128 lines
14 KiB
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128 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.544.6074" ID-GBIF-Dataset="0e5e47a0-f8f3-4039-b0e1-c159fef0b9f6" ID-PMC="PMC4714384" ID-Pensoft-Pub="1313-2970-544-1" ID-PubMed="26798289" ID-ZBK="C5978BD0145B40F8ACDEB27371B7B9A4" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-544-1" ModsDocOrigin="ZooKeys 544" ModsDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" checkinTime="1451243640953" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2015" docId="89859D9006B69871108F2B9945CC6501" docLanguage="en" docName="ZooKeys 544: 1-407" docOrigin="ZooKeys 544" docSource="http://dx.doi.org/10.3897/zookeys.544.6074" docTitle="Mecyclothorax ommatoplax Liebherr, 2015, sp. n." docType="treatment" docUuid="E16F0AD3-9ACF-472E-9D4F-BB4A869C0601" docUuidSource="ZooBank" docVersion="5" lastPageNumber="155" masterDocId="FF92FFC2FF9FFFC7191EFFB8FF91AF5D" masterDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" masterLastPageNumber="407" masterPageNumber="1" pageNumber="153" updateTime="1668162300918" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>544</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>407</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.544.6074</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.544.6074</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-544-1</mods:identifier>
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<mods:identifier type="ZBK">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
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<mods:identifier type="ZooBank">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127853589" LSID="urn:lsid:zoobank.org:act:E16F0AD3-9ACF-472E-9D4F-BB4A869C0601" httpUri="http://treatment.plazi.org/id/89859D9006B69871108F2B9945CC6501" lastPageId="154" lastPageNumber="155" pageId="152" pageNumber="153">
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<subSubSection pageId="152" pageNumber="153" type="multiple">
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<paragraph pageId="152" pageNumber="153">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection pageId="152" pageNumber="153" type="nomenclature">
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<paragraph pageId="152" pageNumber="153">
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(062)
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<taxonomicName LSID="http://zoobank.org/E16F0AD3-9ACF-472E-9D4F-BB4A869C0601" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax ommatoplax" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="ommatoplax">Mecyclothorax ommatoplax</taxonomicName>
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<taxonomicNameLabel pageId="152" pageNumber="153">sp. n.</taxonomicNameLabel>
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Figs 81A, 85
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</paragraph>
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</subSubSection>
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<subSubSection pageId="152" pageNumber="153" type="diagnosis">
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<paragraph pageId="152" pageNumber="153">Diagnosis.</paragraph>
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<paragraph pageId="152" pageNumber="153">
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This species is easily distinguished by the very small, little convex eyes; ocular ratio = 1.29-1.35, ocular lobe ratio = 0.68-0.69, in combination with the granulate isodiametric elytral microsculpture (Fig. 81A). Setal formula 2 2 2 2. Among species of the
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax argutor" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="argutor">Mecyclothorax argutor</taxonomicName>
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group, the beetles are of moderate size, standardized body length 4.8-5.5 mm.
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</paragraph>
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<caption pageId="152" pageNumber="153">
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<paragraph pageId="152" pageNumber="153">
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Figure 81.
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax argutor" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="argutor">Mecyclothorax argutor</taxonomicName>
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group species, dorsal habitus view. A
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax ommatoplax" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="ommatoplax">Mecyclothorax ommatoplax</taxonomicName>
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(Kīpahulu, 1500 m) B
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax semistriatus" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="semistriatus">Mecyclothorax semistriatus</taxonomicName>
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(Honomanu, 1950 m) C
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax refulgens" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="refulgens">Mecyclothorax refulgens</taxonomicName>
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(Kīpahulu, 1860 m) D
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax argutulus" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="argutulus">Mecyclothorax argutulus</taxonomicName>
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(Kuhiwa, 1615 m).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="153" lastPageNumber="154" pageId="152" pageNumber="153" type="description">
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<paragraph pageId="152" pageNumber="153">Description</paragraph>
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<paragraph lastPageId="153" lastPageNumber="154" pageId="152" pageNumber="153">
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(n = 2). Head capsule with frontal grooves broad near clypeus, curved laterally near clypeal juncture, a straight ridgelike lateral carina extended from dorsal mandibular articulation to mesad anterior supraorbital seta; dorsal surface of neck flat to slightly convex; compound eye with approximately 12-13 ommatidia across the maximal horizontal diameter; labral anterior margin medially emarginate 0.2
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<normalizedToken originalValue="×">x</normalizedToken>
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length; antennae filiform, antennomere 3 with sparse, short setae; mentum tooth with sides acute, apex pointed. Pronotum slightly transverse, MPW/PL = 1.17-1.27, constricted basally, MPW/BPW = 1.51-1.58; hind angle right, apex rounded, lateral margin parallel anterad angle; median base depressed relative to disc, covered with rugose wrinkles and round punctures; basal margin broadly, medially extended between laterobasal depressions; median longitudinal impression fine, shallow, crossed by shallow wrin
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<pageBreakToken pageId="153" pageNumber="154" start="start">kles</pageBreakToken>
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; anterior transverse impression broad, shallow, crossed by longitudinal wrinkles; anterior callosity slightly convex, crossed by longitudinal wrinkles from anterior impression; front angles not projected, rounded; pronotal anterior width slightly greater than basal width, APW/BPW = 1.02-1.05; lateral marginal depression narrow, margin upturned to beaded; laterobasal depression moderately broad with low oblique ridge terminated at hind angle. Proepisternum with 6 punctures along hind marginal groove; prosternal process medially concave, margin smoothly upraised. Elytra subquadrate, narrow basally, the basal groove meeting lateral marginal depression at subangulate humerus, MEW/HuW = 1.81-1.94; disc flat, sides moderately depressed; parascutellar seta present; parascutellar striole shallow, smooth, continued onto base; sutural interval slightly more convex than lateral intervals; sutural and 2nd striae of subequal depth from base to apex; discal striae 1-5 continuous, slightly irregular but impunctate, deeper and smooth apically, stria 6 reduced, stria 7 traceable, very shallow, associated intervals broadly, slightly convex; 8th interval slightly convex laterad 7th stria, principally due to deep apical portion of 7th stria; 2 dorsal elytral setae at 0.30
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<normalizedToken originalValue="×">x</normalizedToken>
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and 0.59
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<normalizedToken originalValue="–0.60×">-0.60x</normalizedToken>
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elytral length, setal impressions shallow, about breadth of interval 3; apical and subapical setae present; lateral elytral setae arranged as anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression moderately broad, lined with sculpticells, the margin upturned; subapical sinuation evident, abrupt and brief. Mesepisternum with ~14 punctures in 2-3 rows, metepisternal width to length ratio = 0.67; metepisternum/metepimeron suture distinct. Abdomen with shallow indistinct wrinkles on ventrites 1-3, suture between ventrites 2 and 3 complete; apical female ventrite with 4 equally spaced setae and median trapezoid of 4(-5 unilaterally) subequal, short setae. Legs-metatarsomere 1/metatibial length ratio = 0.19; metatarsomere 4 length along outer lobe 1.4
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<normalizedToken originalValue="×">x</normalizedToken>
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medial tarsomere length, with apical and subapical setae; metatarsal dorsolateral sulci broad, shallow. Microsculpture of vertex an upraised isodiametric mesh in transverse rows; pronotal disc with transverse mesh, median base with granulate isodiametric mesh; elytral apex with distinct transverse mesh; metasternum with distinct transverse mesh; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufobrunneous; antennomere 1 flavous, 2-11 rufoflavous; pronotal disc rufobrunneous, lateral margins narrowly paler in lateral depression, rufoflavous; proepipleuron rufoflavous, proepisternum rufobrunneous with piceous cast; elytral disc reflective rufobrunneous, sutural interval basally rufous, apically flavous, elytral margins broadly rufoflavous, apex broadly rufoflavous to flavous; elytral epipleuron flavous, metepisternum rufobrunneous with piceous cast; abdomen rufobrunneous with piceous cast, apical half of ventrite 6 broadly paler, flavous; metafemur flavous; metatibia rufoflavous.
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</paragraph>
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<paragraph pageId="153" pageNumber="154">Female reproductive tract. The holotype and paratype females were not dissected.</paragraph>
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</subSubSection>
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<subSubSection pageId="153" pageNumber="154" type="holotype">
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<paragraph pageId="153" pageNumber="154">Holotype.</paragraph>
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<paragraph pageId="153" pageNumber="154">
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Female (CUIC) labeled: HI:Maui Haleakala N.P. / Kipahulu Vy.1500 m el. / 9-V-1991 sifting / leaf litter by day // S. Jessel / A.C. Medeiros, / Jr. collectors //
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
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/
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<taxonomicName lsidName="ommatoplax" pageId="153" pageNumber="154" rank="species" species="ommatoplax">ommatoplax</taxonomicName>
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/ ♀ photo / det. J.K. Liebherr 2014 // HOLOTYPE /
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
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/
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<taxonomicName lsidName="ommatoplax" pageId="153" pageNumber="154" rank="species" species="ommatoplax">ommatoplax</taxonomicName>
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/ Liebherr / det. J.K. Liebherr 2015 (black-margined red label).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="154" pageNumber="155" type="paratype">
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<paragraph pageId="154" pageNumber="155">
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<pageBreakToken pageId="154" pageNumber="155" start="start">Paratype</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="154" pageNumber="155">HI: Maui, Kipahulu Vy., West Camp at Dead Pig Bog, 2010 m el., 18-iii-1998, Takumi (BPBM, 1).</paragraph>
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</subSubSection>
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<subSubSection pageId="154" pageNumber="155" type="etymology">
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<paragraph pageId="154" pageNumber="155">Etymology.</paragraph>
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<paragraph pageId="154" pageNumber="155">
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The species epithet compounds the Greek words plax, flat round plate, and ommatos, the genitive of omma, i.e. eye. As Greek,
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<taxonomicName lsidName="ommatoplax" pageId="154" pageNumber="155" rank="species" species="ommatoplax">ommatoplax</taxonomicName>
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is treated as a noun. The epithet refers to the very flat eyes characteristic of this species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="154" pageNumber="155" type="distribution and habitat">
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<paragraph pageId="154" pageNumber="155">Distribution and habitat.</paragraph>
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<paragraph pageId="154" pageNumber="155">
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax ommatoplax" order="Coleoptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="ommatoplax">Mecyclothorax ommatoplax</taxonomicName>
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is distributed in the upper reaches of
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<normalizedToken originalValue="Kīpahulu">Kīpahulu</normalizedToken>
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Valley (Fig. 85). Specimens have been collected in sift samples of leaf and humus litter.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |