683 lines
67 KiB
XML
683 lines
67 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5666990" ID-GBIF-Dataset="d37b958b-b8e2-4ba1-b4ef-836e6b7fa757" ID-GBIF-Taxon="133186524" ID-ISSN="0936-9902" ID-Zenodo-Dep="5666990" checkinTime="1504770362900" checkinUser="plazi" docAuthor="Maurice Kottelat" docDate="2017" docId="8E0587ECFFB8FD7DFD55FA07FDFBFA4C" docLanguage="en" docName="2017 Schistura systomos.pdf" docOrigin="Ichthyol. Explor. Freshwaters 28 (1)" docStyle="DocumentStyle{}" docTitle="Schistura systomos Kottelat, 2017, new species" docType="treatment" docVersion="8" lastPageId="6" lastPageNumber="15" masterDocId="723CFF94FFB9FD7BFFA7FFE6FFC8FFD7" masterDocTitle="Schistura systomos ı a new species of loach from southern Laos (Teleostei: Nemacheilidae)" masterLastPageNumber="16" masterPageNumber="9" pageId="1" pageNumber="10" updateTime="1636544149692" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Schistura systomos ı a new species of loach from southern Laos (Teleostei: Nemacheilidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maurice Kottelat</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Ichthyol. Explor. Freshwaters</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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||
<mods:detail type="volume">
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||
<mods:number>28</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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||
<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>9</mods:start>
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||
<mods:end>16</mods:end>
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||
</mods:extent>
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||
</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.5281/zenodo.886698</mods:identifier>
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<mods:identifier type="GBIF-Dataset">d37b958b-b8e2-4ba1-b4ef-836e6b7fa757</mods:identifier>
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<mods:identifier type="ISSN">0936-9902</mods:identifier>
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<mods:identifier type="Zenodo-Dep">886698</mods:identifier>
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<mods:identifier type="ZooBank">C1E005DB-59E5-47AA-A0B5-E01F15A5E21F</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5666990" ID-GBIF-Taxon="133186524" ID-Zenodo-Dep="5666990" LSID="urn:lsid:plazi:treatment:8E0587ECFFB8FD7DFD55FA07FDFBFA4C" httpUri="http://treatment.plazi.org/id/8E0587ECFFB8FD7DFD55FA07FDFBFA4C" lastPageId="6" lastPageNumber="15" pageId="1" pageNumber="10">
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<heading bold="true" centered="true" fontSize="9" level="2" pageId="1" pageNumber="10" reason="4">
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<subSubSection box="[754,1108,1505,1528]" pageId="1" pageNumber="10" type="nomenclature">
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||
<paragraph blockId="1.[754,1108,1505,1557]" box="[754,1108,1505,1528]" pageId="1" pageNumber="10">
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||
<emphasis bold="true" box="[754,1108,1505,1528]" pageId="1" pageNumber="10">
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||
<taxonomicName box="[754,963,1505,1527]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="10" phylum="Chordata" rank="species" species="systomos">
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<emphasis bold="true" box="[754,963,1505,1527]" italics="true" pageId="1" pageNumber="10">Schistura systomos</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[976,1108,1506,1528]" pageId="1" pageNumber="10" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[876,985,1533,1557]" pageId="1" pageNumber="10" type="description">
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||
<paragraph blockId="1.[754,1108,1505,1557]" box="[876,985,1533,1557]" pageId="1" pageNumber="10">
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||
(
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<figureCitation box="[884,978,1533,1557]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStartId-0="1.[129,161,1032,1051]" captionStartId-1="2.[98,130,1231,1250]" captionTargetBox-0="[153,1168,181,996]" captionTargetBox-1="[119,1132,177,1182]" captionTargetId-0="figure@1.[129,1187,428,739]" captionTargetId-1="figure@2.[98,1156,432,722]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Schistura systomos ı MHNG 2767.056 ı holotypeı 35.0 mm SL; Laos: Champasak Province: Bolaven Plateau: Tad Set waterfall." captionText-1="Fig. 2. Schistura systomos ı CMK 23401 ı paratypes; Laos: Champasak Province: Bolaven Plateau: Tad Set waterfall. aı 35.4 mm SL; bı 21.7 mm SL." httpUri-0="https://zenodo.org/record/886700/files/figure.png" httpUri-1="https://zenodo.org/record/886702/files/figure.png" pageId="1" pageNumber="10">Figs. 1-2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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</heading>
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<subSubSection lastPageId="2" lastPageNumber="11" pageId="1" pageNumber="10" type="materials_examined">
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<paragraph blockId="1.[675,1188,1592,1672]" lastBlockId="2.[98,610,1330,1381]" lastPageId="2" lastPageNumber="11" pageId="1" pageNumber="10">
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<materialsCitation ID-GBIF-Occurrence="1612095095" collectingDate="1919-01" collectionCode="MHNG" collectorName="M. Kottelat & January" country="Laos" county="Bolaven Plateau" elevation="1136" lastPageId="2" lastPageNumber="11" latitude="15.300834" location="Xe Seti" longLatPrecision="21" longitude="106.30639" municipality="Tad Set on Houay Set" pageId="1" pageNumber="10" specimenCode="MHNG 2767.056" specimenCount="1" stateProvince="Champasak Province" typeStatus="holotype">
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<emphasis bold="true" box="[675,784,1592,1613]" pageId="1" pageNumber="10">
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<typeStatus box="[675,779,1592,1613]" pageId="1" pageNumber="10">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode box="[795,975,1592,1614]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="10">MHNG 2767.056</specimenCode>
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ı 35.0 mm SL;
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<collectingCountry box="[1131,1184,1592,1613]" name="Laos" pageId="1" pageNumber="10">Laos</collectingCountry>
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:
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<collectingRegion box="[675,921,1621,1643]" country="Laos" name="Champasak" pageId="1" pageNumber="10">Champasak Province</collectingRegion>
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:
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<collectingCounty box="[938,1128,1621,1643]" pageId="1" pageNumber="10">Bolaven Plateau</collectingCounty>
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:
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<collectingMunicipality pageId="1" pageNumber="10">Tad Set on Houay Set</collectingMunicipality>
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(
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<location LSID="urn:lsid:plazi:treatment:8E0587ECFFB8FD7DFD55FA07FDFBFA4C:03736021FFB8FD7AFCCAF994FC70F950" box="[877,952,1650,1671]" country="Laos" county="Bolaven Plateau" latitude="15.300834" longLatPrecision="21" longitude="106.30639" municipality="Tad Set on Houay Set" name="Xe Seti" pageId="1" pageNumber="10" stateProvince="Champasak Province">Xe Setı</location>
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tributary of
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<location LSID="urn:lsid:plazi:treatment:8E0587ECFFB8FD7DFD55FA07FDFBFA4C:03736021FFB8FD7AFBE3F994FB5DF950" box="[1092,1173,1650,1671]" country="Laos" county="Bolaven Plateau" latitude="15.300834" longLatPrecision="21" longitude="106.30639" municipality="Tad Set on Houay Set" name="Xe Don" pageId="1" pageNumber="10" stateProvince="Champasak Province">Xe Don</location>
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)ı
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<emphasis bold="true" box="[106,121,696,1211]" pageId="2" pageNumber="11">a b</emphasis>
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near
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<location LSID="urn:lsid:plazi:treatment:8E0587ECFFB8FD7DFD55FA07FDFBFA4C:03736021FFBBFD79FF30FAD4FE96FA9F" box="[151,350,1330,1352]" country="Laos" county="Bolaven Plateau" latitude="15.300834" longLatPrecision="21" longitude="106.30639" municipality="Tad Set on Houay Set" name="Ban Nonghinkhaoi" pageId="2" pageNumber="11" stateProvince="Champasak Province">Ban Nonghinkhaoı</location>
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<geoCoordinate box="[355,474,1330,1352]" direction="north" orientation="latitude" pageId="2" pageNumber="11" precision="15" value="15.300834">15°18'03" N</geoCoordinate>
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<geoCoordinate box="[479,607,1330,1352]" direction="east" orientation="longitude" pageId="2" pageNumber="11" precision="15" value="106.30639">106°18'23" E</geoCoordinate>
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;
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<elevation box="[98,210,1359,1381]" metricMagnitude="3" metricUnit="m" metricValue="1.136" pageId="2" pageNumber="11" unit="m" value="1136.0">
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<quantity box="[98,173,1359,1381]" metricMagnitude="3" metricUnit="m" metricValue="1.136" pageId="2" pageNumber="11" unit="m" value="1136.0">1136 m</quantity>
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asl
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</elevation>
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;
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<collectorName box="[221,340,1359,1381]" pageId="2" pageNumber="11">M. Kottelat</collectorName>
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et al.
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<collectingDate box="[397,435,1359,1381]" pageId="2" pageNumber="11" value="1919-01">
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<date box="[397,435,1359,1381]" pageId="2" pageNumber="11" value="1919-01">ı 19</date>
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</collectingDate>
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<collectorName box="[442,525,1359,1380]" pageId="2" pageNumber="11">January</collectorName>
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2013.
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</materialsCitation>
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</paragraph>
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<caption httpUri="https://zenodo.org/record/886702/files/figure.png" pageId="2" pageNumber="11" targetBox="[119,1132,177,1182]" targetPageId="2">
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<paragraph blockId="2.[98,1158,1231,1276]" pageId="2" pageNumber="11">
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<emphasis bold="true" box="[98,155,1231,1250]" pageId="2" pageNumber="11">Fig. 2.</emphasis>
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<taxonomicName box="[166,331,1231,1250]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="11" phylum="Chordata" rank="species" species="systomos">
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<emphasis box="[166,331,1231,1250]" italics="true" pageId="2" pageNumber="11">
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||
Schistura
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<taxonomicName box="[253,331,1232,1250]" pageId="2" pageNumber="11" rank="species" species="systomos">systomos</taxonomicName>
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</emphasis>
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||
</taxonomicName>
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ı CMK 23401ı paratypes; Laos: Champasak Province: Bolaven Plateau: Tad Set waterfall.
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<emphasis bold="true" box="[98,114,1257,1276]" pageId="2" pageNumber="11">a,</emphasis>
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35.4 mm SL;
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<emphasis bold="true" box="[246,264,1257,1276]" pageId="2" pageNumber="11">b,</emphasis>
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21.7 mm SL.
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</paragraph>
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</caption>
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<paragraph blockId="2.[98,610,1415,1496]" pageId="2" pageNumber="11">
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<materialsCitation ID-GBIF-Occurrence="1612095096" collectingDate="6 i" collectingDateMax="2 i" collectingDateMin="6 i" collectionCode="MHNG" collectorName="M. Kottelat & January" country="Laos" county="Bolaven Plateau" elevation="1136" latitude="15.300834" location="Xe Seti" longLatPrecision="21" longitude="106.30639" municipality="Tad Set on Houay Set" pageId="2" pageNumber="11" specimenCount="1" stateProvince="Champasak Province" typeStatus="paratype">
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||
<emphasis bold="true" box="[98,211,1417,1438]" pageId="2" pageNumber="11">
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<typeStatus box="[98,206,1417,1438]" pageId="2" pageNumber="11">Paratypes</typeStatus>
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.
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</emphasis>
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<accessionNumber box="[223,350,1416,1437]" httpUri="http://www.uniprot.org/uniprot/CMK23401" pageId="2" pageNumber="11">CMK 23401</accessionNumber>
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||
ı
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||
<collectingDate box="[366,384,1416,1438]" pageId="2" pageNumber="11">
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||
<date box="[366,384,1416,1438]" pageId="2" pageNumber="11">6ı</date>
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||
</collectingDate>
|
||
<quantity box="[394,539,1415,1438]" metricMagnitude="-2" metricUnit="m" metricValue="2.8549999999999995" metricValueMax="3.54" metricValueMin="2.17" pageId="2" pageNumber="11" unit="mm" value="28.549999999999997" valueMax="35.4" valueMin="21.7">21.7-35.4 mm</quantity>
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SL;
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||
<collectingDate box="[592,610,1416,1437]" pageId="2" pageNumber="11">
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<date box="[592,610,1416,1437]" pageId="2" pageNumber="11">2ı</date>
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||
</collectingDate>
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||
ethanol-fixedı
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||
<quantity box="[258,405,1444,1467]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" metricValueMax="3.16" metricValueMin="2.86" pageId="2" pageNumber="11" unit="mm" value="30.1" valueMax="31.6" valueMin="28.6">28.6-31.6 mm</quantity>
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SL; same data as holotype.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="3" lastPageNumber="12" pageId="2" pageNumber="11" type="diagnosis">
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||
<paragraph blockId="2.[98,611,1531,1669]" lastBlockId="2.[644,1157,1330,1669]" pageId="2" pageNumber="11">
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||
<emphasis bold="true" box="[98,216,1531,1553]" pageId="2" pageNumber="11">Diagnosis.</emphasis>
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||
<taxonomicName box="[228,422,1532,1553]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="11" phylum="Chordata" rank="species" species="systomos">
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<emphasis box="[228,422,1532,1553]" italics="true" pageId="2" pageNumber="11">Schistura systomos</emphasis>
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</taxonomicName>
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||
is distinguished from the other species of the genus in Southeast Asia by having a small mouth (gape about 2.9- 3.1 times in maximum head width;
|
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<figureCitation box="[487,543,1618,1640]" captionStart="Fig. 3" captionStartId="5.[129,161,552,571]" captionTargetBox="[154,1174,181,537]" captionTargetId="figure@5.[661,1186,156,536]" captionTargetPageId="5" captionText="Fig. 3. Ventral view of mouth of Schistura species: aı narrow mouth in S. systomos ı MHNG 2767.056 ı holotypeı 35.0 mm SL; and bı wide mouth in other Schistura species (here S. clatrata ı CMK 23398 ı 81.5 mm SL; collected together with S. systomos)." httpUri="https://zenodo.org/record/886704/files/figure.png" pageId="2" pageNumber="11">Fig. 3</figureCitation>
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||
a)ı occupying only the median half of head width at its position (vs less than about two timesı lateral extremity of lips reaching or almost reaching sides of head in ventral view;
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||
<figureCitation box="[891,948,1388,1410]" captionStart="Fig. 3" captionStartId="5.[129,161,552,571]" captionTargetBox="[154,1174,181,537]" captionTargetId="figure@5.[661,1186,156,536]" captionTargetPageId="5" captionText="Fig. 3. Ventral view of mouth of Schistura species: aı narrow mouth in S. systomos ı MHNG 2767.056 ı holotypeı 35.0 mm SL; and bı wide mouth in other Schistura species (here S. clatrata ı CMK 23398 ı 81.5 mm SL; collected together with S. systomos)." httpUri="https://zenodo.org/record/886704/files/figure.png" pageId="2" pageNumber="11">Fig. 3</figureCitation>
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b); a blunt snout (vs. rounded or pointed) resulting in a relatively deep infraorbital area; head about circular in crosssection (vs. depressed)ı lower surface flat only in middle (vs. whole surface flat); infraorbital area roundedı not expanded laterally and leaving the eyes partly or almost visible in ventral view (vs. infraorbital area vertical or expanded laterally or swollenı eyes visible only in dorsal view).
|
||
</paragraph>
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||
<paragraph blockId="2.[644,1157,1330,1669]" lastBlockId="3.[675,1187,162,472]" lastPageId="3" lastPageNumber="12" pageId="2" pageNumber="11">
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||
Additional diagnostic charactersı not unique to the species are: small size at maturity (
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<quantity box="[567,640,162,184]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="12" unit="mm" value="35.0">35 mm</quantity>
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SL and possibly less); wide interorbital area (interorbital distance about 1.7-2.1 times eye diameter); depth of caudal peduncle 1.5-1.7 times in its length; 8
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||
<emphasis bold="true" box="[219,223,277,289]" pageId="3" pageNumber="12">1</emphasis>
|
||
/
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||
<emphasis bold="true" box="[231,238,285,297]" pageId="3" pageNumber="12">2</emphasis>
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||
branched dorsal-fin rays; pelvic-fin origin below or slightly in front of dorsal-fin origin; pelvic axillary lobe varying from rudimentary and fused to small and free; 9 +8 branched caudal-fin rays; no suborbital flap; 14-17 bars (4-6 predorsalı 3-4 subdorsalı 6-9 postdorsal) on bodyı of regular width and shapeı wider than interspacesı continuous over back with contralateral bars under dorsal fin and on caudal peduncle; on predorsal areaı bars irregularly shaped and setı dissociated in blotches and short stripes. Pattern at caudal-fin base made of a black vertically elongated blotch occupying median
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||
<emphasis bold="true" box="[874,878,248,260]" pageId="3" pageNumber="12">1</emphasis>
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||
/
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||
<emphasis bold="true" box="[887,894,256,268]" pageId="3" pageNumber="12">3</emphasis>
|
||
of caudal baseı not reaching dorsal and ventral midlines; overimposed to itı a dark brown bar extending ventrally beyond the base of the lower unbranched principal rayı and extending dorsally to base of branched ray 5; and a small blotch from above the base of the upper unbranched principal rayı reaching or not the dorsal midline.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12" type="description">
|
||
<paragraph blockId="3.[675,1188,506,1682]" pageId="3" pageNumber="12">
|
||
<emphasis bold="true" box="[675,813,508,530]" pageId="3" pageNumber="12">Description.</emphasis>
|
||
See
|
||
<figureCitation box="[873,1004,506,530]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStartId-0="1.[129,161,1032,1051]" captionStartId-1="2.[98,130,1231,1250]" captionTargetBox-0="[153,1168,181,996]" captionTargetBox-1="[119,1132,177,1182]" captionTargetId-0="figure@1.[129,1187,428,739]" captionTargetId-1="figure@2.[98,1156,432,722]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Schistura systomos ı MHNG 2767.056 ı holotypeı 35.0 mm SL; Laos: Champasak Province: Bolaven Plateau: Tad Set waterfall." captionText-1="Fig. 2. Schistura systomos ı CMK 23401 ı paratypes; Laos: Champasak Province: Bolaven Plateau: Tad Set waterfall. aı 35.4 mm SL; bı 21.7 mm SL." httpUri-0="https://zenodo.org/record/886700/files/figure.png" httpUri-1="https://zenodo.org/record/886702/files/figure.png" pageId="3" pageNumber="12">Figures 1-2</figureCitation>
|
||
for general appearance and
|
||
<tableCitation box="[826,905,536,558]" captionStart="Table 1" captionStartId="3.[128,182,588,607]" captionTargetBox="[128,636,700,1667]" captionText="Table 1. Morphometric data of holotype and two largest paratypes of Schistura systomos. Hı holotype; Pı paratype." httpUri="http://table.plazi.org/id/52D36672FFBAFD78FF27FDAAFF0CFD44" pageId="3" pageNumber="12" tableUuid="52D36672FFBAFD78FF27FDAAFF0CFD44">Table 1</tableCitation>
|
||
for morphometric data of
|
||
<typeStatus box="[675,769,565,587]" pageId="3" pageNumber="12">holotype</typeStatus>
|
||
and 2 largest
|
||
<typeStatus box="[921,1029,566,587]" pageId="3" pageNumber="12">paratypes</typeStatus>
|
||
.
|
||
<collectionCodeEnum box="[1039,1058,565,586]" pageId="3" pageNumber="12">A</collectionCodeEnum>
|
||
moderately elongate nemacheilid with body depth slowly increasing up to slightly in front of dorsal-fin origin. Behind dorsal finı body depth almost uniform until caudal-fin base. Dorsal profile continuous between head and body. Head about circular in cross sectionı lower surface flat only in middle; infraorbital area roundedı not expanded laterally. Body slightly compressed anteriorly to compressed posteriorly. Interorbital area arched. Eye dorso-lateralı at edge of cranium and partly or almost visible from below because infraorbital area not expanded laterally; diameter 1.7-2.1 times in interorbital width; in lateral viewı eye located lower than profile of head. Cheeks not swollen. Snout roundedı blunt. Depth of caudal peduncle 1.5-1.7 times in its lengthı uniform; low dorsal keel on posterior fourth of post-dorsal area; no ventral keel. Dorsal keel continuous with upper margin of caudal fin. Largest recorded size
|
||
<quantity box="[675,765,1141,1163]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="3" pageNumber="12" unit="mm" value="35.4">35.4 mm</quantity>
|
||
<collectionCode box="[772,802,1141,1162]" country="Sierra Leone" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14608" name="University of Sierra Leone, Njala University College" pageId="3" pageNumber="12">SL</collectionCode>
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||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="52D36672FFBAFD78FF27FDAAFF0CFD44" httpUri="http://table.plazi.org/id/52D36672FFBAFD78FF27FDAAFF0CFD44" pageId="3" pageNumber="12" targetBox="[128,636,700,1667]" targetIsTable="true" targetPageId="3">
|
||
<paragraph blockId="3.[128,641,588,659]" pageId="3" pageNumber="12">
|
||
<emphasis bold="true" box="[128,202,588,607]" pageId="3" pageNumber="12">Table 1.</emphasis>
|
||
Morphometric data of holotype and two largest paratypes of
|
||
<taxonomicName box="[263,436,614,633]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="12" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[263,436,614,633]" italics="true" pageId="3" pageNumber="12">
|
||
Schistura
|
||
<taxonomicName box="[357,436,614,632]" pageId="3" pageNumber="12" rank="species" species="systomos">systomos</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
. Hı holotype; Pı paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="3" pageNumber="12">
|
||
<table box="[128,636,700,1667]" gridcols="4" gridrows="36" pageId="3" pageNumber="12">
|
||
<tr box="[128,636,700,719]" gridrow="0" pageId="3" pageNumber="12" rowspan-0="1">
|
||
<th box="[475,523,700,719]" gridcol="1" gridrow="0" pageId="3" pageNumber="12">P</th>
|
||
<th box="[532,580,700,719]" gridcol="2" gridrow="0" pageId="3" pageNumber="12">H</th>
|
||
<th box="[588,636,700,719]" gridcol="3" gridrow="0" pageId="3" pageNumber="12">P</th>
|
||
</tr>
|
||
<tr box="[128,636,735,754]" gridrow="1" pageId="3" pageNumber="12">
|
||
<th box="[128,466,735,754]" gridcol="0" gridrow="1" pageId="3" pageNumber="12">Standard length (mm)</th>
|
||
<td box="[475,523,735,754]" gridcol="1" gridrow="1" pageId="3" pageNumber="12">34.7</td>
|
||
<td box="[532,580,735,754]" gridcol="2" gridrow="1" pageId="3" pageNumber="12">35.0</td>
|
||
<td box="[588,636,735,754]" gridcol="3" gridrow="1" pageId="3" pageNumber="12">35.4</td>
|
||
</tr>
|
||
<tr box="[128,636,760,780]" gridrow="2" pageId="3" pageNumber="12">
|
||
<th box="[128,466,760,780]" gridcol="0" gridrow="2" pageId="3" pageNumber="12">Total length (mm)</th>
|
||
<td box="[475,523,760,780]" gridcol="1" gridrow="2" pageId="3" pageNumber="12">42.0</td>
|
||
<td box="[532,580,760,780]" gridcol="2" gridrow="2" pageId="3" pageNumber="12">42.3</td>
|
||
<td box="[588,636,760,780]" gridcol="3" gridrow="2" pageId="3" pageNumber="12">42.5</td>
|
||
</tr>
|
||
<tr box="[128,636,794,813]" gridrow="3" pageId="3" pageNumber="12" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th box="[128,466,794,813]" gridcol="0" gridrow="3" pageId="3" pageNumber="12">
|
||
<emphasis bold="true" box="[128,416,794,813]" pageId="3" pageNumber="12">In percent of standard length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[128,636,819,838]" gridrow="4" pageId="3" pageNumber="12">
|
||
<th box="[128,466,819,838]" gridcol="0" gridrow="4" pageId="3" pageNumber="12">Total length</th>
|
||
<td box="[475,523,819,838]" gridcol="1" gridrow="4" pageId="3" pageNumber="12">121.1</td>
|
||
<td box="[532,580,819,838]" gridcol="2" gridrow="4" pageId="3" pageNumber="12">120.7</td>
|
||
<td box="[588,636,819,838]" gridcol="3" gridrow="4" pageId="3" pageNumber="12">119.9</td>
|
||
</tr>
|
||
<tr box="[128,636,845,864]" gridrow="5" pageId="3" pageNumber="12">
|
||
<th box="[128,466,845,864]" gridcol="0" gridrow="5" pageId="3" pageNumber="12">Head length (dorsal)</th>
|
||
<td box="[475,523,845,864]" gridcol="1" gridrow="5" pageId="3" pageNumber="12">21.3</td>
|
||
<td box="[532,580,845,864]" gridcol="2" gridrow="5" pageId="3" pageNumber="12">22.1</td>
|
||
<td box="[588,636,845,864]" gridcol="3" gridrow="5" pageId="3" pageNumber="12">19.8</td>
|
||
</tr>
|
||
<tr box="[128,636,870,889]" gridrow="6" pageId="3" pageNumber="12">
|
||
<th box="[128,466,870,889]" gridcol="0" gridrow="6" pageId="3" pageNumber="12">Head length (lateral)</th>
|
||
<td box="[475,523,870,889]" gridcol="1" gridrow="6" pageId="3" pageNumber="12">22.6</td>
|
||
<td box="[532,580,870,889]" gridcol="2" gridrow="6" pageId="3" pageNumber="12">22.3</td>
|
||
<td box="[588,636,870,889]" gridcol="3" gridrow="6" pageId="3" pageNumber="12">24.2</td>
|
||
</tr>
|
||
<tr box="[128,636,895,914]" gridrow="7" pageId="3" pageNumber="12">
|
||
<th box="[128,466,895,914]" gridcol="0" gridrow="7" pageId="3" pageNumber="12">Predorsal length</th>
|
||
<td box="[475,523,895,914]" gridcol="1" gridrow="7" pageId="3" pageNumber="12">54.5</td>
|
||
<td box="[532,580,895,914]" gridcol="2" gridrow="7" pageId="3" pageNumber="12">54.9</td>
|
||
<td box="[588,636,895,914]" gridcol="3" gridrow="7" pageId="3" pageNumber="12">55.1</td>
|
||
</tr>
|
||
<tr box="[128,636,921,940]" gridrow="8" pageId="3" pageNumber="12">
|
||
<th box="[128,466,921,940]" gridcol="0" gridrow="8" pageId="3" pageNumber="12">Prepelvic length</th>
|
||
<td box="[475,523,921,940]" gridcol="1" gridrow="8" pageId="3" pageNumber="12">50.8</td>
|
||
<td box="[532,580,921,940]" gridcol="2" gridrow="8" pageId="3" pageNumber="12">51.1</td>
|
||
<td box="[588,636,921,940]" gridcol="3" gridrow="8" pageId="3" pageNumber="12">51.6</td>
|
||
</tr>
|
||
<tr box="[128,636,946,966]" gridrow="9" pageId="3" pageNumber="12">
|
||
<th box="[128,466,946,966]" gridcol="0" gridrow="9" pageId="3" pageNumber="12">Pre-anus length</th>
|
||
<td box="[475,523,946,966]" gridcol="1" gridrow="9" pageId="3" pageNumber="12">65.7</td>
|
||
<td box="[532,580,946,966]" gridcol="2" gridrow="9" pageId="3" pageNumber="12">67.4</td>
|
||
<td box="[588,636,946,966]" gridcol="3" gridrow="9" pageId="3" pageNumber="12">66.4</td>
|
||
</tr>
|
||
<tr box="[128,636,972,991]" gridrow="10" pageId="3" pageNumber="12">
|
||
<th box="[128,466,972,991]" gridcol="0" gridrow="10" pageId="3" pageNumber="12">Pre-anal length</th>
|
||
<td box="[475,523,972,991]" gridcol="1" gridrow="10" pageId="3" pageNumber="12">74.6</td>
|
||
<td box="[532,580,972,991]" gridcol="2" gridrow="10" pageId="3" pageNumber="12">76.2</td>
|
||
<td box="[588,636,972,991]" gridcol="3" gridrow="10" pageId="3" pageNumber="12">76.3</td>
|
||
</tr>
|
||
<tr box="[128,636,997,1016]" gridrow="11" pageId="3" pageNumber="12">
|
||
<th box="[128,466,997,1016]" gridcol="0" gridrow="11" pageId="3" pageNumber="12">Head depth</th>
|
||
<td box="[475,523,997,1016]" gridcol="1" gridrow="11" pageId="3" pageNumber="12">12.4</td>
|
||
<td box="[532,580,997,1016]" gridcol="2" gridrow="11" pageId="3" pageNumber="12">12.8</td>
|
||
<td box="[588,636,997,1016]" gridcol="3" gridrow="11" pageId="3" pageNumber="12">12.5</td>
|
||
</tr>
|
||
<tr box="[128,636,1022,1042]" gridrow="12" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1022,1042]" gridcol="0" gridrow="12" pageId="3" pageNumber="12">Body depth at dorsal-fin origin</th>
|
||
<td box="[475,523,1022,1042]" gridcol="1" gridrow="12" pageId="3" pageNumber="12">15.1</td>
|
||
<td box="[532,580,1022,1042]" gridcol="2" gridrow="12" pageId="3" pageNumber="12">19.1</td>
|
||
<td box="[588,636,1022,1042]" gridcol="3" gridrow="12" pageId="3" pageNumber="12">17.1</td>
|
||
</tr>
|
||
<tr box="[128,636,1048,1067]" gridrow="13" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1048,1067]" gridcol="0" gridrow="13" pageId="3" pageNumber="12">Depth of caudal peduncle</th>
|
||
<td box="[475,523,1048,1067]" gridcol="1" gridrow="13" pageId="3" pageNumber="12">11.2</td>
|
||
<td box="[532,580,1048,1067]" gridcol="2" gridrow="13" pageId="3" pageNumber="12">11.1</td>
|
||
<td box="[588,636,1048,1067]" gridcol="3" gridrow="13" pageId="3" pageNumber="12">11.2</td>
|
||
</tr>
|
||
<tr box="[128,636,1073,1092]" gridrow="14" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1073,1092]" gridcol="0" gridrow="14" pageId="3" pageNumber="12">Length of caudal peduncle</th>
|
||
<td box="[475,523,1073,1092]" gridcol="1" gridrow="14" pageId="3" pageNumber="12">18.5</td>
|
||
<td box="[532,580,1073,1092]" gridcol="2" gridrow="14" pageId="3" pageNumber="12">17.4</td>
|
||
<td box="[588,636,1073,1092]" gridcol="3" gridrow="14" pageId="3" pageNumber="12">17.8</td>
|
||
</tr>
|
||
<tr box="[128,636,1099,1118]" gridrow="15" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1099,1118]" gridcol="0" gridrow="15" pageId="3" pageNumber="12">Head width</th>
|
||
<td box="[475,523,1099,1118]" gridcol="1" gridrow="15" pageId="3" pageNumber="12">14.4</td>
|
||
<td box="[532,580,1099,1118]" gridcol="2" gridrow="15" pageId="3" pageNumber="12">14.6</td>
|
||
<td box="[588,636,1099,1118]" gridcol="3" gridrow="15" pageId="3" pageNumber="12">14.9</td>
|
||
</tr>
|
||
<tr box="[128,636,1124,1144]" gridrow="16" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1124,1144]" gridcol="0" gridrow="16" pageId="3" pageNumber="12">Body width at dorsal-fin origin</th>
|
||
<td box="[475,523,1124,1144]" gridcol="1" gridrow="16" pageId="3" pageNumber="12">11.8</td>
|
||
<td box="[532,580,1124,1144]" gridcol="2" gridrow="16" pageId="3" pageNumber="12">15.4</td>
|
||
<td box="[588,636,1124,1144]" gridcol="3" gridrow="16" pageId="3" pageNumber="12">13.4</td>
|
||
</tr>
|
||
<tr box="[128,636,1150,1169]" gridrow="17" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1150,1169]" gridcol="0" gridrow="17" pageId="3" pageNumber="12">Snout length</th>
|
||
<td box="[475,523,1150,1169]" gridcol="1" gridrow="17" pageId="3" pageNumber="12">9.6</td>
|
||
<td box="[532,580,1150,1169]" gridcol="2" gridrow="17" pageId="3" pageNumber="12">9.6</td>
|
||
<td box="[588,636,1150,1169]" gridcol="3" gridrow="17" pageId="3" pageNumber="12">10.4</td>
|
||
</tr>
|
||
<tr box="[128,636,1175,1194]" gridrow="18" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1175,1194]" gridcol="0" gridrow="18" pageId="3" pageNumber="12">Eye diameter</th>
|
||
<td box="[475,523,1175,1194]" gridcol="1" gridrow="18" pageId="3" pageNumber="12">4.6</td>
|
||
<td box="[532,580,1175,1194]" gridcol="2" gridrow="18" pageId="3" pageNumber="12">4.6</td>
|
||
<td box="[588,636,1175,1194]" gridcol="3" gridrow="18" pageId="3" pageNumber="12">4.8</td>
|
||
</tr>
|
||
<tr box="[128,636,1200,1220]" gridrow="19" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1200,1220]" gridcol="0" gridrow="19" pageId="3" pageNumber="12">Interorbital width</th>
|
||
<td box="[475,523,1200,1220]" gridcol="1" gridrow="19" pageId="3" pageNumber="12">8.3</td>
|
||
<td box="[532,580,1200,1220]" gridcol="2" gridrow="19" pageId="3" pageNumber="12">9.8</td>
|
||
<td box="[588,636,1200,1220]" gridcol="3" gridrow="19" pageId="3" pageNumber="12">8.2</td>
|
||
</tr>
|
||
<tr box="[128,636,1226,1245]" gridrow="20" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1226,1245]" gridcol="0" gridrow="20" pageId="3" pageNumber="12">Length of dorsal fin</th>
|
||
<td box="[475,523,1226,1245]" gridcol="1" gridrow="20" pageId="3" pageNumber="12">18.2</td>
|
||
<td box="[532,580,1226,1245]" gridcol="2" gridrow="20" pageId="3" pageNumber="12">18.4</td>
|
||
<td box="[588,636,1226,1245]" gridcol="3" gridrow="20" pageId="3" pageNumber="12">15.8</td>
|
||
</tr>
|
||
<tr box="[128,636,1251,1296]" gridrow="21" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1251,1296]" gridcol="0" gridrow="21" pageId="3" pageNumber="12">Length of upper caudal-fin lobe Length of median caudal-fin rays</th>
|
||
<td box="[475,523,1251,1296]" gridcol="1" gridrow="21" pageId="3" pageNumber="12">20.4 15.8</td>
|
||
<td box="[532,580,1251,1296]" gridcol="2" gridrow="21" pageId="3" pageNumber="12">22.4 16.6</td>
|
||
<td box="[588,636,1251,1296]" gridcol="3" gridrow="21" pageId="3" pageNumber="12">19.6 15.7</td>
|
||
</tr>
|
||
<tr box="[128,636,1302,1321]" gridrow="22" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1302,1321]" gridcol="0" gridrow="22" pageId="3" pageNumber="12">Length of lower caudal-fin lobe</th>
|
||
<td box="[475,523,1302,1321]" gridcol="1" gridrow="22" pageId="3" pageNumber="12">20.2</td>
|
||
<td box="[532,580,1302,1321]" gridcol="2" gridrow="22" pageId="3" pageNumber="12">21.1</td>
|
||
<td box="[588,636,1302,1321]" gridcol="3" gridrow="22" pageId="3" pageNumber="12">19.8</td>
|
||
</tr>
|
||
<tr box="[128,636,1327,1347]" gridrow="23" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1327,1347]" gridcol="0" gridrow="23" pageId="3" pageNumber="12">Length of anal fin</th>
|
||
<td box="[475,523,1327,1347]" gridcol="1" gridrow="23" pageId="3" pageNumber="12">18.0</td>
|
||
<td box="[532,580,1327,1347]" gridcol="2" gridrow="23" pageId="3" pageNumber="12">17.8</td>
|
||
<td box="[588,636,1327,1347]" gridcol="3" gridrow="23" pageId="3" pageNumber="12">16.6</td>
|
||
</tr>
|
||
<tr box="[128,636,1353,1372]" gridrow="24" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1353,1372]" gridcol="0" gridrow="24" pageId="3" pageNumber="12">Length of pelvic fin</th>
|
||
<td box="[475,523,1353,1372]" gridcol="1" gridrow="24" pageId="3" pageNumber="12">16.6</td>
|
||
<td box="[532,580,1353,1372]" gridcol="2" gridrow="24" pageId="3" pageNumber="12">17.3</td>
|
||
<td box="[588,636,1353,1372]" gridcol="3" gridrow="24" pageId="3" pageNumber="12">15.2</td>
|
||
</tr>
|
||
<tr box="[128,636,1378,1397]" gridrow="25" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1378,1397]" gridcol="0" gridrow="25" pageId="3" pageNumber="12">Length of pectoral fin</th>
|
||
<td box="[475,523,1378,1397]" gridcol="1" gridrow="25" pageId="3" pageNumber="12">18.5</td>
|
||
<td box="[532,580,1378,1397]" gridcol="2" gridrow="25" pageId="3" pageNumber="12">17.9</td>
|
||
<td box="[588,636,1378,1397]" gridcol="3" gridrow="25" pageId="3" pageNumber="12">17.5</td>
|
||
</tr>
|
||
<tr box="[128,636,1412,1431]" gridrow="26" pageId="3" pageNumber="12" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th box="[128,466,1412,1431]" gridcol="0" gridrow="26" pageId="3" pageNumber="12">
|
||
<emphasis bold="true" box="[128,445,1412,1431]" pageId="3" pageNumber="12">In percent of dorsal head length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[128,636,1437,1456]" gridrow="27" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1437,1456]" gridcol="0" gridrow="27" pageId="3" pageNumber="12">Snout length</th>
|
||
<td box="[475,523,1437,1456]" gridcol="1" gridrow="27" pageId="3" pageNumber="12">45</td>
|
||
<td box="[532,580,1437,1456]" gridcol="2" gridrow="27" pageId="3" pageNumber="12">43</td>
|
||
<td box="[588,636,1437,1456]" gridcol="3" gridrow="27" pageId="3" pageNumber="12">53</td>
|
||
</tr>
|
||
<tr box="[128,636,1462,1482]" gridrow="28" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1462,1482]" gridcol="0" gridrow="28" pageId="3" pageNumber="12">Eye diameter</th>
|
||
<td box="[475,523,1462,1482]" gridcol="1" gridrow="28" pageId="3" pageNumber="12">21</td>
|
||
<td box="[532,580,1462,1482]" gridcol="2" gridrow="28" pageId="3" pageNumber="12">66</td>
|
||
<td box="[588,636,1462,1482]" gridcol="3" gridrow="28" pageId="3" pageNumber="12">75</td>
|
||
</tr>
|
||
<tr box="[128,636,1488,1507]" gridrow="29" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1488,1507]" gridcol="0" gridrow="29" pageId="3" pageNumber="12">Interorbital width</th>
|
||
<td box="[475,523,1488,1507]" gridcol="1" gridrow="29" pageId="3" pageNumber="12">39</td>
|
||
<td box="[532,580,1488,1507]" gridcol="2" gridrow="29" pageId="3" pageNumber="12">44</td>
|
||
<td box="[588,636,1488,1507]" gridcol="3" gridrow="29" pageId="3" pageNumber="12">41</td>
|
||
</tr>
|
||
<tr box="[128,636,1513,1532]" gridrow="30" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1513,1532]" gridcol="0" gridrow="30" pageId="3" pageNumber="12">Mouth width</th>
|
||
<td box="[475,523,1513,1532]" gridcol="1" gridrow="30" pageId="3" pageNumber="12">23</td>
|
||
<td box="[532,580,1513,1532]" gridcol="2" gridrow="30" pageId="3" pageNumber="12">21</td>
|
||
<td box="[588,636,1513,1532]" gridcol="3" gridrow="30" pageId="3" pageNumber="12">24</td>
|
||
</tr>
|
||
<tr box="[128,636,1547,1566]" gridrow="31" pageId="3" pageNumber="12" rowspan-1="1" rowspan-2="1" rowspan-3="1">
|
||
<th box="[128,466,1547,1566]" gridcol="0" gridrow="31" pageId="3" pageNumber="12">
|
||
<emphasis bold="true" box="[128,446,1547,1566]" pageId="3" pageNumber="12">In percent of lateral head length</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[128,636,1572,1591]" gridrow="32" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1572,1591]" gridcol="0" gridrow="32" pageId="3" pageNumber="12">Snout length</th>
|
||
<td box="[475,523,1572,1591]" gridcol="1" gridrow="32" pageId="3" pageNumber="12">43</td>
|
||
<td box="[532,580,1572,1591]" gridcol="2" gridrow="32" pageId="3" pageNumber="12">43</td>
|
||
<td box="[588,636,1572,1591]" gridcol="3" gridrow="32" pageId="3" pageNumber="12">43</td>
|
||
</tr>
|
||
<tr box="[128,636,1597,1617]" gridrow="33" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1597,1617]" gridcol="0" gridrow="33" pageId="3" pageNumber="12">Eye diameter</th>
|
||
<td box="[475,523,1597,1617]" gridcol="1" gridrow="33" pageId="3" pageNumber="12">20</td>
|
||
<td box="[532,580,1597,1617]" gridcol="2" gridrow="33" pageId="3" pageNumber="12">21</td>
|
||
<td box="[588,636,1597,1617]" gridcol="3" gridrow="33" pageId="3" pageNumber="12">20</td>
|
||
</tr>
|
||
<tr box="[128,636,1623,1642]" gridrow="34" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1623,1642]" gridcol="0" gridrow="34" pageId="3" pageNumber="12">Interorbital width</th>
|
||
<td box="[475,523,1623,1642]" gridcol="1" gridrow="34" pageId="3" pageNumber="12">37</td>
|
||
<td box="[532,580,1623,1642]" gridcol="2" gridrow="34" pageId="3" pageNumber="12">44</td>
|
||
<td box="[588,636,1623,1642]" gridcol="3" gridrow="34" pageId="3" pageNumber="12">34</td>
|
||
</tr>
|
||
<tr box="[128,636,1648,1667]" gridrow="35" pageId="3" pageNumber="12">
|
||
<th box="[128,466,1648,1667]" gridcol="0" gridrow="35" pageId="3" pageNumber="12">Mouth width</th>
|
||
<td box="[475,523,1648,1667]" gridcol="1" gridrow="35" pageId="3" pageNumber="12">22</td>
|
||
<td box="[532,580,1648,1667]" gridcol="2" gridrow="35" pageId="3" pageNumber="12">21</td>
|
||
<td box="[588,636,1648,1667]" gridcol="3" gridrow="35" pageId="3" pageNumber="12">20</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="3.[675,1188,506,1682]" pageId="3" pageNumber="12">
|
||
Dorsal fin with 4 unbranched and 8
|
||
<emphasis bold="true" box="[1132,1136,1169,1181]" pageId="3" pageNumber="12">1</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[1145,1152,1178,1190]" pageId="3" pageNumber="12">2</emphasis>
|
||
(7) branched rays; distal margin slightly convex or straight. Second branched ray longest. Pectoral fin with 1 unbranched and 8 (1) or 9 (6*) branched raysı roundedı reaching about halfway of distance to pelvic-fin base. No axillary pectoral lobe. Pelvic fin with 1 unbranched and 7 (7) branched raysı reaching to anusı rounded; posterior margin convex; origin below or slightly in front of dorsalfin origin; axillary lobe from rudimentary and fused to small and free. Anus situated about 1.5 eye diameter in front of anal fin. Anal fin with 3 unbranched and 5
|
||
<emphasis bold="true" box="[896,900,1515,1527]" pageId="3" pageNumber="12">1</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[908,915,1524,1536]" pageId="3" pageNumber="12">2</emphasis>
|
||
(7) branched rays; distal margin convex. Caudal fin with 9 + 8 (7) branched raysı emarginateı lobes roundedı of equal length.
|
||
</paragraph>
|
||
<paragraph blockId="3.[675,1188,506,1682]" lastBlockId="4.[98,611,160,1221]" lastPageId="4" lastPageNumber="13" pageId="3" pageNumber="12">
|
||
Body covered by embedded scalesı except on predorsal area and on belly in front of anus. Lateral line completeı pores difficult to count with accuracyı about 80-
|
||
<date box="[363,393,162,183]" pageId="4" pageNumber="13" value="1990-01">90ı</date>
|
||
anterior most 16-29 conspicuousı remainder indistinct (fixation artefactı pores hidden in midlateral groove resulting from dehydratation by ethanol). In anterior half of bodyı a small auxiliary pore or papillae above and below some lateral line tubes (usually by pair). Cephalic lateral line system with 6 supraorbitalı 4 + 11-12 infraorbitalı 8-9 preoperculo-mandibular and 3 supratemporal pores.
|
||
</paragraph>
|
||
<paragraph blockId="4.[98,611,160,1221]" pageId="4" pageNumber="13">
|
||
Anterior nare pierced in front side of a pointed triangular flap-like tube. Posterior nare adjacent to anterior one. Mouth archedı narrowı gape about 2.9-3.1 times in maximum head widthı occupying only median half of head width and lateral extremity of lips by far not reaching lateral outline of head in ventral view; gape about two times wider than long (
|
||
<figureCitation box="[355,414,623,645]" captionStart="Fig. 3" captionStartId="5.[129,161,552,571]" captionTargetBox="[154,1174,181,537]" captionTargetId="figure@5.[661,1186,156,536]" captionTargetPageId="5" captionText="Fig. 3. Ventral view of mouth of Schistura species: aı narrow mouth in S. systomos ı MHNG 2767.056 ı holotypeı 35.0 mm SL; and bı wide mouth in other Schistura species (here S. clatrata ı CMK 23398 ı 81.5 mm SL; collected together with S. systomos)." httpUri="https://zenodo.org/record/886704/files/figure.png" pageId="4" pageNumber="13">Fig. 3</figureCitation>
|
||
a). Lips thick. Upper lip with a rounded median notch in which processus dentiformis is visibleı with numerous fine and shallow furrowsı edge finely crenulated. Processus dentiformis presentı well-developed. Lower lip with conspicuous V-shaped median interruption; broad median part with two or three sulciı narrower lateral partı shortı with a few fine and shallow furrows. Tip of lower jaw exposed. A shallow median concavity in lower jaw in which processus dentiformis is fitting. Inner rostral barbel not reaching corner of mouth; outer one reaching slightly beyond corner of mouth or vertical of anterior margin of eye. Maxillary barbel reaching below vertical of posterior half of eye. Intestine thin (about
|
||
<quantity box="[401,479,1055,1076]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="13" unit="mm" value="0.3">0.3 mm</quantity>
|
||
diameter in
|
||
<quantity box="[98,188,1084,1106]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="4" pageNumber="13" unit="mm" value="35.4">35.4 mm</quantity>
|
||
SL specimen)ı straightı without bend behind stomach (
|
||
<figureCitation box="[283,343,1112,1135]" captionStart="Fig. 4" captionStartId="5.[128,163,1632,1651]" captionTargetBox="[222,546,1038,1613]" captionTargetId="figure@5.[222,546,1037,1613]" captionTargetPageId="5" captionText="Fig. 4. Schistura systomos ı CMK 23401 ı paratypeı 35.4 mm SL; digestive tract. Scale bar 1 mm." httpUri="https://zenodo.org/record/886706/files/figure.png" pageId="4" pageNumber="13">Fig. 4</figureCitation>
|
||
); stomach comparatively smallı about
|
||
<quantity box="[237,297,1141,1163]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="13" unit="mm" value="3.0">3 mm</quantity>
|
||
longı body of stomach wideı swollen. Air bladder without posterior chamber in body cavity.
|
||
</paragraph>
|
||
<paragraph blockId="4.[98,611,1256,1365]" pageId="4" pageNumber="13">
|
||
<emphasis bold="true" box="[98,320,1256,1278]" pageId="4" pageNumber="13">Sexual dimorphism.</emphasis>
|
||
None observed. No specimen with suborbital flapı groove or slit. No modified pectoral raysı no tuberclesı no patches of unculi. Ripe females deeper bodied.
|
||
</paragraph>
|
||
<paragraph blockId="4.[98,611,1400,1653]" lastBlockId="4.[644,1158,162,1250]" pageId="4" pageNumber="13">
|
||
<emphasis bold="true" box="[98,221,1400,1422]" pageId="4" pageNumber="13">Coloration.</emphasis>
|
||
After three weeks in formalin. Head and body background colour pale yellowish brownı throat and belly whitish to pale yellowish; except otherwise statedı all markings dark brown to blackish. Head without distinct pattern. Body with 14-17 bars (4-6 predorsalı 3-4 subdorsalı 6-9 postdorsal)ı reaching downwards to level of pectoral finı or to ventral midline on caudal peduncleı continuous over back with contralateral bars under dorsal fin and on caudal peduncleı of regular width and shapeı wider than interspacesı anterior bars slightly broader than posterior ones. On predorsal areaı bars irregularly shaped and setı dissociated in blotches and short stripes and usually not meeting contralaterals.
|
||
</paragraph>
|
||
<paragraph blockId="4.[644,1158,162,1250]" pageId="4" pageNumber="13">
|
||
A conspicuous black blotch on caudal-fin base. Black inner pigments forming a vertically elongated blotch occupying median
|
||
<emphasis bold="true" box="[1106,1110,392,404]" pageId="4" pageNumber="13">1</emphasis>
|
||
/
|
||
<emphasis bold="true" box="[1118,1125,400,412]" pageId="4" pageNumber="13">3</emphasis>
|
||
of caudal-fin baseı not reaching dorsal and ventral midlines; overimposedı a dark brown bar extending ventrally beyond base of lower unbranched principal rayı dorsally to base of branched ray 5 (from top). A smaller blotch from above base of upper unbranched principal rayı reaching or not dorsal midline. Space between bar on caudal-fin base and preceeding bar paler than rest of body background coloration. A wide inner axial stripe made of chevron-shaped elements present but faintly markedı not visible in front of dorsal fin.
|
||
</paragraph>
|
||
<paragraph blockId="4.[644,1158,162,1250]" pageId="4" pageNumber="13">
|
||
Dorsal fin hyalineı with a small black blotch at base of simple and first branched raysı followed by two or three blackish blotches on or near base of following raysı space between blotches hyaline; on median third of rays (near branching in branched rays)ı black pigments along edge of rays and between segments. Caudal fin hyalineı on proximal two thirds of principal raysı black pigments along edge of rays and between segments. Anal fin hyalineı with black blotch on base of simple rays; on median third of rays (near branching in branched rays)ı black pigments along edge of rays and between segments. Pelvic and pectoral fins hyaline; on median third of rays (near branching in branched rays)ı black pigments along edge of rays and between segments. In smallest specimen (
|
||
<quantity box="[651,742,1199,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="4" pageNumber="13" unit="mm" value="21.7">21.7 mm</quantity>
|
||
SL;
|
||
<figureCitation box="[795,852,1199,1221]" captionStart="Fig. 2" captionStartId="2.[98,130,1231,1250]" captionTargetBox="[119,1132,177,1182]" captionTargetId="figure@2.[98,1156,432,722]" captionTargetPageId="2" captionText="Fig. 2. Schistura systomos ı CMK 23401 ı paratypes; Laos: Champasak Province: Bolaven Plateau: Tad Set waterfall. aı 35.4 mm SL; bı 21.7 mm SL." httpUri="https://zenodo.org/record/886702/files/figure.png" pageId="4" pageNumber="13">Fig. 2</figureCitation>
|
||
b)ı body greyishı bars more faintly markedı more irregularly shaped.
|
||
</paragraph>
|
||
<paragraph blockId="4.[644,1157,1285,1422]" pageId="4" pageNumber="13">
|
||
<emphasis bold="true" box="[644,839,1285,1307]" pageId="4" pageNumber="13">Notes on biology.</emphasis>
|
||
A dissected female (
|
||
<quantity box="[1065,1156,1285,1307]" metricMagnitude="-2" metricUnit="m" metricValue="3.54" pageId="4" pageNumber="13" unit="mm" value="35.4">35.4 mm</quantity>
|
||
SL) had one ripe ovary with 14 yellow ova about
|
||
<quantity box="[644,765,1341,1364]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="4" pageNumber="13" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2-1.4 mm</quantity>
|
||
diameter and about 31 whitish oocytes. Its stomach and intestine contained remains of unidentified insect larvae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="13" type="distribution">
|
||
<paragraph blockId="4.[644,1157,1458,1624]" pageId="4" pageNumber="13">
|
||
<emphasis bold="true" box="[644,795,1458,1480]" pageId="4" pageNumber="13">Distribution.</emphasis>
|
||
<taxonomicName box="[807,1009,1458,1480]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="13" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[807,1009,1458,1480]" italics="true" pageId="4" pageNumber="13">Schistura systomos</emphasis>
|
||
</taxonomicName>
|
||
is presently known only from the
|
||
<typeStatus box="[890,937,1487,1508]" pageId="4" pageNumber="13">type</typeStatus>
|
||
localityı a waterfall on Xe Setı a tributary of Xe Doneı near edge of Bolaven Plateau (
|
||
<figureCitation box="[825,886,1544,1566]" captionStart="Fig. 5" captionStartId="6.[98,130,824,843]" captionTargetBox="[98,1156,156,809]" captionTargetId="figure@6.[98,1156,156,809]" captionTargetPageId="6" captionText="Fig. 5. Laos: Champasak Province: Bolaven Plateau: Tad Set on Houay Set; type locality of Schistura systomos; 19 January 2013." httpUri="https://zenodo.org/record/886708/files/figure.png" pageId="4" pageNumber="13">Fig. 5</figureCitation>
|
||
). It was collected between stones and blocksı in about
|
||
<quantity box="[929,1040,1572,1595]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="4" pageNumber="13" unit="cm" value="75.0" valueMax="100.0" valueMin="50.0">50-100 cm</quantity>
|
||
depth (dry season) below the waterfall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="14" type="etymology">
|
||
<paragraph blockId="5.[136,148,511,532]" lastBlockId="5.[129,641,676,757]" pageId="5" pageNumber="14">
|
||
<emphasis bold="true" pageId="5" pageNumber="14">a Etymology.</emphasis>
|
||
From the classical Greek σύστοµος (
|
||
<taxonomicName box="[136,236,706,728]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="systomos">systomos</taxonomicName>
|
||
)ı meaning with a narrow mouth. An adjectiveı indeclinable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption httpUri="https://zenodo.org/record/886704/files/figure.png" pageId="5" pageNumber="14" targetBox="[154,1174,181,537]" targetPageId="5">
|
||
<paragraph blockId="5.[128,1187,552,623]" pageId="5" pageNumber="14">
|
||
<emphasis bold="true" box="[129,186,552,571]" pageId="5" pageNumber="14">Fig. 3.</emphasis>
|
||
Ventral view of mouth of
|
||
<taxonomicName box="[446,528,552,571]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis box="[446,528,552,571]" italics="true" pageId="5" pageNumber="14">Schistura</emphasis>
|
||
</taxonomicName>
|
||
species:
|
||
<emphasis bold="true" box="[618,634,552,571]" pageId="5" pageNumber="14">a,</emphasis>
|
||
narrow mouth in
|
||
<taxonomicName box="[813,914,553,572]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[813,914,553,572]" italics="true" pageId="5" pageNumber="14">
|
||
S.
|
||
<taxonomicName box="[836,914,553,571]" pageId="5" pageNumber="14" rank="species" species="systomos">systomos</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
ı MHNG 2767.056ı holotypeı 35.0 mm SL; and
|
||
<emphasis bold="true" box="[300,318,578,597]" pageId="5" pageNumber="14">b,</emphasis>
|
||
wide mouth in other
|
||
<taxonomicName box="[534,616,578,597]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis box="[534,616,578,597]" italics="true" pageId="5" pageNumber="14">Schistura</emphasis>
|
||
</taxonomicName>
|
||
species (here
|
||
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[753,842,578,597]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="clatrata">
|
||
<emphasis box="[753,842,578,597]" italics="true" pageId="5" pageNumber="14">
|
||
S.
|
||
<taxonomicName box="[776,842,578,597]" pageId="5" pageNumber="14" rank="species" species="clatrata">clatrata</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
ı CMK 23398ı 81.5 mm SL; collected together with
|
||
<taxonomicName box="[262,363,604,623]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[262,363,604,623]" italics="true" pageId="5" pageNumber="14">
|
||
S.
|
||
<taxonomicName box="[285,363,604,622]" pageId="5" pageNumber="14" rank="species" species="systomos">systomos</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection lastPageId="6" lastPageNumber="15" pageId="5" pageNumber="14" type="discussion">
|
||
<paragraph blockId="5.[129,641,792,987]" lastBlockId="5.[675,1188,677,1678]" pageId="5" pageNumber="14">
|
||
<emphasis bold="true" box="[129,234,793,814]" pageId="5" pageNumber="14">Remarks.</emphasis>
|
||
<taxonomicName box="[246,444,792,814]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[246,444,792,814]" italics="true" pageId="5" pageNumber="14">Schistura systomos</emphasis>
|
||
</taxonomicName>
|
||
has the smallest mouth in the genus
|
||
<taxonomicName box="[333,425,821,842]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis box="[333,425,821,842]" italics="true" pageId="5" pageNumber="14">Schistura</emphasis>
|
||
</taxonomicName>
|
||
and possibly among loaches of the family
|
||
<taxonomicName box="[353,516,850,872]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
. The mouth occupies only the median half of head width at its location (
|
||
<figureCitation box="[258,316,908,930]" captionStart="Fig. 3" captionStartId="5.[129,161,552,571]" captionTargetBox="[154,1174,181,537]" captionTargetId="figure@5.[661,1186,156,536]" captionTargetPageId="5" captionText="Fig. 3. Ventral view of mouth of Schistura species: aı narrow mouth in S. systomos ı MHNG 2767.056 ı holotypeı 35.0 mm SL; and bı wide mouth in other Schistura species (here S. clatrata ı CMK 23398 ı 81.5 mm SL; collected together with S. systomos)." httpUri="https://zenodo.org/record/886704/files/figure.png" pageId="5" pageNumber="14">Fig. 3</figureCitation>
|
||
a)ı while in the other species it occupies most of head widthı with the lipsı when seen in ventral viewı extending to or almost to the lateral outline of the head (
|
||
<figureCitation box="[951,1009,677,699]" captionStart="Fig. 3" captionStartId="5.[129,161,552,571]" captionTargetBox="[154,1174,181,537]" captionTargetId="figure@5.[661,1186,156,536]" captionTargetPageId="5" captionText="Fig. 3. Ventral view of mouth of Schistura species: aı narrow mouth in S. systomos ı MHNG 2767.056 ı holotypeı 35.0 mm SL; and bı wide mouth in other Schistura species (here S. clatrata ı CMK 23398 ı 81.5 mm SL; collected together with S. systomos)." httpUri="https://zenodo.org/record/886704/files/figure.png" pageId="5" pageNumber="14">Fig. 3</figureCitation>
|
||
b). As mentioned earlierı the genus
|
||
<taxonomicName box="[859,951,706,727]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis box="[859,951,706,727]" italics="true" pageId="5" pageNumber="14">Schistura</emphasis>
|
||
</taxonomicName>
|
||
is very diverse andı as presently understoodı polyphyletic. A number of species group can be distinguished by morphological characters and by colour patternı most of which will likely be recognised as distinct genera. Two species of small size (
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1990" box="[955,1049,850,871]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="geisleri">
|
||
<emphasis box="[955,1049,850,871]" italics="true" pageId="5" pageNumber="14">S. geisleri</emphasis>
|
||
</taxonomicName>
|
||
ı
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1998" box="[1061,1179,850,871]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="isostigma">
|
||
<emphasis box="[1061,1179,850,871]" italics="true" pageId="5" pageNumber="14">S. isostigma</emphasis>
|
||
</taxonomicName>
|
||
) have the mouth somewhat narrower than other
|
||
<taxonomicName box="[675,768,907,928]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis box="[675,768,907,928]" italics="true" pageId="5" pageNumber="14">Schistura</emphasis>
|
||
</taxonomicName>
|
||
(not quantified) but they are distinguished by the pale yellow background and the colour pattern made of a midlateral row of dark brown to black blotches and a row of saddles along dorsal midline (
|
||
<bibRefCitation author="Kottelati M." bookContentInfo="262 pp." box="[908,1059,1022,1044]" journalOrPublisher="Pfeili Muncheni" pageId="5" pageNumber="14" refString="Kottelati M. 1990. Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailandi Burmai Laosi Cambodia and southern Viet Nam. Pfeili Muncheni 262 pp." title="Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailandi Burmai Laosi Cambodia and southern Viet Nam" type="book" year="1990" yearSuffix="i">
|
||
Kottelatı
|
||
<date box="[1005,1059,1023,1044]" pageId="5" pageNumber="14" value="1990-01">1990ı</date>
|
||
</bibRefCitation>
|
||
<bibRefCitation author="Kottelati M." box="[1065,1117,1023,1045]" journalOrPublisher="Ichthyological Exploration of Freshwatersi" pageId="5" pageNumber="14" pagination="1 - 128" part="9" refString="- 1998. Fishes of the Nam Theun and Xe Bangfai basinsi Laosi with diagnoses of twenty-two new species (Teleostei: Cyprinidaei Balitoridaei Cobitidaei Coiidae and Odontobutidae). Ichthyological Exploration of Freshwatersi 9: 1 - 128." title="Fishes of the Nam Theun and Xe Bangfai basinsi Laosi with diagnoses of twenty-two new species (Teleostei: Cyprinidaei Balitoridaei Cobitidaei Coiidae and Odontobutidae)" type="journal article" year="1998">1998</bibRefCitation>
|
||
). They also have a suborbital flap in male that is lacking in
|
||
<taxonomicName box="[702,816,1080,1102]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[702,816,1080,1102]" italics="true" pageId="5" pageNumber="14">S. systomos</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[675,1188,677,1678]" lastBlockId="6.[98,611,923,1435]" lastPageId="6" lastPageNumber="15" pageId="5" pageNumber="14">
|
||
Similarlyı two species of
|
||
<taxonomicName box="[1029,1188,1109,1130]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis box="[1029,1188,1109,1130]" italics="true" pageId="5" pageNumber="14">Physoschistura</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName baseAuthorityName="Annandale" baseAuthorityYear="1918" box="[683,813,1138,1159]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="brunneana">
|
||
<emphasis box="[683,813,1138,1159]" italics="true" pageId="5" pageNumber="14">P. brunneana</emphasis>
|
||
</taxonomicName>
|
||
ı
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1990" box="[828,1026,1138,1159]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="pseudobrunneana">
|
||
<emphasis box="[828,1026,1138,1159]" italics="true" pageId="5" pageNumber="14">P. pseudobrunneana</emphasis>
|
||
</taxonomicName>
|
||
) also have the mouth somewhat narrower than species of
|
||
<taxonomicName class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis italics="true" pageId="5" pageNumber="14">Schistura</emphasis>
|
||
</taxonomicName>
|
||
(not quantified)ı slightly compressed and deep body and headı and the eyes partly or almost visible in ventral view;
|
||
<taxonomicName box="[896,1010,1253,1275]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[896,1010,1253,1275]" italics="true" pageId="5" pageNumber="14">S. systomos</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by the absence of a posterior chamber of the air bladder in the body cavity (vs. presence)ı the absence of a suborbital flap in male (vs. presence)ı the arched mouth (vs. U-shaped)ı and the lower lip with V-shaped median interruptionı the median part with two or three sulciı the lateral part shortı with a few shallow furrows (vs. with wide median interruption forming two lateral triangular pads with deep sulci).
|
||
<taxonomicName box="[858,1005,1512,1533]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis box="[858,1005,1512,1533]" italics="true" pageId="5" pageNumber="14">Physoschistura</emphasis>
|
||
</taxonomicName>
|
||
is another genus to which a number of species have been added in recent years (Chen et al.
|
||
<date box="[964,1029,1570,1592]" pageId="5" pageNumber="14" value="2011-01">ı 2011</date>
|
||
;
|
||
<bibRefCitation author="Lokeshwori Y. & W. Vishwanath" journalOrPublisher="Zootaxai" pageId="5" pageNumber="14" pagination="95 - 102" part="3586" refString="Lokeshwori Y. & W. Vishwanath. 2012 a. A new loach of the genus Physoschistura Banarescu & Nalbant (Teleostei: Nemacheilidae) from Chindwin basini Manipuri India. Zootaxai 3586: 95 - 102." title="A new loach of the genus Physoschistura Banarescu & Nalbant (Teleostei: Nemacheilidae) from Chindwin basini Manipuri India" type="journal article" year="2012">Lokeshwor & Vishwanathı 2012</bibRefCitation>
|
||
a- b;
|
||
<bibRefCitation author="Suvarnarakshai A." box="[912,1132,1598,1621]" journalOrPublisher="Zootaxai" pageId="5" pageNumber="14" pagination="236 - 248" part="3736" refString="Suvarnarakshai A. 2013. A new species of Physoschistura (Pisces: Nemacheilidae) from northern Thailand. Zootaxai 3736: 236 - 248." title="A new species of Physoschistura (Pisces: Nemacheilidae) from northern Thailand" type="journal article" year="2013">Suvarnarakshaı 2013</bibRefCitation>
|
||
) that do not seem related to the
|
||
<typeStatus box="[961,1008,1628,1649]" pageId="5" pageNumber="14">type</typeStatus>
|
||
species (
|
||
<taxonomicName baseAuthorityName="Annandale" baseAuthorityYear="1918" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="brunneana">
|
||
<emphasis italics="true" pageId="5" pageNumber="14">P. brunneana</emphasis>
|
||
</taxonomicName>
|
||
) and seem to represent one or more distinct genera. A number of species placed in
|
||
<taxonomicName box="[485,575,924,945]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis box="[485,575,924,945]" italics="true" pageId="6" pageNumber="15">Schistura</emphasis>
|
||
</taxonomicName>
|
||
are apparently also related to these
|
||
<emphasis box="[441,604,952,973]" italics="true" pageId="6" pageNumber="15">‘Physoschistura’</emphasis>
|
||
ı for example
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1990" box="[232,326,981,1002]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="species" species="geisleri">
|
||
<emphasis box="[232,326,981,1002]" italics="true" pageId="6" pageNumber="15">S. geisleri</emphasis>
|
||
</taxonomicName>
|
||
ı
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1998" box="[339,456,981,1003]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="species" species="isostigma">
|
||
<emphasis box="[339,456,981,1003]" italics="true" pageId="6" pageNumber="15">S. isostigma</emphasis>
|
||
</taxonomicName>
|
||
ı
|
||
<taxonomicName authorityName="Kottelat" authorityYear="1990" box="[469,610,981,1003]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="species" species="maepaiensis">
|
||
<emphasis box="[469,610,981,1003]" italics="true" pageId="6" pageNumber="15">S. maepaiensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="S. Q. Zhu & S. H. Wang" baseAuthorityYear="1985" box="[152,349,1010,1031]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="species" species="shuangjiangensis">
|
||
<emphasis box="[152,349,1010,1031]" italics="true" pageId="6" pageNumber="15">S. shuangjiangensis</emphasis>
|
||
</taxonomicName>
|
||
. There is no character suggesting that
|
||
<taxonomicName box="[265,378,1039,1060]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[265,378,1039,1060]" italics="true" pageId="6" pageNumber="15">S. systomos</emphasis>
|
||
</taxonomicName>
|
||
could be more closely related to any of these
|
||
<emphasis box="[328,491,1068,1089]" italics="true" pageId="6" pageNumber="15">‘Physoschistura’</emphasis>
|
||
rather than to
|
||
<emphasis box="[124,232,1096,1117]" italics="true" pageId="6" pageNumber="15">‘Schistura’</emphasis>
|
||
. In factı
|
||
<taxonomicName box="[319,432,1097,1118]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[319,432,1097,1118]" italics="true" pageId="6" pageNumber="15">S. systomos</emphasis>
|
||
</taxonomicName>
|
||
likely represents a genus of its own. It relationships will be addressed in a forthcoming review of the genera of Southeast Asian
|
||
<taxonomicName box="[304,467,1183,1205]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/886706/files/figure.png" pageId="5" pageNumber="14" targetBox="[222,546,1038,1613]" targetPageId="5">
|
||
<paragraph blockId="5.[128,640,1632,1677]" pageId="5" pageNumber="14">
|
||
<emphasis bold="true" box="[128,191,1632,1651]" pageId="5" pageNumber="14">Fig. 4.</emphasis>
|
||
<taxonomicName box="[202,386,1632,1651]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[202,386,1632,1651]" italics="true" pageId="5" pageNumber="14">
|
||
<taxonomicName box="[202,290,1632,1651]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="14" phylum="Chordata" rank="genus">Schistura</taxonomicName>
|
||
<taxonomicName box="[303,386,1632,1650]" pageId="5" pageNumber="14" rank="species" species="systomos">systomos</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
ı CMK 23401ı paratypeı 35.4 mm SL; digestive tract. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption httpUri="https://zenodo.org/record/886708/files/figure.png" pageId="6" pageNumber="15" targetBox="[98,1156,156,809]" targetPageId="6">
|
||
<paragraph blockId="6.[98,1156,824,869]" pageId="6" pageNumber="15">
|
||
<emphasis bold="true" box="[98,151,824,844]" pageId="6" pageNumber="15">Fig. 5</emphasis>
|
||
. Laos: Champasak Province: Bolaven Plateau: Tad Set on Houay Set; type locality of
|
||
<taxonomicName box="[983,1151,824,843]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[983,1151,824,843]" italics="true" pageId="6" pageNumber="15">Schistura systomos</emphasis>
|
||
</taxonomicName>
|
||
; 19 January 2013.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[98,611,923,1435]" pageId="6" pageNumber="15">
|
||
Compared to other nemacheilidsı the stomach of
|
||
<taxonomicName box="[123,235,1241,1262]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="15" phylum="Chordata" rank="species" species="systomos">
|
||
<emphasis box="[123,235,1241,1262]" italics="true" pageId="6" pageNumber="15">S. systomos</emphasis>
|
||
</taxonomicName>
|
||
appears proportionally smaller (not quantified) and the intestine is very thin and straight. In the great majority of nemacheilidsı the intestine has at least one conspicuous bendı a loop or a complex coiling. The stomach and intestine of the two dissected specimens contained remains of insect larvae that could not be identified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |