat, 1877" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Neritidae" genus="Theodoxia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Theodoxia jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">Theodoxia jordani</taxonomicName>
: 
<bibRefCitation author="Germain, L" journalOrPublisher="Bailliere et fils, Paris" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/13187112" refId="B59" refString="Germain, L, 1921. Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J.-B. Bailliere et fils, Paris, https://biodiversitylibrary.org/page/13187112" title="Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J. - B." url="https://biodiversitylibrary.org/page/13187112" year="1921">Germain 1921</bibRefCitation>
: 511-514.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
Ponsoti 
<bibRefCitation author="Pallary, P" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle, deuxieme serie" pageId="0" pageNumber="25" pagination="286 - 290" publicationUrl="http://gallica.bnf.fr/ark:/12148/bpt6k5532113m/f44.image.r" refId="B117" refString="Pallary, P, 1930. Mollusques aquatiques nouveaux du Levant. Bulletin du Museum National d'Histoire Naturelle, deuxieme serie 2 (3): 286 - 290, http://gallica.bnf.fr/ark:/12148/bpt6k5532113m/f44.image.r" title="Mollusques aquatiques nouveaux du Levant." url="http://gallica.bnf.fr/ark:/12148/bpt6k5532113m/f44.image.r" volume="2" year="1930">Pallary 1930</bibRefCitation>
: 286-287, fig. 1.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani var. unicarinatus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="jordani" variety="unicarinatus">Theodoxus (Neritaea) jordani var. unicarinatus</taxonomicName>
<bibRefCitation author="Picard, L" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="25" pagination="105 - 139" publicationUrl="http://www.zobodat.at/pdf/Archiv-fuer-Molluskenkunde_66_0105-0139.pdf" refId="B122" refString="Picard, L, 1934. Mollusken der levantinischen Stufe Nordpalaestinas (Jordantal). Archiv fuer Molluskenkunde 66 (3): 105 - 139, http://www.zobodat.at/pdf/Archiv-fuer-Molluskenkunde_66_0105-0139.pdf" title="Mollusken der levantinischen Stufe Nordpalaestinas (Jordantal)." url="http://www.zobodat.at/pdf/Archiv-fuer-Molluskenkunde_66_0105-0139.pdf" volume="66" year="1934">Picard 1934</bibRefCitation>
: 107-111, pl. 7, figs 1-4.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani var. bicarinatus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="jordani" variety="bicarinatus">Theodoxus (Neritaea) jordani var. bicarinatus</taxonomicName>
<bibRefCitation author="Picard, L" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="25" pagination="105 - 139" publicationUrl="http://www.zobodat.at/pdf/Archiv-fuer-Molluskenkunde_66_0105-0139.pdf" refId="B122" refString="Picard, L, 1934. Mollusken der levantinischen Stufe Nordpalaestinas (Jordantal). Archiv fuer Molluskenkunde 66 (3): 105 - 139, http://www.zobodat.at/pdf/Archiv-fuer-Molluskenkunde_66_0105-0139.pdf" title="Mollusken der levantinischen Stufe Nordpalaestinas (Jordantal)." url="http://www.zobodat.at/pdf/Archiv-fuer-Molluskenkunde_66_0105-0139.pdf" volume="66" year="1934">Picard 1934</bibRefCitation>
: 111-112, pl. 7, figs 5-8.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
(
<taxonomicName authority="Roth, 1855" authorityName="Roth" authorityYear="1855" class="Gastropoda" family="Neritidae" genus="Neritaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neritaea" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritaea</taxonomicName>
) Gombaulti 
<bibRefCitation author="Pallary, P" journalOrPublisher="Memoires de l'Institut d'Egypte" pageId="0" pageNumber="25" pagination="1 - 141" refId="B118" refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141" title="Deuxieme addition a la faune malacologique de la Syrie." volume="39" year="1939">Pallary 1939</bibRefCitation>
: 107, pl. 4, figs 53-56.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina homsensis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="homsensis">Neritina (Neritaea) homsensis</taxonomicName>
<bibRefCitation author="Pallary, P" journalOrPublisher="Memoires de l'Institut d'Egypte" pageId="0" pageNumber="25" pagination="1 - 141" refId="B118" refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141" title="Deuxieme addition a la faune malacologique de la Syrie." volume="39" year="1939">Pallary 1939</bibRefCitation>
: 108, pl. 4, figs 57-61.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina homsensis var. major" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="homsensis" variety="major">Neritina (Neritaea) homsensis var. major</taxonomicName>
<bibRefCitation author="Pallary, P" journalOrPublisher="Memoires de l'Institut d'Egypte" pageId="0" pageNumber="25" pagination="1 - 141" refId="B118" refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141" title="Deuxieme addition a la faune malacologique de la Syrie." volume="39" year="1939">Pallary 1939</bibRefCitation>
: 109 (
<emphasis bold="true" pageId="0" pageNumber="25">non</emphasis>
<taxonomicName authorityName="Recluz" authorityYear="1850" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina wallisiarum var. major" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="wallisiarum" variety="major">Neritina wallisiarum var. major</taxonomicName>
<normalizedToken originalValue="Récluz">Recluz</normalizedToken>
, 1850).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina homsensis var. minor" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="homsensis" variety="minor">Neritina (Neritaea) homsensis var. minor</taxonomicName>
<bibRefCitation author="Pallary, P" journalOrPublisher="Memoires de l'Institut d'Egypte" pageId="0" pageNumber="25" pagination="1 - 141" refId="B118" refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141" title="Deuxieme addition a la faune malacologique de la Syrie." volume="39" year="1939">Pallary 1939</bibRefCitation>
: 109 (nomen nudum, 
<emphasis bold="true" pageId="0" pageNumber="25">non</emphasis>
<taxonomicName authorityName="Metcalf" authorityYear="1904" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina minor" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="minor">Neritina minor</taxonomicName>
Menke, 1828).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
(
<taxonomicName authority="Roth, 1855" authorityName="Roth" authorityYear="1855" class="Gastropoda" family="Neritidae" genus="Neritaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neritaea" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritaea</taxonomicName>
) Ponsoti: 
<bibRefCitation author="Pallary, P" journalOrPublisher="Memoires de l'Institut d'Egypte" pageId="0" pageNumber="25" pagination="1 - 141" refId="B118" refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141" title="Deuxieme addition a la faune malacologique de la Syrie." volume="39" year="1939">Pallary 1939</bibRefCitation>
: 109-110, pl. 4, figs 44-46.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName baseAuthorityName="G. B. Sowerby I" baseAuthorityYear="1836" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">Theodoxus (Neritaea) jordani</taxonomicName>
: 
<bibRefCitation author="Tchernov, E" journalOrPublisher="Malacologia" pageId="0" pageNumber="25" pagination="147 - 184" publicationUrl="https://biodiversitylibrary.org/page/13122806" refId="B154" refString="Tchernov, E, 1975. The molluscs of the Sea of Galilee. Malacologia 15 (1): 147 - 184, https://biodiversitylibrary.org/page/13122806" title="The molluscs of the Sea of Galilee." url="https://biodiversitylibrary.org/page/13122806" volume="15" year="1975">Tchernov 1975</bibRefCitation>
: 153; 
<bibRefCitation DOI="https://doi.org/10.1080/09397140.1993.10637638" author="Schuett, H" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" pagination="69 - 112" refId="B143" refString="Schuett, H, Ortal, R, 1993. A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria). Zoology in the Middle East 8 (1): 69 - 112, DOI: https://doi.org/10.1080/09397140.1993.10637638" title="A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria)." url="https://doi.org/10.1080/09397140.1993.10637638" volume="8" year="1993">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
and Ortal 1993
</bibRefCitation>
: 78-79, pl. 3, figs 37-41; 
<bibRefCitation author="Bandel, K" journalOrPublisher="Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg" pageId="0" pageNumber="25" pagination="65 - 164" refId="B20" refString="Bandel, K, 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg 85: 65 - 164" title="The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)." volume="85" year="2001">Bandel 2001</bibRefCitation>
: 84-86, figs 12-20, 32-40; 
<bibRefCitation author="Amr, ZS" journalOrPublisher="Denisia" pageId="0" pageNumber="25" pagination="221 - 227" publicationUrl="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" refId="B10" refString="Amr, ZS, Abu Baker, M, 2004. Freshwater snails of Jordan. Denisia 14: 221 - 227, http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" title="Freshwater snails of Jordan." url="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" volume="14" year="2004">Amr and Abu Baker 2004</bibRefCitation>
: 221-222, fig. 1.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus cinctella" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="cinctella">Theodoxus (Neritaea) cinctella</taxonomicName>
: 
<bibRefCitation DOI="https://doi.org/10.1080/09397140.1989.10637575" author="Schuett, H" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" pagination="55 - 58" refId="B144" refString="Schuett, H, Sesen, R, 1989a. The freshwater molluscs of Ceylanpinar. Zoology in the Middle East 3 (1): 55 - 58, DOI: https://doi.org/10.1080/09397140.1989.10637575" title="The freshwater molluscs of Ceylanpinar." url="https://doi.org/10.1080/09397140.1989.10637575" volume="3" year="1989 a">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
and 
<normalizedToken originalValue="Şeşen">Sesen</normalizedToken>
1989a
</bibRefCitation>
: 56-57.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1874" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus cinctellus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="cinctellus">Theodoxus (Neritaea) cinctellus</taxonomicName>
: 
<bibRefCitation author="Schuett, H" journalOrPublisher="Basteria" pageId="0" pageNumber="25" pagination="39 - 46" refId="B145" refString="Schuett, H, Sesen, R, 1989b. Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae). Basteria 53 (1-3): 39 - 46" title="Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae)." volume="53" year="1989 b">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
and 
<normalizedToken originalValue="Şeşen">Sesen</normalizedToken>
1989b
</bibRefCitation>
: 45-46; 
<bibRefCitation author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="877 - 900" refId="B161" refString="Yildirim, MZ, 1999. Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular. Turkish Journal of Zoology 23 (3): 877 - 900" title="Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular." volume="23" year="1999">
<normalizedToken originalValue="Yıldırım">Yildirim</normalizedToken>
1999
</bibRefCitation>
: 886.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authority="tricarinatus" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">Theodoxus (Neritaea) jordani tricarinatus</taxonomicName>
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
in 
<bibRefCitation DOI="https://doi.org/10.1080/09397140.1993.10637638" author="Schuett, H" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" pagination="69 - 112" refId="B143" refString="Schuett, H, Ortal, R, 1993. A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria). Zoology in the Middle East 8 (1): 69 - 112, DOI: https://doi.org/10.1080/09397140.1993.10637638" title="A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria)." url="https://doi.org/10.1080/09397140.1993.10637638" volume="8" year="1993">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
and Ortal 1993
</bibRefCitation>
: 79, pl. 3, fig. 42.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus pliocostulatus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="pliocostulatus">Theodoxus (Neritaea) pliocostulatus</taxonomicName>
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
in 
<bibRefCitation DOI="https://doi.org/10.1080/09397140.1993.10637638" author="Schuett, H" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" pagination="69 - 112" refId="B143" refString="Schuett, H, Ortal, R, 1993. A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria). Zoology in the Middle East 8 (1): 69 - 112, DOI: https://doi.org/10.1080/09397140.1993.10637638" title="A preliminary correlation between the Plio-Pleistocene malacofaunas of the Jordan Valley (Israel) and the Orontes Valley (Syria)." url="https://doi.org/10.1080/09397140.1993.10637638" volume="8" year="1993">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
and Ortal 1993
</bibRefCitation>
: 79-80, pl. 3, fig. 43.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName baseAuthorityName="Reeve" baseAuthorityYear="1865" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus niloticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="niloticus">Theodoxus niloticus</taxonomicName>
: 
<bibRefCitation author="Brown, DS" journalOrPublisher="Taylor &amp; Francis, London" pageId="0" pageNumber="25" refId="B33" refString="Brown, DS, 1994. Freshwater snails of Africa and their medical importance. Taylor &amp; Francis, London" title="Freshwater snails of Africa and their medical importance." year="1994">Brown 1994</bibRefCitation>
: 45, figs 16b-c.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">Theodoxus jordani</taxonomicName>
: 
<bibRefCitation author="Yildirim, MZ" journalOrPublisher="Turkish Journal of Zoology" pageId="0" pageNumber="25" pagination="877 - 900" refId="B161" refString="Yildirim, MZ, 1999. Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular. Turkish Journal of Zoology 23 (3): 877 - 900" title="Tuerkiye Prosobranchia (Gastropoda: Mollusca) tuerleri ve zoocografik yayilislari. 1. Tatli ve aci sular." volume="23" year="1999">
<normalizedToken originalValue="Yıldırım">Yildirim</normalizedToken>
1999
</bibRefCitation>
: 886; 
<bibRefCitation author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
: 2993; 
<bibRefCitation author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
: 44, figs 30.
</paragraph>
<paragraph pageId="0" pageNumber="25">
non 
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina mesopotamica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="mesopotamica">Neritina mesopotamica</taxonomicName>
: 
<bibRefCitation author="Mansoorian, AB" journalOrPublisher="Iranian International Journal of Science" pageId="0" pageNumber="25" pagination="96 - 103" refId="B95" refString="Mansoorian, AB, 2001. Freshwater Gastropod of Khuzestan Province, South-West Iran. Iranian International Journal of Science 2 (2): 96 - 103" title="Freshwater Gastropod of Khuzestan Province, South-West Iran." volume="2" year="2001">Mansoorian 2001</bibRefCitation>
: 4, figs 1-4 (= 
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina schlaeflii" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="schlaeflii">Neritina schlaeflii</taxonomicName>
Mousson, 1874).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName baseAuthorityName="Sowerby" baseAuthorityYear="1849" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus macrii" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="macrii">Theodoxus macrii</taxonomicName>
[sic]: 
<bibRefCitation author="Amr, ZS" journalOrPublisher="Denisia" pageId="0" pageNumber="25" pagination="221 - 227" publicationUrl="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" refId="B10" refString="Amr, ZS, Abu Baker, M, 2004. Freshwater snails of Jordan. Denisia 14: 221 - 227, http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" title="Freshwater snails of Jordan." url="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" volume="14" year="2004">Amr and Abu Baker 2004</bibRefCitation>
: 222, fig. 2; 
<bibRefCitation author="Handal, EN" journalOrPublisher="Jordan Journal of Natural History" pageId="0" pageNumber="25" pagination="23 - 29" refId="B70" refString="Handal, EN, Amr, Z, Qumsiyeh, MB, 2015. Some records of freshwater snail from the Occupied Palestinian territories. Jordan Journal of Natural History 1 (2): 23 - 29" title="Some records of freshwater snail from the Occupied Palestinian territories." volume="1" year="2015">Handal et al. 2015</bibRefCitation>
: 25-26, fig. 1B (
<emphasis bold="true" pageId="0" pageNumber="25">non</emphasis>
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina macri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="macri">Neritina macri</taxonomicName>
G.B. Sowerby 
<collectionCode>II</collectionCode>
, 1849).
</paragraph>
<paragraph pageId="0" pageNumber="25">
? 
<taxonomicName authorityName="Martens" authorityYear="1874" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina cinctellus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="cinctellus">Neritina cinctellus</taxonomicName>
: 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
and 
<normalizedToken originalValue="Pešić">Pesic</normalizedToken>
2012
</bibRefCitation>
: 13-14, fig. 2a, c.
</paragraph>
<paragraph pageId="0" pageNumber="25">
? 
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina mesopotamica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="mesopotamica">Neritina mesopotamica</taxonomicName>
: 
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
and 
<normalizedToken originalValue="Pešić">Pesic</normalizedToken>
2012
</bibRefCitation>
: 14-16.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName baseAuthorityName="Mousson" baseAuthorityYear="1874" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus euphraticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euphraticus">Theodoxus euphraticus</taxonomicName>
: 
<bibRefCitation author="Mansoorian, AB" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B96" refString="Mansoorian, AB, Samaee, A, 2012. Gastropoda of the freshwaters of Iran. Privately published, Tehran" title="Gastropoda of the freshwaters of Iran." year="2012">Mansoorian and Samaee 2012</bibRefCitation>
: 50-57 (partim, only material from Chaharmahal and Bakhtiari, Fars, Kermanshah, and Khuzestan provinces).
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Mousson" authorityYear="1874" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina euphratica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="euphratica">Neritina euphratica</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
and 
<normalizedToken originalValue="Pešić">Pesic</normalizedToken>
2012
</bibRefCitation>
: 16, fig. 2d, e.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">Theodoxus cf. jordani</taxonomicName>
: 
<bibRefCitation author="Alhejoj, I" journalOrPublisher="Freiberger Forschungshefte: Palaeontologie, Stratigraphie, Fazies C" pageId="0" pageNumber="25" pagination="141 - 173" refId="B7" refString="Alhejoj, I, Bandel, K, 2013. Mollusks of the Pleistocene Al-Qarn Formation of the Jordan Rift-Valley in Jordan. Freiberger Forschungshefte: Palaeontologie, Stratigraphie, Fazies C 545 (21): 141 - 173" title="Mollusks of the Pleistocene Al-Qarn Formation of the Jordan Rift-Valley in Jordan." volume="545" year="2013">Alhejoj and Bandel 2013</bibRefCitation>
: 146-147, pl. 1, figs 5-8.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Eichhorst" authorityYear="2016" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus octagonus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="octagonus">Theodoxus octagonus</taxonomicName>
<bibRefCitation author="Eichhorst, TE" journalOrPublisher="ConchBooks, Hackenheim, Germany" pageId="0" pageNumber="25" refId="B48" refString="Eichhorst, TE, 2016. Neritidae of the World. Vol. 2. ConchBooks, Hackenheim, Germany" title="Neritidae of the World. Vol. 2." year="2016">Eichhorst 2016</bibRefCitation>
: 940, pl. 293, figs 1-8.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName authorityName="Eichhorst" authorityYear="2016" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus octagonus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="octagonus">Theodoxus (Neritaea) octagonus</taxonomicName>
: 
<bibRefCitation author="Mienis, HK" journalOrPublisher="Ellipsaria" pageId="0" pageNumber="25" pagination="37 - 38" refId="B102" refString="Mienis, HK, Rittner, O, 2017. Some information concerning Theodoxus (Neritaea) octagonus Eichhorst, 2016, an endemic subterranean species from Israel. Ellipsaria 19 (2): 37 - 38" title="Some information concerning Theodoxus (Neritaea) octagonus Eichhorst, 2016, an endemic subterranean species from Israel." volume="19" year="2017">Mienis and Rittner 2017</bibRefCitation>
: 37, figs 1-3.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus mesopotamicus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="mesopotamicus">Theodoxus mesopotamicus</taxonomicName>
: 
<bibRefCitation author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
2019
</bibRefCitation>
: 47, fig. 34.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName baseAuthorityName="Martens" baseAuthorityYear="1874" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus cinctellus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="cinctellus">Theodoxus cinctellus</taxonomicName>
: 
<bibRefCitation author="Guerlek, ME" journalOrPublisher="Fresenius Enviromental Bulletin" pageId="0" pageNumber="25" pagination="2992 - 3013" refId="B69" refString="Guerlek, ME, Sahin, SK, Doekuemcue, N, Yildirim, MZ, 2019. Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia). Fresenius Enviromental Bulletin 28 (4): 2992 - 3013" title="Checklist of the freshwater mollusca of Turkey (Mollusca: Gastropoda, Bivalvia)." volume="28" year="2019">
<normalizedToken originalValue="Gürlek">Guerlek</normalizedToken>
et al. 2019
</bibRefCitation>
: 2993.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="type locality">
<paragraph pageId="0" pageNumber="25">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="25">River Jordan.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="type material">
<paragraph pageId="0" pageNumber="25">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="25">
<materialsCitation collectionCode="ZMZ" country="Turkey" typeStatus="type">
The 
<typeStatus>type</typeStatus>
material of 
<taxonomicName genus="N." lsidName="N. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">N. jordani</emphasis>
</taxonomicName>
could unfortunately not be traced. The 
<typeStatus>type</typeStatus>
material of the taxa introduced by Mousson are stored in 
<collectionCode collectionName="ZMZ">ZMZ</collectionCode>
, including 
<specimenCount type="generic">16 syntypes</specimenCount>
of 
<taxonomicName genus="N." lsidName="N. bellardii" pageId="0" pageNumber="25" rank="species" species="bellardii">
<emphasis italics="true" pageId="0" pageNumber="25">N. bellardii</emphasis>
</taxonomicName>
(coll. no. 528918)
</materialsCitation>
, 
<materialsCitation country="Turkey">
<specimenCount type="generic">13 syntypes</specimenCount>
of 
<taxonomicName genus="N." lsidName="N. euphratica" pageId="0" pageNumber="25" rank="species" species="euphratica">
<emphasis italics="true" pageId="0" pageNumber="25">N. euphratica</emphasis>
</taxonomicName>
(coll. no. 528916)
</materialsCitation>
, 
<materialsCitation country="Turkey">
<specimenCount type="generic">20 syntypes</specimenCount>
of 
<taxonomicName genus="N." lsidName="N. jordani var. turris" pageId="0" pageNumber="25" rank="variety" species="jordani" variety="turris">N. jordani var. turris</taxonomicName>
(coll. no. 528930)
</materialsCitation>
, 
<materialsCitation country="Turkey">
<specimenCount type="generic">5 syntypes</specimenCount>
of 
<taxonomicName genus="N." lsidName="N. karasuna" pageId="0" pageNumber="25" rank="species" species="karasuna">
<emphasis italics="true" pageId="0" pageNumber="25">N. karasuna</emphasis>
</taxonomicName>
(coll. no. 528937)
</materialsCitation>
, 
<materialsCitation country="Turkey">
and 
<specimenCount type="generic">56 syntypes</specimenCount>
of 
<taxonomicName genus="N." lsidName="N. meridionalis var. mesopotamica" pageId="0" pageNumber="25" rank="variety" species="meridionalis" variety="mesopotamica">N. meridionalis var. mesopotamica</taxonomicName>
(coll. no. 528912-528914).
</materialsCitation>
<materialsCitation collectionCode="ZMB" country="Turkey">
Additionally, 
<specimenCount type="generic">64 syntypes</specimenCount>
of 
<taxonomicName genus="N." lsidName="N. cinctella" pageId="0" pageNumber="25" rank="species" species="cinctella">
<emphasis italics="true" pageId="0" pageNumber="25">N. cinctella</emphasis>
</taxonomicName>
are stored in 
<collectionCode collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
(coll. no. 
<collectionCode collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
/Moll 21732, 21735, 66639)
</materialsCitation>
, 
<materialsCitation collectionCode="MHNG" country="Turkey">
<specimenCount type="generic">2 syntypes</specimenCount>
of 
<taxonomicName genus="N." lsidName="N. michonii" pageId="0" pageNumber="25" rank="species" species="michonii">
<emphasis italics="true" pageId="0" pageNumber="25">N. michonii</emphasis>
</taxonomicName>
are stored in 
<collectionCode collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
(MHNG-MOLL-111706-111707)
</materialsCitation>
, 
<materialsCitation collectionCode="MNHM" country="Turkey" typeStatus="paratype">
<specimenCount type="generic">2 syntypes</specimenCount>
of 
<taxonomicName genus="N." lsidName="N. ponsoti" pageId="0" pageNumber="25" rank="species" species="ponsoti">
<emphasis italics="true" pageId="0" pageNumber="25">N. ponsoti</emphasis>
</taxonomicName>
are stored in 
<collectionCode collectionName="United States, Colorado, Colorado Springs, John May Museum of Natural History" country="USA" httpUri="http://grbio.org/cool/q3kg-qwj4" name="John May Museum of Natural History" type="">MNHM</collectionCode>
(coll. no. MNHN-IM-2000-32754) and another one (labelled as 
<normalizedToken originalValue="“paratype”">
&quot;
<typeStatus>paratype</typeStatus>
&quot;
</normalizedToken>
) is stored in 
<collectionCode collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" type="Museum">RBINS</collectionCode>
(coll. no. 
<collectionCode country="Russia" httpUri="http://grbio.org/cool/etcv-5c5s" name="Mus. Tinro, Vladyvostok" type="">MT</collectionCode>
820)
</materialsCitation>
, 
<materialsCitation collectionCode="BMSM" country="Turkey" typeStatus="holotype">
and the 
<typeStatus>holotype</typeStatus>
and two 
<typeStatus>paratypes</typeStatus>
of 
<taxonomicName genus="T." lsidName="T. octagonus" pageId="0" pageNumber="25" rank="species" species="octagonus">
<emphasis italics="true" pageId="0" pageNumber="25">T. octagonus</emphasis>
</taxonomicName>
are stored in 
<collectionCode collectionName="BMSM">BMSM</collectionCode>
(coll. no. 
<collectionCode collectionName="BMSM">BMSM</collectionCode>
93544-93545)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="remarks">
<paragraph pageId="0" pageNumber="25">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="25">
A number of nominal species have already been synonymised under 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
based on similarities in shell morphology and overlapping distribution ranges. These include 
<taxonomicName genus="N." lsidName="N. michonii" pageId="0" pageNumber="25" rank="species" species="michonii">
<emphasis italics="true" pageId="0" pageNumber="25">N. michonii</emphasis>
</taxonomicName>
Bourguignat, 1852, 
<taxonomicName genus="N." lsidName="N. karasuna" pageId="0" pageNumber="25" rank="species" species="karasuna">
<emphasis italics="true" pageId="0" pageNumber="25">N. karasuna</emphasis>
</taxonomicName>
Mousson, 1874, and 
<taxonomicName genus="N." lsidName="N. orontis" pageId="0" pageNumber="25" rank="species" species="orontis">
<emphasis italics="true" pageId="0" pageNumber="25">N. orontis</emphasis>
</taxonomicName>
Blanckenhorn, 1897 (
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
, 
<bibRefCitation DOI="https://doi.org/10.1007/978-3-319-51856-5_17" author="Alhejoj, I" journalOrPublisher="Freiberger Forschungshefte: Palaeontologie, Stratigraphie, Fazies C" pageId="0" pageNumber="25" refId="B8" refString="Alhejoj, I, Bandel, K, Salameh, E, 2017. Aquatic mollusks: occurrences, identification and their use as bioindicators of environmental conditions (salinity, trace elements and pollution parameters) in Jordan. In Abdalla O, Kacimov A, Chen M, Al-Maktoumi A, Al-Hosni T and Clark I (Eds) Water resources in arid areas: the way forward. Springer, Cham, 295-318. https://doi.org/10.1007/978-3-319-51856-5_17" title="Aquatic mollusks: occurrences, identification and their use as bioindicators of environmental conditions (salinity, trace elements and pollution parameters) in Jordan. In Abdalla O, Kacimov A, Chen M, Al-Maktoumi A, Al-Hosni T and Clark I (Eds) Water resources in arid areas: the way forward. Springer, Cham, 295 - 318." url="https://doi.org/10.1007/978-3-319-51856-5_17" year="2017">Alhejoj et al. 2017</bibRefCitation>
). Moreover, 
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. (2019a)</bibRefCitation>
suggested that 
<taxonomicName genus="T." lsidName="T. niloticus" pageId="0" pageNumber="25" rank="species" species="niloticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. niloticus</emphasis>
</taxonomicName>
(Reeve, 1856) from the Nile River, Egypt (which has been considered a potential synonym of 
<taxonomicName genus="N." lsidName="N. africana" pageId="0" pageNumber="25" rank="species" species="africana">
<emphasis italics="true" pageId="0" pageNumber="25">N. africana</emphasis>
</taxonomicName>
Reeve, 1856; 
<bibRefCitation author="Brown, DS" journalOrPublisher="Taylor &amp; Francis, London" pageId="0" pageNumber="25" refId="B33" refString="Brown, DS, 1994. Freshwater snails of Africa and their medical importance. Taylor &amp; Francis, London" title="Freshwater snails of Africa and their medical importance." year="1994">Brown 1994</bibRefCitation>
) is conspecific with 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
on the basis of molecular data-something additionally supported by 
<bibRefCitation author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
(2019)
</bibRefCitation>
. Our phylogenetic analyses expand upon this; we show that 
<taxonomicName genus="T." lsidName="T. mesopotamicus" pageId="0" pageNumber="25" rank="species" species="mesopotamicus">
<emphasis italics="true" pageId="0" pageNumber="25">T. mesopotamicus</emphasis>
</taxonomicName>
(including the synonyms 
<taxonomicName genus="N." lsidName="N. cinctella" pageId="0" pageNumber="25" rank="species" species="cinctella">
<emphasis italics="true" pageId="0" pageNumber="25">N. cinctella</emphasis>
</taxonomicName>
Martens, 1874, 
<taxonomicName genus="N." lsidName="N. anatolica var. mesopotamica" pageId="0" pageNumber="25" rank="variety" species="anatolica" variety="mesopotamica">N. anatolica var. mesopotamica</taxonomicName>
Martens, 1874 (
<bibRefCitation DOI="https://doi.org/10.1080/09397140.1989.10637575" author="Schuett, H" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" pagination="55 - 58" refId="B144" refString="Schuett, H, Sesen, R, 1989a. The freshwater molluscs of Ceylanpinar. Zoology in the Middle East 3 (1): 55 - 58, DOI: https://doi.org/10.1080/09397140.1989.10637575" title="The freshwater molluscs of Ceylanpinar." url="https://doi.org/10.1080/09397140.1989.10637575" volume="3" year="1989 a">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
and 
<normalizedToken originalValue="Şeşen">Sesen</normalizedToken>
1989a
</bibRefCitation>
, 
<bibRefCitation author="Schuett, H" journalOrPublisher="Basteria" pageId="0" pageNumber="25" pagination="39 - 46" refId="B145" refString="Schuett, H, Sesen, R, 1989b. Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae). Basteria 53 (1-3): 39 - 46" title="Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae)." volume="53" year="1989 b">1989b</bibRefCitation>
), and 
<taxonomicName genus="T." lsidName="T. euphraticus" pageId="0" pageNumber="25" rank="species" species="euphraticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. euphraticus</emphasis>
</taxonomicName>
should be considered conspecific with 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
). These results support the earlier notion of 
<bibRefCitation author="Roth, G" editor="Krupp, F" journalOrPublisher="Dr. ludwig Reichert Verlag, Wiesbaden, Germany" pageId="0" pageNumber="25" pagination="72 - 79" refId="B136" refString="Roth, G, 1987. Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae). In: Krupp, F, Schneider, W, Kinzelbach, R, Eds., Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East. Dr. ludwig Reichert Verlag, Wiesbaden, Germany: 72 - 79" title="Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae)." volumeTitle="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East." year="1987">Roth (1987)</bibRefCitation>
, who suggested that 
<taxonomicName genus="T." lsidName="T. euphraticus" pageId="0" pageNumber="25" rank="species" species="euphraticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. euphraticus</emphasis>
</taxonomicName>
, 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
, 
<taxonomicName genus="T." lsidName="T. mesopotamicus" pageId="0" pageNumber="25" rank="species" species="mesopotamicus">
<emphasis italics="true" pageId="0" pageNumber="25">T. mesopotamicus</emphasis>
</taxonomicName>
, and 
<taxonomicName genus="T." lsidName="T. niloticus" pageId="0" pageNumber="25" rank="species" species="niloticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. niloticus</emphasis>
</taxonomicName>
may be conspecific based on morphological similarities, particularly concerning the opercula (Figs 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure15" httpUri="https://binary.pensoft.net/fig/376464" pageId="0" pageNumber="25">15</figureCitation>
- 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure17" httpUri="https://binary.pensoft.net/fig/376466" pageId="0" pageNumber="25">17</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.96.48312.figure15" httpUri="https://binary.pensoft.net/fig/376464" pageId="0" pageNumber="25" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 15.</emphasis>
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus jordani</emphasis>
</taxonomicName>
(G.B. Sowerby I, 1836). 
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
. Specimen from the Avakas Gorge, Peyia, Cyprus (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
18321) incorporated into the phylogeny (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
); 
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">H</emphasis>
. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
20777); 
<emphasis bold="true" pageId="0" pageNumber="25">I</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">L</emphasis>
. Specimen from the Orontes River, Antakya, Hatay, Turkey (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
24175) used in the phylogeny (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
); 
<emphasis bold="true" pageId="0" pageNumber="25">M</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">O</emphasis>
. Specimen (
<taxonomicName genus="T." lsidName="T. euphraticus" pageId="0" pageNumber="25" rank="species" species="euphraticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. euphraticus</emphasis>
</taxonomicName>
-morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
22233); 
<emphasis bold="true" pageId="0" pageNumber="25">P</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">R</emphasis>
. Topotype of 
<taxonomicName genus="N." lsidName="N. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">N. jordani</emphasis>
</taxonomicName>
from the River Jordan, Israel (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
24177) incorporated into the phylogeny (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
); 
<emphasis bold="true" pageId="0" pageNumber="25">S</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="25">T</emphasis>
. Specimen (
<taxonomicName genus="T." lsidName="T. euphraticus" pageId="0" pageNumber="25" rank="species" species="euphraticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. euphraticus</emphasis>
</taxonomicName>
-morphotype) from the Karun River, Ahvaz, Iran (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
21681); 
<emphasis bold="true" pageId="0" pageNumber="25">U</emphasis>
. Specimen (
<taxonomicName genus="T." lsidName="T. niloticus" pageId="0" pageNumber="25" rank="species" species="niloticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. niloticus</emphasis>
</taxonomicName>
-morphotype) collected in the Nile River, Egypt (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
24178) used in the phylogeny (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
); 
<emphasis bold="true" pageId="0" pageNumber="25">V</emphasis>
. Specimen (
<taxonomicName genus="T." lsidName="T. euphraticus" pageId="0" pageNumber="25" rank="species" species="euphraticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. euphraticus</emphasis>
</taxonomicName>
-morphotype) from the Karun River, Ahvaz, Iran (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
21682). Scale bars: 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<caption doi="10.3897/zse.96.48312.figure16" httpUri="https://binary.pensoft.net/fig/376465" pageId="0" pageNumber="25" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 16.</emphasis>
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus jordani</emphasis>
</taxonomicName>
(G.B. Sowerby I, 1836). 
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="25">B</emphasis>
. 
<typeStatus>Syntype</typeStatus>
of 
<taxonomicName genus="N." lsidName="N. euphratica" pageId="0" pageNumber="25" rank="species" species="euphratica">
<emphasis italics="true" pageId="0" pageNumber="25">N. euphratica</emphasis>
</taxonomicName>
from the Euphrates River at Samawah, Iraq (
<collectionCode collectionName="ZMZ">ZMZ</collectionCode>
528916); 
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
. 
<typeStatus>Syntype</typeStatus>
of 
<taxonomicName genus="N." lsidName="N. michonii" pageId="0" pageNumber="25" rank="species" species="michonii">
<emphasis italics="true" pageId="0" pageNumber="25">N. michonii</emphasis>
</taxonomicName>
from Syria (MHNG-MOLL-111706); 
<emphasis bold="true" pageId="0" pageNumber="25">F</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">H</emphasis>
. 
<typeStatus>Syntype</typeStatus>
of 
<taxonomicName genus="N." lsidName="N. karasuna" pageId="0" pageNumber="25" rank="species" species="karasuna">
<emphasis italics="true" pageId="0" pageNumber="25">N. karasuna</emphasis>
</taxonomicName>
collected from the Karasu River, south-eastern Turkey (
<collectionCode collectionName="ZMZ">ZMZ</collectionCode>
528937); 
<emphasis bold="true" pageId="0" pageNumber="25">I</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="25">J</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="25">K</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="25">L</emphasis>
. Two 
<typeStatus>syntypes</typeStatus>
of 
<taxonomicName genus="N." lsidName="N. ponsoti" pageId="0" pageNumber="25" rank="species" species="ponsoti">
<emphasis italics="true" pageId="0" pageNumber="25">N. ponsoti</emphasis>
</taxonomicName>
from Lake 
<normalizedToken originalValue="Muzayrīb">Muzayrīb</normalizedToken>
, Syria (MNHN-IM-2000-32754); 
<emphasis bold="true" pageId="0" pageNumber="25">M</emphasis>
. 
<typeStatus>Syntype</typeStatus>
(labelled as 
<typeStatus>paratype</typeStatus>
) of 
<taxonomicName genus="N." lsidName="N. ponsoti" pageId="0" pageNumber="25" rank="species" species="ponsoti">
<emphasis italics="true" pageId="0" pageNumber="25">N. ponsoti</emphasis>
</taxonomicName>
from Lake 
<normalizedToken originalValue="Muzayrīb">Muzayrīb</normalizedToken>
(
<collectionCode country="Russia" httpUri="http://grbio.org/cool/etcv-5c5s" name="Mus. Tinro, Vladyvostok" type="">MT</collectionCode>
820, 
<collectionCode collectionName="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" type="Museum">RBINS</collectionCode>
); 
<emphasis bold="true" pageId="0" pageNumber="25">N</emphasis>
. 
<typeStatus>Syntype</typeStatus>
of 
<taxonomicName genus="N." lsidName="N. jordani var. turris" pageId="0" pageNumber="25" rank="variety" species="jordani" variety="turris">N. jordani var. turris</taxonomicName>
collected in the Sea of Galilee (
<collectionCode collectionName="ZMZ">ZMZ</collectionCode>
528930); 
<emphasis bold="true" pageId="0" pageNumber="25">O</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="25">P</emphasis>
. 
<typeStatus>Syntype</typeStatus>
of 
<taxonomicName genus="N." lsidName="N. bellardii" pageId="0" pageNumber="25" rank="species" species="bellardii">
<emphasis italics="true" pageId="0" pageNumber="25">N. bellardii</emphasis>
</taxonomicName>
from the Litani River, Beqaa valley, Lebanon (
<collectionCode collectionName="ZMZ">ZMZ</collectionCode>
528918). Scale bars: 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<caption doi="10.3897/zse.96.48312.figure17" httpUri="https://binary.pensoft.net/fig/376466" pageId="0" pageNumber="25" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figure 17.</emphasis>
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus jordani</emphasis>
</taxonomicName>
(G.B. Sowerby I, 1836). 
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
. 
<typeStatus>Syntype</typeStatus>
of 
<taxonomicName genus="N." lsidName="N. meridionalis var. mesopotamica" pageId="0" pageNumber="25" rank="variety" species="meridionalis" variety="mesopotamica">N. meridionalis var. mesopotamica</taxonomicName>
from near 
<normalizedToken originalValue="Diyarbakır">Diyarbakir</normalizedToken>
, Turkey (
<collectionCode collectionName="ZMZ">ZMZ</collectionCode>
528914); 
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
. Specimen (
<taxonomicName genus="T." lsidName="T. mesopotamicus" pageId="0" pageNumber="25" rank="species" species="mesopotamicus">
<emphasis italics="true" pageId="0" pageNumber="25">T. mesopotamicus</emphasis>
</taxonomicName>
-morphotype) collected in 
<normalizedToken originalValue="Şanlıurfa">Sanliurfa</normalizedToken>
, Turkey (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
23426) and used in the phylogeny (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
); 
<emphasis bold="true" pageId="0" pageNumber="25">F</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">H</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="25">K</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="25">M</emphasis>
. Two 
<typeStatus>syntypes</typeStatus>
of 
<taxonomicName genus="N." lsidName="N. cinctella" pageId="0" pageNumber="25" rank="species" species="cinctella">
<emphasis italics="true" pageId="0" pageNumber="25">N. cinctella</emphasis>
</taxonomicName>
collected at the source of River Chabur near Ras al-Ayn, Syria (
<collectionCode collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
/Moll 21732); 
<emphasis bold="true" pageId="0" pageNumber="25">I</emphasis>
, 
<emphasis bold="true" pageId="0" pageNumber="25">J</emphasis>
. Topotype of 
<taxonomicName genus="N." lsidName="N. cinctella" pageId="0" pageNumber="25" rank="species" species="cinctella">
<emphasis italics="true" pageId="0" pageNumber="25">N. cinctella</emphasis>
</taxonomicName>
from the same locality (
<collectionCode collectionName="UGSB">UGSB</collectionCode>
18762). Scale bars: 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="25">
Although no molecular data could be incorporated, we further synonymise 
<taxonomicName genus="T." lsidName="T. octagonus" pageId="0" pageNumber="25" rank="species" species="octagonus">
<emphasis italics="true" pageId="0" pageNumber="25">T. octagonus</emphasis>
</taxonomicName>
Eichhorst, 2016 with 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
. While 
<taxonomicName genus="T." lsidName="T. octagonus" pageId="0" pageNumber="25" rank="species" species="octagonus">
<emphasis italics="true" pageId="0" pageNumber="25">T. octagonus</emphasis>
</taxonomicName>
maintains a subterranean lifestyle, the species shares a number of characteristics with 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
besides overlapping ranges; the operculum structures of the two species are near identical with otherwise large pseudo-apophyses and both occasionally show keeling of the shell. Moreover, key characters used to distinguish 
<taxonomicName genus="T." lsidName="T. octagonus" pageId="0" pageNumber="25" rank="species" species="octagonus">
<emphasis italics="true" pageId="0" pageNumber="25">T. octagonus</emphasis>
</taxonomicName>
, such as the pronounced aperture and colouration of the shells, may be mitigated when intraspecific phenotypic plasticity of both species is considered (
<bibRefCitation DOI="https://doi.org/10.1007/BF00346406" author="Heller, J" journalOrPublisher="Oecologia" pageId="0" pageNumber="25" pagination="98 - 104" refId="B71" refString="Heller, J, 1979. Visual versus non-visual selection of shell colour in an Israeli freshwater snail. Oecologia 44 (1): 98 - 104, DOI: https://doi.org/10.1007/BF00346406" title="Visual versus non-visual selection of shell colour in an Israeli freshwater snail." url="https://doi.org/10.1007/BF00346406" volume="44" year="1979">Heller 1979</bibRefCitation>
; 
<bibRefCitation author="Bandel, K" journalOrPublisher="Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg" pageId="0" pageNumber="25" pagination="65 - 164" refId="B20" refString="Bandel, K, 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg 85: 65 - 164" title="The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)." volume="85" year="2001">Bandel 2001</bibRefCitation>
; 
<bibRefCitation author="Amr, Z" journalOrPublisher="Jordan Journal of Natural History" pageId="0" pageNumber="25" pagination="83 - 115" refId="B11" refString="Amr, Z, Nasarat, H, Neubert, E, 2014. Notes on the current and past freshwater snail fauna of Jordan. Jordan Journal of Natural History 1: 83 - 115" title="Notes on the current and past freshwater snail fauna of Jordan." volume="1" year="2014">Amr et al. 2014</bibRefCitation>
; 
<bibRefCitation author="Eichhorst, TE" journalOrPublisher="ConchBooks, Hackenheim, Germany" pageId="0" pageNumber="25" refId="B48" refString="Eichhorst, TE, 2016. Neritidae of the World. Vol. 2. ConchBooks, Hackenheim, Germany" title="Neritidae of the World. Vol. 2." year="2016">Eichhorst 2016</bibRefCitation>
; 
<bibRefCitation author="Mienis, HK" journalOrPublisher="Ellipsaria" pageId="0" pageNumber="25" pagination="37 - 38" refId="B102" refString="Mienis, HK, Rittner, O, 2017. Some information concerning Theodoxus (Neritaea) octagonus Eichhorst, 2016, an endemic subterranean species from Israel. Ellipsaria 19 (2): 37 - 38" title="Some information concerning Theodoxus (Neritaea) octagonus Eichhorst, 2016, an endemic subterranean species from Israel." volume="19" year="2017">Mienis and Rittner 2017</bibRefCitation>
; Figs 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure15" httpUri="https://binary.pensoft.net/fig/376464" pageId="0" pageNumber="25">15</figureCitation>
- 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure17" httpUri="https://binary.pensoft.net/fig/376466" pageId="0" pageNumber="25">17</figureCitation>
). Likewise, the taxa 
<taxonomicName genus="N." lsidName="N. bellardii" pageId="0" pageNumber="25" rank="species" species="bellardii">
<emphasis italics="true" pageId="0" pageNumber="25">N. bellardii</emphasis>
</taxonomicName>
Mousson, 1854, 
<taxonomicName genus="N." lsidName="N. jordani var. turris" pageId="0" pageNumber="25" rank="variety" species="jordani" variety="turris">N. jordani var. turris</taxonomicName>
Mousson, 1861, 
<taxonomicName genus="N." lsidName="N. ponsoti" pageId="0" pageNumber="25" rank="species" species="ponsoti">
<emphasis italics="true" pageId="0" pageNumber="25">N. ponsoti</emphasis>
</taxonomicName>
Pallary, 1930, 
<taxonomicName genus="N." lsidName="N. homsensis" pageId="0" pageNumber="25" rank="species" species="homsensis">
<emphasis italics="true" pageId="0" pageNumber="25">N. homsensis</emphasis>
</taxonomicName>
Pallary, 1939, 
<taxonomicName genus="T." lsidName="T. jordani var. unicarinatus" pageId="0" pageNumber="25" rank="variety" species="jordani" variety="unicarinatus">T. jordani var. unicarinatus</taxonomicName>
Picard, 1934, 
<taxonomicName genus="T." lsidName="T. jordani var. bicarinatus" pageId="0" pageNumber="25" rank="variety" species="jordani" variety="bicarinatus">T. jordani var. bicarinatus</taxonomicName>
Picard, 1934, 
<taxonomicName genus="N." lsidName="N. gombaulti" pageId="0" pageNumber="25" rank="species" species="gombaulti">
<emphasis italics="true" pageId="0" pageNumber="25">N. gombaulti</emphasis>
</taxonomicName>
Pallary, 1939, 
<taxonomicName authority="tricarinatus" genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani tricarinatus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
in 
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
&amp; Ortal, 1993, and 
<taxonomicName genus="T." lsidName="T. pliocostulatus" pageId="0" pageNumber="25" rank="species" species="pliocostulatus">
<emphasis italics="true" pageId="0" pageNumber="25">T. pliocostulatus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
in 
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
&amp; Ortal, 1993 are well within the variability of 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
and are considered junior synonyms herein as well.
</paragraph>
<paragraph pageId="0" pageNumber="25">
Regarding the phylogenetic placement of 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
, 
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2006.06.026" author="Bunje, PME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="25" pagination="373 - 387" refId="B35" refString="Bunje, PME, Lindberg, DR, 2007. Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development. Molecular Phylogenetics and Evolution 42 (2): 373 - 387, DOI: https://doi.org/10.1016/j.ympev.2006.06.026" title="Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development." url="https://doi.org/10.1016/j.ympev.2006.06.026" volume="42" year="2007">Bunje and Lindberg (2007)</bibRefCitation>
found 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
to form a sister species to 
<taxonomicName genus="T." lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
<emphasis italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
</taxonomicName>
. 
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. (2019a)</bibRefCitation>
reaffirmed this clade but found further support for a number of other species not included by 
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2006.06.026" author="Bunje, PME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="25" pagination="373 - 387" refId="B35" refString="Bunje, PME, Lindberg, DR, 2007. Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development. Molecular Phylogenetics and Evolution 42 (2): 373 - 387, DOI: https://doi.org/10.1016/j.ympev.2006.06.026" title="Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development." url="https://doi.org/10.1016/j.ympev.2006.06.026" volume="42" year="2007">Bunje and Lindberg (2007)</bibRefCitation>
, including 
<taxonomicName genus="T." lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
<emphasis italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
</taxonomicName>
, which 
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. (2019a)</bibRefCitation>
found to be the closest sister species to 
<taxonomicName genus="T." lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
</taxonomicName>
(also see Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
). 
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus jordani</emphasis>
</taxonomicName>
and 
<taxonomicName genus="T." lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
<emphasis italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
</taxonomicName>
likely diverged in the early Pleistocene (
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
; Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the &quot; Systematic checklist &quot; for more information). Small green (PP ≥ 0.95) and red (PP &lt;0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="25">
Notes on synonymy and homonymy: 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Novitates Conchologicae, Supplement" pageId="0" pageNumber="25" pagination="1 - 127" publicationUrl="https://biodiversitylibrary.org/page/12980992" refId="B98" refString="Martens, Ev, 1874. Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates Conchologicae, Supplement 5: 1 - 127, https://biodiversitylibrary.org/page/12980992" title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht." url="https://biodiversitylibrary.org/page/12980992" volume="5" year="1874">Martens (1874)</bibRefCitation>
described two taxa from the Chabur River near Ras al-Aynin Syria, 
<taxonomicName authorityName="Martens" authorityYear="1874" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina cinctella" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="cinctella">
<emphasis italics="true" pageId="0" pageNumber="25">Neritina cinctella</emphasis>
</taxonomicName>
Martens, 1874 and 
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina anatolica var. mesopotamica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="variety" species="anatolica" variety="mesopotamica">Neritina anatolica var. mesopotamica</taxonomicName>
Martens, 1874. The latter variety is a junior homonym of 
<taxonomicName genus="N." lsidName="N. meridionalis var. mesopotamica" pageId="0" pageNumber="25" rank="variety" species="meridionalis" variety="mesopotamica">N. meridionalis var. mesopotamica</taxonomicName>
Mousson, 1874, which was published earlier (see also the discussion in the postface of 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Novitates Conchologicae, Supplement" pageId="0" pageNumber="25" pagination="1 - 127" publicationUrl="https://biodiversitylibrary.org/page/12980992" refId="B98" refString="Martens, Ev, 1874. Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht. Novitates Conchologicae, Supplement 5: 1 - 127, https://biodiversitylibrary.org/page/12980992" title="Ueber vorderasiatische Conchylien nach den Sammlungen des Prof. Hausknecht." url="https://biodiversitylibrary.org/page/12980992" volume="5" year="1874">Martens 1874</bibRefCitation>
and 
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer &amp; Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer &amp; Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
). Based on newly collected material, 
<bibRefCitation DOI="https://doi.org/10.1080/09397140.1989.10637575" author="Schuett, H" journalOrPublisher="Zoology in the Middle East" pageId="0" pageNumber="25" pagination="55 - 58" refId="B144" refString="Schuett, H, Sesen, R, 1989a. The freshwater molluscs of Ceylanpinar. Zoology in the Middle East 3 (1): 55 - 58, DOI: https://doi.org/10.1080/09397140.1989.10637575" title="The freshwater molluscs of Ceylanpinar." url="https://doi.org/10.1080/09397140.1989.10637575" volume="3" year="1989 a">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
and 
<normalizedToken originalValue="Şeşen">Sesen</normalizedToken>
(1989a
</bibRefCitation>
, 
<bibRefCitation author="Schuett, H" journalOrPublisher="Basteria" pageId="0" pageNumber="25" pagination="39 - 46" refId="B145" refString="Schuett, H, Sesen, R, 1989b. Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae). Basteria 53 (1-3): 39 - 46" title="Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae)." volume="53" year="1989 b">1989b</bibRefCitation>
) considered all three taxa (
<taxonomicName authorityName="Martens" authorityYear="1874" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina cinctella" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="cinctella">
<emphasis italics="true" pageId="0" pageNumber="25">Neritina cinctella</emphasis>
</taxonomicName>
, 
<taxonomicName genus="N." lsidName="N. anatolica var. mesopotamica" pageId="0" pageNumber="25" rank="variety" species="anatolica" variety="mesopotamica">N. anatolica var. mesopotamica</taxonomicName>
, and 
<taxonomicName genus="N." lsidName="N. meridionalis var. mesopotamica" pageId="0" pageNumber="25" rank="variety" species="meridionalis" variety="mesopotamica">N. meridionalis var. mesopotamica</taxonomicName>
) to belong to the same species given multiple intermediate forms. Although they were aware of the homonymy issue and that 
<normalizedToken originalValue="Mousson’s">Mousson's</normalizedToken>
name has priority, they erroneously chose 
<taxonomicName genus="T." lsidName="T. cinctellus" pageId="0" pageNumber="25" rank="species" species="cinctellus">
<emphasis italics="true" pageId="0" pageNumber="25">T. cinctellus</emphasis>
</taxonomicName>
as the name of the species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="distribution">
<paragraph pageId="0" pageNumber="25">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="25">
<taxonomicName class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus jordani" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="jordani">
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus jordani</emphasis>
</taxonomicName>
is common throughout southern Anatolia, the Levant, Mesopotamia, and parts of the Middle East, extending to at least southern Iran, although it is probably not found eastward of the Zagros Mountains (
<bibRefCitation author="Roth, G" editor="Krupp, F" journalOrPublisher="Dr. ludwig Reichert Verlag, Wiesbaden, Germany" pageId="0" pageNumber="25" pagination="72 - 79" refId="B136" refString="Roth, G, 1987. Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae). In: Krupp, F, Schneider, W, Kinzelbach, R, Eds., Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East. Dr. ludwig Reichert Verlag, Wiesbaden, Germany: 72 - 79" title="Data on the distribution and faunal history of the genus Theodoxus in the Middle East (Gastropoda: Neritidae)." volumeTitle="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East." year="1987">Roth 1987</bibRefCitation>
; 
<bibRefCitation author="Schuett, H" journalOrPublisher="Basteria" pageId="0" pageNumber="25" pagination="39 - 46" refId="B145" refString="Schuett, H, Sesen, R, 1989b. Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae). Basteria 53 (1-3): 39 - 46" title="Theodoxus in South-eastern Anatolia, Turkey (Gastropoda Prosobranchia, Neritidae)." volume="53" year="1989 b">
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
and 
<normalizedToken originalValue="Şeşen">Sesen</normalizedToken>
1989b
</bibRefCitation>
; 
<bibRefCitation author="Bandel, K" journalOrPublisher="Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg" pageId="0" pageNumber="25" pagination="65 - 164" refId="B20" refString="Bandel, K, 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg 85: 65 - 164" title="The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)." volume="85" year="2001">Bandel 2001</bibRefCitation>
; 
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
; Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3A</figureCitation>
). It is also present on Cyprus and in the Nile River system, Egypt (
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
). Some records of this species from Iran and Iraq (e.g. 
<taxonomicName genus="N." lsidName="N. mesopotamica" pageId="0" pageNumber="25" rank="species" species="mesopotamica">
<emphasis italics="true" pageId="0" pageNumber="25">N. mesopotamica</emphasis>
</taxonomicName>
sensu 
<bibRefCitation author="Mansoorian, AB" journalOrPublisher="Iranian International Journal of Science" pageId="0" pageNumber="25" pagination="96 - 103" refId="B95" refString="Mansoorian, AB, 2001. Freshwater Gastropod of Khuzestan Province, South-West Iran. Iranian International Journal of Science 2 (2): 96 - 103" title="Freshwater Gastropod of Khuzestan Province, South-West Iran." volume="2" year="2001">Mansoorian 2001</bibRefCitation>
) may be misidentifications of 
<taxonomicName genus="N." lsidName="N. schlaeflii" pageId="0" pageNumber="25" rank="species" species="schlaeflii">
<emphasis italics="true" pageId="0" pageNumber="25">N. schlaeflii</emphasis>
</taxonomicName>
Mousson, 1874 which inhabits the lower reaches of the Euphrates, Karun, and Tigris rivers, as well as other brackish water systems around the Persian Gulf (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.219.3406" author="Gloeer, P" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="11 - 61" refId="B63" refString="Gloeer, P, Pesic, V, 2012. The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species. ZooKeys 219: 11 - 61, DOI: https://doi.org/10.3897/zookeys.219.3406" title="The freshwater snails (Gastropoda) of Iran, with descriptions of two new genera and eight new species." url="https://doi.org/10.3897/zookeys.219.3406" volume="219" year="2012">
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
and 
<normalizedToken originalValue="Pešić">Pesic</normalizedToken>
2012
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>