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<mods:title id="0A92195894ADECD2DAACB50FAFD9F2B4">Three new Chaerilus from Malaysia (Tioman Island) and Thailand (Scorpiones Chaerilidae) with a review of C cimrmani C sejnai and C tichyi</mods:title>
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<taxonomicName id="26DD4D0502028F41FC12FB8CFB42D1EC" ID-CoL="T9ZJ" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[992,1214,1149,1176]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">Chaerilus majkusi</taxonomicName>
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(
<figureCitation id="79E62A0302028F41FC9FFB6EFC21D1C2" box="[877,989,1183,1206]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–12" captionStart-2="Figures 13–20" captionStart-3="Figures 21–32" captionStart-4="Figures 33–36" captionStartId-0="4.[192,278,1612,1635]" captionStartId-1="5.[192,278,1754,1777]" captionStartId-2="6.[192,278,1627,1650]" captionStartId-3="7.[192,278,1343,1366]" captionStartId-4="8.[192,278,921,944]" captionTargetBox-0="[206,1423,198,1574]" captionTargetBox-1="[191,1440,191,1716]" captionTargetBox-2="[192,1439,204,1588]" captionTargetBox-3="[215,1420,199,1311]" captionTargetBox-4="[199,1426,199,856]" captionTargetId-0="figure-19@4.[192,1439,192,1582]" captionTargetId-1="figure-6@5.[192,1439,192,1715]" captionTargetId-2="figure-19@6.[192,1439,192,1597]" captionTargetId-3="figure-6@7.[192,1437,192,1313]" captionTargetId-4="figure-19@8.[192,1437,192,860]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="Figures 1–4: Chaerilus majkusi sp. n. Figures 1–2. Holotype male, dorsal (1) and ventral (2) views. Figures 3–4. Paratype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm." captionText-1="Figures 5–12: Chaerilus majkusi sp. n. Figures 5, 7, 9–12. Holotype male, carapace and tergites I–IV (5), sternopectinal region and sternites (7), left legs I–IV, retrolateral aspect (9–12). Figures 6, 8. Paratype female, carapace and tergites I–III (6), sternopectinal region and sternites III–V (8)." captionText-2="Figures 13–20: Chaerilus majkusi sp. n. Figures 13, 15–17. Holotype male, metasoma V and telson lateral (13), metasoma and telson, lateral (15), ventral (16), and dorsal (17) views. Figures 14, 18–20. Paratype female, metasoma V and telson lateral (14), metasoma and telson, lateral (18), ventral (19), and dorsal (20) views. Scale bar: 10 mm (15–20)." captionText-3="Figures 21–32: Chaerilus majkusi sp. n. Figures 21–24. Paratype female, pedipalp chela, dorsal (21) and externodorsal (22) views, movable (23) and fixed (24) fingers. Figures 25–32. Holotype male, pedipalp chela, dorsal (25), externodorsal (26), and ventrointernal (27) views. Pedipalp patella, dorsal (28), external (29) and ventral (30) views. Pedipalp femur and trochanter, dorsal (31) and ventral (32) views. The trichobothrial pattern is indicated in Figures 25–31." captionText-4="Figures 33–36: Figures 33–34. Chaerilus majkusi sp. n., male paratype, left chelicera dorsal (33) and ventral (34) views. Figures 35–36. Chaerilus stockmannorum sp. n., male paratype, right chelicera dorsal (35) and ventral (36) views." pageId="3" pageNumber="2">Figs. 1–34</figureCitation>
, 
<figureCitation id="79E62A0302028F41FC18FB6EFBFBD1C2" box="[1002,1031,1183,1206]" captionStart="Figures 37–38" captionStartId="9.[192,278,1830,1853]" captionTargetBox="[191,1440,212,1801]" captionTargetId="figure-42@9.[192,1439,213,1800]" captionTargetPageId="9" captionText="Figures 37–38: Chaerilus majkusi sp. n., female paratype (37) and Chaerilus sejnai Kovařík, 2005, male (38) in vivo habitus." pageId="3" pageNumber="2">37</figureCitation>
, 
<figureCitation id="79E62A0302028F41FBE0FB6EFBA9D1C2" box="[1042,1109,1183,1206]" captionStart="Figures 39–40" captionStartId="10.[192,278,1696,1719]" captionTargetBox="[191,1440,167,1666]" captionTargetId="figure-19@10.[192,1439,168,1666]" captionTargetPageId="10" captionText="Figures 39–40: Chaerilus majkusi sp. n., male holotype (39) and female paratype with newborns (40) in vivo habitus." pageId="3" pageNumber="2">39–40</figureCitation>
, 
<figureCitation id="79E62A0302028F41FB90FB6EFB81D1C2" box="[1122,1149,1183,1206]" captionStart="Figures 43–44" captionStartId="12.[192,278,1767,1790]" captionTargetBox="[191,1440,199,1729]" captionTargetId="figure-75@12.[192,1439,200,1728]" captionTargetPageId="12" captionText="Figures 43–44: Chaerilus, collection localities. Figure 43. C. majkusi sp. n., type locality. Figure 44. C. neradorum sp. n., type locality." pageId="3" pageNumber="2">43</figureCitation>
, 
<figureCitation id="79E62A0302028F41FB79FB6EFB4FD1C2" box="[1163,1203,1183,1206]" captionStart="Figures 108–114" captionStartId="23.[192,278,1730,1753]" captionTargetBox="[191,1440,191,1701]" captionTargetId="figure-6@23.[192,1439,192,1700]" captionTargetPageId="23" captionText="Figures 108–114: Hemispermatophores of Chaerilus species. Figure 108. C.cimrmani. Figure 109. C.majkusi sp. n., male holotype. Figure 110. C.tichyi. Figures 111–112. C.stockmannorum sp. n., male holotype. Figures 113–114. C.hofereki, samples from two different individuals, capsule region and distal lamina (113) and whole hemispermatophore (114, reproduced from Kovařík et al., 2015). Convex views shown in all figures except Fig 112 (anterior view). Left hemispermatophore shown in all figures except for Fig. 108, which is right hemispermatophore shown in mirror image for comparison. Scale bars: 500 µm (108–109, 113–114), 1 mm (110–112)." pageId="3" pageNumber="2">109</figureCitation>
, 
<figureCitation id="79E62A0302028F41FB32FB6EFAE1D1C2" box="[1216,1309,1183,1206]" captionStart="Figures 115–123" captionStartId="24.[192,278,1474,1497]" captionTargetBox="[191,1437,191,1445]" captionTargetId="figure-383@24.[192,1437,192,1444]" captionTargetPageId="24" captionText="Figures 115–123: The chromosomes of Chaerilus species. Figures 115–116. C. majkusi sp. n., mitotic metaphase (115) and meiotic metaphase I (116). Figures 117–118. C. neradorum sp. n., mitotic metaphase (117) and meiotic metaphase I (118). Figures 119–120. C. stockmannorum sp. n., mitotic metaphase (119) and meiotic metaphase I (120), arrows show chromosomes in pentavalent and arrowheads show chromosomes in trivalent. Figure 121. Mitotic metaphase of C. cimrmani. Figure 122. Meiotic metaphase I of C. sejnai. Figure 123. Mitotic metaphase of C. tichyi. Scale bar: 10 µm." pageId="3" pageNumber="2">115–116</figureCitation>
, 
<tableCitation id="AC5F033D02028F41FAD9FB6EFA86D1C2" box="[1323,1402,1183,1206]" captionStart="Table 1" captionStartId="17.[192,257,818,841]" captionText="Table 1: Comparative measurements of Chaerilus majkusi sp. n. and Chaerilus neradorum sp. n. types. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="3" pageNumber="2">Table 1</tableCitation>
) 
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<specimenCount id="F7DBFD0F02028F41FC21FA00FC08D37C" box="[979,1012,1521,1544]" pageId="3" pageNumber="2" type="male">1♂</specimenCount>
juv (
<typeStatus id="3E66882402028F41FBDBFA00FB6DD37C" box="[1065,1169,1521,1544]" pageId="3" pageNumber="2" type="paratype">paratypes</typeStatus>
), leg. 
<collectorName id="4C28535002028F41FB28FA00FAA2D37C" box="[1242,1374,1521,1544]" pageId="3" pageNumber="2">D. Ho-ferek</collectorName>
</materialsCitation>
&amp; 
<materialsCitation id="51B53CDB02028F41FA7AFA00FB74D310" collectionCode="R" collectorName="D. Hoferek &amp; F. Kovarik" country="Malaysia" location="Majkus" pageId="3" pageNumber="2" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<collectionCode id="87CCAE4302028F41FA7AFA00FA66D37C" box="[1416,1434,1521,1544]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="2">R</collectionCode>
. 
<location id="E402605D02028F41FCBAF9FEFC64D352" LSID="urn:lsid:plazi:treatment:6974879002028F4AFC12FB8CFB36D3F6:E402605D02028F41FCBAF9FEFC64D352" box="[840,920,1551,1574]" country="Malaysia" name="Majkus" pageId="3" pageNumber="2">Majkus</location>
, 
<specimenCount id="F7DBFD0F02028F41FC55F9FEFC34D352" box="[935,968,1551,1574]" pageId="3" pageNumber="2" type="male">3♂</specimenCount>
<specimenCount id="F7DBFD0F02028F41FC3AF9FEFC15D352" box="[968,1001,1551,1574]" pageId="3" pageNumber="2" type="female">2♀</specimenCount>
8 juvs. (
<typeStatus id="3E66882402028F41FBB1F9FEFB50D352" box="[1091,1196,1551,1574]" pageId="3" pageNumber="2" type="paratype">paratypes</typeStatus>
, 1454–1456, offspring of the female 
<typeStatus id="3E66882402028F41FC16F9DFFBBCD331" box="[996,1088,1582,1605]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
, some of them still alive), bred by 
<collectorName id="4C28535002028F41FC9BF9BCFC1DD310" box="[873,993,1613,1636]" pageId="3" pageNumber="2">D. Hoferek</collectorName>
and 
<collectorName id="4C28535002028F41FBE7F9BCFB74D310" box="[1045,1160,1613,1636]" pageId="3" pageNumber="2">F. Kovařík</collectorName>
</materialsCitation>
, 
<collectionCode id="87CCAE4302028F41FB64F9BCFB20D310" box="[1174,1244,1613,1636]" pageId="3" pageNumber="2">FKCP</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="A9C7650D02028F41FCBAF976FBDDD3AF" pageId="3" pageNumber="2" type="etymology">
<paragraph id="E162368602028F41FCBAF976FBDDD3AF" blockId="3.[840,1440,1671,1755]" pageId="3" pageNumber="2">
ETYMOLOGY. A patronym in honor of Zdeněk Majkus, 
<collectingCountry id="99CA761602028F41FCB8F957FC07D3C9" box="[842,1019,1702,1725]" name="Czech Republic" pageId="3" pageNumber="2">Czech Republic</collectingCountry>
, for his friendship and lifelong dedication to arachnids.
</paragraph>
</subSubSection>
<subSubSection id="A9C7650D02028F46FCBAF90FFAF9D271" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="diagnosis">
<paragraph id="E162368602028F46FCBAF90FFAF9D271" blockId="3.[840,1441,1790,1905]" lastBlockId="4.[840,1440,1712,1797]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
DIAGNOSIS. Total length 
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. Two pairs of lateral eyes and one pair of median eyes. Male differs from female in having pedipalp chela much wider. Chela length/width ratio in male 1.87; in female 2.28. Movable finger of pedipalp with 16 imbricated granule rows. Fingers straight in both sexes. Chela of pedipalp with 7–8 carinae. Pectinal teeth number 
<quantity id="26259B6302058F46FD80F91FFD60D271" box="[626,668,1774,1797]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="3" unit="in" value="6.0">6 in</quantity>
males, 
<quantity id="26259B6302058F46FD1DF91FFCE4D271" box="[751,792,1774,1797]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="3" unit="in" value="4.0">4 in</quantity>
females. Carapace granulated. Mesosomal tergites granulated mainly in median area. All sternites smooth without carinae and granules. First metasomal segment with 6 or 8 carinae, second to fourth segments with 6 carinae, ventral metasomal carinae absent. All metasomal segments very sparsely granulated.
</paragraph>
</subSubSection>
<caption id="B5A2660E02058F46FF32F9BDFD0BD30F" pageId="4" pageNumber="3" startId="4.[192,278,1612,1635]" targetBox="[206,1423,198,1574]" targetPageId="4">
<paragraph id="E162368602058F46FF32F9BDFD0BD30F" blockId="4.[192,1440,1612,1659]" pageId="4" pageNumber="3">
<emphasis id="D3A9EA9402058F46FF32F9BDFEACD317" bold="true" box="[192,336,1612,1635]" pageId="4" pageNumber="3">Figures 1–4:</emphasis>
<taxonomicName id="26DD4D0502058F46FEAAF9BFFDF6D317" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[344,522,1614,1635]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
<emphasis id="D3A9EA9402058F46FEAAF9BFFDF6D317" box="[344,522,1614,1635]" italics="true" pageId="4" pageNumber="3">Chaerilus majkusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C89A57EF02058F46FDE1F9BFFDB7D317" box="[531,587,1614,1635]" pageId="4" pageNumber="3" rank="species">
<emphasis id="D3A9EA9402058F46FDE1F9BFFDD6D317" bold="true" box="[531,554,1614,1635]" pageId="4" pageNumber="3">sp</emphasis>
. 
<emphasis id="D3A9EA9402058F46FDCAF9BFFDB9D317" bold="true" box="[568,581,1614,1635]" pageId="4" pageNumber="3">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="D3A9EA9402058F46FDA6F9BFFD31D317" bold="true" box="[596,717,1614,1635]" pageId="4" pageNumber="3">Figures 1–2</emphasis>
. Holotype male, dorsal (1) and ventral (2) views. 
<emphasis id="D3A9EA9402058F46FB36F9BFFAC2D317" bold="true" box="[1220,1342,1614,1635]" pageId="4" pageNumber="3">Figures 3–4</emphasis>
. Paratype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm.
</paragraph>
</caption>
<subSubSection id="A9C7650D02058F4AFCBAF8DAFB36D3F6" lastPageId="8" lastPageNumber="7" pageId="4" pageNumber="3" type="description">
<paragraph id="E162368602058F46FCBAF8DAFA66D215" blockId="4.[840,1441,1835,1889]" pageId="4" pageNumber="3">
DESCRIPTION. Total length 
<quantity id="26259B6302058F46FB81F8DAFB1AD236" box="[1139,1254,1835,1858]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="4" pageNumber="3" unit="mm" value="27.0" valueMax="29.0" valueMin="25.0">25–29 mm</quantity>
. Two well developed pairs of lateral eyes and one pair of median eyes
</paragraph>
<caption id="B5A2660E02048F47FF32F92BFD90D257" pageId="5" pageNumber="4" startId="5.[192,278,1754,1777]" targetBox="[191,1440,191,1716]" targetPageId="5">
<paragraph id="E162368602048F47FF32F92BFD90D257" blockId="5.[192,1440,1754,1827]" pageId="5" pageNumber="4">
<emphasis id="D3A9EA9402048F47FF32F92BFEA7D385" bold="true" box="[192,347,1754,1777]" pageId="5" pageNumber="4">Figures 5–12:</emphasis>
<taxonomicName id="26DD4D0502048F47FE93F92DFDECD385" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[353,528,1756,1777]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
<emphasis id="D3A9EA9402048F47FE93F92DFDECD385" box="[353,528,1756,1777]" italics="true" pageId="5" pageNumber="4">Chaerilus majkusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C89A57EF02048F47FDE4F92DFDB0D385" box="[534,588,1756,1777]" pageId="5" pageNumber="4" rank="species">
<emphasis id="D3A9EA9402048F47FDE4F92DFDD1D385" bold="true" box="[534,557,1756,1777]" pageId="5" pageNumber="4">sp</emphasis>
. 
<emphasis id="D3A9EA9402048F47FDCBF92DFDBAD385" bold="true" box="[569,582,1756,1777]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="D3A9EA9402048F47FDA0F92DFD4ED385" bold="true" box="[594,690,1756,1777]" pageId="5" pageNumber="4">Figures 5</emphasis>
, 
<emphasis id="D3A9EA9402048F47FD4CF92DFD36D385" bold="true" box="[702,714,1756,1777]" pageId="5" pageNumber="4">7</emphasis>
, 
<emphasis id="D3A9EA9402048F47FD24F92DFCFAD385" bold="true" box="[726,774,1756,1777]" pageId="5" pageNumber="4">9–12</emphasis>
. Holotype male, carapace and tergites I–IV (5), sternopectinal region and sternites (7), left legs I–IV, retrolateral aspect (9–12). 
<emphasis id="D3A9EA9402048F47FCD3F904FC7AD27E" bold="true" box="[801,902,1781,1802]" pageId="5" pageNumber="4">Figures 6</emphasis>
, 
<emphasis id="D3A9EA9402048F47FC6AF904FC58D27E" bold="true" box="[920,932,1781,1802]" pageId="5" pageNumber="4">8</emphasis>
. Paratype female, carapace and tergites I–III (6), sternopectinal region and sternites III–V (8).
</paragraph>
</caption>
<caption id="B5A2660E02078F44FF32F9AAFB99D3D0" pageId="6" pageNumber="5" startId="6.[192,278,1627,1650]" targetBox="[192,1439,204,1588]" targetPageId="6">
<paragraph id="E162368602078F44FF32F9AAFB99D3D0" blockId="6.[192,1440,1627,1700]" pageId="6" pageNumber="5">
<emphasis id="D3A9EA9402078F44FF32F9AAFE94D306" bold="true" box="[192,360,1627,1650]" pageId="6" pageNumber="5">Figures 13–20:</emphasis>
<taxonomicName id="26DD4D0502078F44FE9CF9ACFDE2D306" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[366,542,1629,1650]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
<emphasis id="D3A9EA9402078F44FE9CF9ACFDE2D306" box="[366,542,1629,1650]" italics="true" pageId="6" pageNumber="5">Chaerilus majkusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C89A57EF02078F44FDD6F9ACFDA6D306" box="[548,602,1629,1650]" pageId="6" pageNumber="5" rank="species">
<emphasis id="D3A9EA9402078F44FDD6F9ACFDC7D306" bold="true" box="[548,571,1629,1650]" pageId="6" pageNumber="5">sp</emphasis>
. 
<emphasis id="D3A9EA9402078F44FDB4F9ACFDAFD306" bold="true" box="[582,595,1629,1650]" pageId="6" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="D3A9EA9402078F44FD92F9ACFD37D306" bold="true" box="[608,715,1629,1650]" pageId="6" pageNumber="5">Figures 13</emphasis>
, 
<emphasis id="D3A9EA9402078F44FD25F9ACFCEFD306" bold="true" box="[727,787,1629,1650]" pageId="6" pageNumber="5">15–17</emphasis>
. Holotype male, metasoma V and telson lateral (13), metasoma and telson, lateral (15), ventral (16), and dorsal (17) views. 
<emphasis id="D3A9EA9402078F44FD2EF987FCBBD3FF" bold="true" box="[732,839,1654,1675]" pageId="6" pageNumber="5">Figures 14</emphasis>
, 
<emphasis id="D3A9EA9402078F44FCA6F987FC6CD3FF" bold="true" box="[852,912,1654,1675]" pageId="6" pageNumber="5">18–20</emphasis>
. Paratype female, metasoma V and telson lateral (14), metasoma and telson, lateral (18), ventral (19), and dorsal (20) views. Scale bar: 10 mm (15–20).
</paragraph>
</caption>
<paragraph id="E162368602078F44FF32F938FAD9D269" blockId="6.[192,793,1737,1913]" lastBlockId="6.[840,1441,1737,1913]" pageId="6" pageNumber="5">
(
<figureCitation id="79E62A0302078F44FF35F938FEC8D394" box="[199,308,1737,1760]" captionStart="Figures 5–12" captionStartId="5.[192,278,1754,1777]" captionTargetBox="[191,1440,191,1716]" captionTargetId="figure-6@5.[192,1439,192,1715]" captionTargetPageId="5" captionText="Figures 5–12: Chaerilus majkusi sp. n. Figures 5, 7, 9–12. Holotype male, carapace and tergites I–IV (5), sternopectinal region and sternites (7), left legs I–IV, retrolateral aspect (9–12). Figures 6, 8. Paratype female, carapace and tergites I–III (6), sternopectinal region and sternites III–V (8)." pageId="6" pageNumber="5">Figs. 5–6</figureCitation>
). The chelicerae (
<figureCitation id="79E62A0302078F44FDF2F938FD7FD394" box="[512,643,1737,1760]" captionStart="Figures 33–36" captionStartId="8.[192,278,921,944]" captionTargetBox="[199,1426,199,856]" captionTargetId="figure-19@8.[192,1437,192,860]" captionTargetPageId="8" captionText="Figures 33–36: Figures 33–34. Chaerilus majkusi sp. n., male paratype, left chelicera dorsal (33) and ventral (34) views. Figures 35–36. Chaerilus stockmannorum sp. n., male paratype, right chelicera dorsal (35) and ventral (36) views." pageId="6" pageNumber="5">Figs. 33–34</figureCitation>
) are smooth, yellow and strongly reticulate, anteriorly black. The male differs from the female in having pedipalp chela much wider with short fingers. The chela length/width ratio in the male 1.87; in the female 2.28. The male has relatively larger pectines (
<figureCitation id="79E62A0302078F44FE16F893FD82D20D" box="[484,638,1890,1913]" captionStart="Figures 5–12" captionStartId="5.[192,278,1754,1777]" captionTargetBox="[191,1440,191,1716]" captionTargetId="figure-6@5.[192,1439,192,1715]" captionTargetPageId="5" captionText="Figures 5–12: Chaerilus majkusi sp. n. Figures 5, 7, 9–12. Holotype male, carapace and tergites I–IV (5), sternopectinal region and sternites (7), left legs I–IV, retrolateral aspect (9–12). Figures 6, 8. Paratype female, carapace and tergites I–III (6), sternopectinal region and sternites III–V (8)." pageId="6" pageNumber="5">Figs. 7 and 8</figureCitation>
). There is no sexual dimorphism in shape of metasoma and telson. For the position and distribution of trichobothria, see 
<figureCitation id="79E62A0302078F44FCBAF8F7FC35D269" box="[840,969,1798,1821]" captionStart="Figures 21–32" captionStartId="7.[192,278,1343,1366]" captionTargetBox="[215,1420,199,1311]" captionTargetId="figure-6@7.[192,1437,192,1313]" captionTargetPageId="7" captionText="Figures 21–32: Chaerilus majkusi sp. n. Figures 21–24. Paratype female, pedipalp chela, dorsal (21) and externodorsal (22) views, movable (23) and fixed (24) fingers. Figures 25–32. Holotype male, pedipalp chela, dorsal (25), externodorsal (26), and ventrointernal (27) views. Pedipalp patella, dorsal (28), external (29) and ventral (30) views. Pedipalp femur and trochanter, dorsal (31) and ventral (32) views. The trichobothrial pattern is indicated in Figures 25–31." pageId="6" pageNumber="5">Figs. 25–31</figureCitation>
. For measurements, see 
<tableCitation id="AC5F033D02078F44FB3CF8F7FAD9D269" box="[1230,1317,1798,1821]" captionStart="Table 1" captionStartId="17.[192,257,818,841]" captionText="Table 1: Comparative measurements of Chaerilus majkusi sp. n. and Chaerilus neradorum sp. n. types. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="6" pageNumber="5">Table 1.</tableCitation>
</paragraph>
<paragraph id="E162368602078F44FCBAF8D4FC23D20D" blockId="6.[840,1441,1737,1913]" pageId="6" pageNumber="5">
COLORATION (
<figureCitation id="79E62A0302078F44FC16F8D4FBB5D248" box="[996,1097,1829,1852]" captionStart="Figures 1–4" captionStartId="4.[192,278,1612,1635]" captionTargetBox="[206,1423,198,1574]" captionTargetId="figure-19@4.[192,1439,192,1582]" captionTargetPageId="4" captionText="Figures 1–4: Chaerilus majkusi sp. n. Figures 1–2. Holotype male, dorsal (1) and ventral (2) views. Figures 3–4. Paratype female, dorsal (3) and ventral (4) views. Scale bar: 10 mm." pageId="6" pageNumber="5">Figs. 1–4</figureCitation>
). The color is yellowish orange to brown, spotted. Older specimens are darker (see 
<figureCitation id="79E62A0302078F44FA84F8B5FC98D20D" captionStart="Figures 39–40" captionStartId="10.[192,278,1696,1719]" captionTargetBox="[191,1440,167,1666]" captionTargetId="figure-19@10.[192,1439,168,1666]" captionTargetPageId="10" captionText="Figures 39–40: Chaerilus majkusi sp. n., male holotype (39) and female paratype with newborns (40) in vivo habitus." pageId="6" pageNumber="5">Fig. 39</figureCitation>
versus 40).
</paragraph>
<caption id="B5A2660E02068F45FF32FACEFBD0D0D4" pageId="7" pageNumber="6" startId="7.[192,278,1343,1366]" targetBox="[215,1420,199,1311]" targetPageId="7">
<paragraph id="E162368602068F45FF32FACEFBD0D0D4" blockId="7.[192,1440,1343,1440]" pageId="7" pageNumber="6">
<emphasis id="D3A9EA9402068F45FF32FACEFE96D022" bold="true" box="[192,362,1343,1366]" pageId="7" pageNumber="6">Figures 21–32:</emphasis>
<taxonomicName id="26DD4D0502068F45FE86FAB0FDDAD022" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[372,550,1345,1366]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
<emphasis id="D3A9EA9402068F45FE86FAB0FDDAD022" box="[372,550,1345,1366]" italics="true" pageId="7" pageNumber="6">Chaerilus majkusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C89A57EF02068F45FDDEFAB0FD9FD022" box="[556,611,1345,1366]" pageId="7" pageNumber="6" rank="species">
<emphasis id="D3A9EA9402068F45FDDEFAB0FDBFD022" bold="true" box="[556,579,1345,1366]" pageId="7" pageNumber="6">sp</emphasis>
. 
<emphasis id="D3A9EA9402068F45FDA2FAB0FDA1D022" bold="true" box="[592,605,1345,1366]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="D3A9EA9402068F45FD99FAB0FD01D022" bold="true" box="[619,765,1345,1366]" pageId="7" pageNumber="6">Figures 21–24</emphasis>
. Paratype female, pedipalp chela, dorsal (21) and externodorsal (22) views, movable (23) and fixed (24) fingers. 
<emphasis id="D3A9EA9402068F45FD81FAABFCFED01B" bold="true" box="[627,770,1370,1391]" pageId="7" pageNumber="6">Figures 25–32</emphasis>
. Holotype male, pedipalp chela, dorsal (25), externodorsal (26), and ventrointernal (27) views. Pedipalp patella, dorsal (28), external (29) and ventral (30) views. Pedipalp femur and trochanter, dorsal (31) and ventral (32) views. The trichobothrial pattern is indicated in Figures 25–31.
</paragraph>
</caption>
<paragraph id="E162368602068F45FF32FA24FD48D3B7" blockId="7.[192,793,1493,1883]" pageId="7" pageNumber="6">
CARAPACE AND MESOSOMA (
<figureCitation id="79E62A0302068F45FE04FA24FDA2D098" box="[502,606,1493,1516]" captionStart="Figures 5–12" captionStartId="5.[192,278,1754,1777]" captionTargetBox="[191,1440,191,1716]" captionTargetId="figure-6@5.[192,1439,192,1715]" captionTargetPageId="5" captionText="Figures 5–12: Chaerilus majkusi sp. n. Figures 5, 7, 9–12. Holotype male, carapace and tergites I–IV (5), sternopectinal region and sternites (7), left legs I–IV, retrolateral aspect (9–12). Figures 6, 8. Paratype female, carapace and tergites I–III (6), sternopectinal region and sternites III–V (8)." pageId="7" pageNumber="6">Figs. 5–8</figureCitation>
). The carapace is covered by large granules that do not form carinae and are absent from two strips in the median area. The anterior margin of the carapace is almost straight to weakly concave. The mesosomal tergites are irregularly granulated. All sternites are smooth without carinae and granulation. Sternite V with smooth patch indicated. Pectinal teeth number 
<quantity id="26259B6302068F45FE5DF95DFE25D3B7" box="[431,473,1708,1731]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="7" pageNumber="6" unit="in" value="6.0">6 in</quantity>
males, 
<quantity id="26259B6302068F45FDD8F95DFDAFD3B7" box="[554,595,1708,1731]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="6" unit="in" value="4.0">4 in</quantity>
females.
</paragraph>
<paragraph id="E162368602068F45FF32F93BFB14D3F0" blockId="7.[192,793,1493,1883]" lastBlockId="7.[840,1441,1493,1881]" pageId="7" pageNumber="6">
METASOMA AND TELSON (
<figureCitation id="79E62A0302068F45FDF2F93BFD71D395" box="[512,653,1738,1761]" captionStart="Figures 13–20" captionStartId="6.[192,278,1627,1650]" captionTargetBox="[192,1439,204,1588]" captionTargetId="figure-19@6.[192,1439,192,1597]" captionTargetPageId="6" captionText="Figures 13–20: Chaerilus majkusi sp. n. Figures 13, 15–17. Holotype male, metasoma V and telson lateral (13), metasoma and telson, lateral (15), ventral (16), and dorsal (17) views. Figures 14, 18–20. Paratype female, metasoma V and telson lateral (14), metasoma and telson, lateral (18), ventral (19), and dorsal (20) views. Scale bar: 10 mm (15–20)." pageId="7" pageNumber="6">Figs. 13–20</figureCitation>
). The first metasomal segment bears 6 or 8 incomplete carinae, the second to fourth bear six incomplete carinae, and the fifth segment bears four carinae. All carinae are composed of sparse, large granules. The spaces between carinae are very sparsely, irregularly granulated, more so on lateral surfaces of all segments and ventral surfaces of segments IV–V. Metasomal segments I–III are ventrally smooth without carinae and granules. All segments are very sparsely hirsute. The telson is elongate, smooth and very sparsely hirsute.
</paragraph>
<paragraph id="E162368602068F4AFCBAF97DFD8FD136" blockId="7.[840,1441,1493,1881]" lastBlockId="8.[192,793,1006,1305]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
PEDIPALPS (
<figureCitation id="79E62A0302068F45FC3FF97DFBAED3D7" box="[973,1106,1676,1699]" captionStart="Figures 21–32" captionStartId="7.[192,278,1343,1366]" captionTargetBox="[215,1420,199,1311]" captionTargetId="figure-6@7.[192,1437,192,1313]" captionTargetPageId="7" captionText="Figures 21–32: Chaerilus majkusi sp. n. Figures 21–24. Paratype female, pedipalp chela, dorsal (21) and externodorsal (22) views, movable (23) and fixed (24) fingers. Figures 25–32. Holotype male, pedipalp chela, dorsal (25), externodorsal (26), and ventrointernal (27) views. Pedipalp patella, dorsal (28), external (29) and ventral (30) views. Pedipalp femur and trochanter, dorsal (31) and ventral (32) views. The trichobothrial pattern is indicated in Figures 25–31." pageId="7" pageNumber="6">Figs. 21–32</figureCitation>
). The pedipalp chela is stout with swollen manus in the male. The movable finger bears 16 and the fixed finger 15 imbricated rows of granules. The chela has seven or eight mostly granulated carinae. The carina on the externodorsal surface of the manus may be incomplete. The dorsal and internal surfaces of chela with reticulate granulation patterns. The patella is smooth with dorsal carina developed and granulated, other carinae are absent or weakly indicated. The femur has four carinae.
</paragraph>
<caption id="B5A2660E02098F4AFF32FC68FAEDD6BC" pageId="8" pageNumber="7" startId="8.[192,278,921,944]" targetBox="[199,1426,199,856]" targetPageId="8">
<paragraph id="E162368602098F4AFF32FC68FAEDD6BC" blockId="8.[192,1440,921,968]" pageId="8" pageNumber="7">
<emphasis id="D3A9EA9402098F4AFF32FC68FDF5D6DB" bold="true" box="[192,521,921,944]" pageId="8" pageNumber="7">Figures 33–36: Figures 33–34</emphasis>
. 
<taxonomicName id="26DD4D0502098F4AFDEBFC6BFD37D6DB" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[537,715,922,943]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
<emphasis id="D3A9EA9402098F4AFDEBFC6BFD37D6DB" box="[537,715,922,943]" italics="true" pageId="8" pageNumber="7">Chaerilus majkusi</emphasis>
</taxonomicName>
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<emphasis id="D3A9EA9402098F4AFD27FC6BFD10D6DB" bold="true" box="[725,748,922,943]" pageId="8" pageNumber="7">sp</emphasis>
. 
<emphasis id="D3A9EA9402098F4AFD09FC6BFCF4D6DB" bold="true" box="[763,776,922,943]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
, male paratype, left chelicera dorsal (33) and ventral (34) views. 
<emphasis id="D3A9EA9402098F4AFF32FC42FEACD6BC" bold="true" box="[192,336,947,968]" pageId="8" pageNumber="7">Figures 35–36</emphasis>
. 
<taxonomicName id="26DD4D0502098F4AFEA9FC42FDA5D6BC" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[347,601,947,968]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="stockmannorum" status="sp. nov.">
<emphasis id="D3A9EA9402098F4AFEA9FC42FDA5D6BC" box="[347,601,947,968]" italics="true" pageId="8" pageNumber="7">Chaerilus stockmannorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C89A57EF02098F4AFDADFC42FD68D6BC" box="[607,660,947,968]" pageId="8" pageNumber="7" rank="species">
<emphasis id="D3A9EA9402098F4AFDADFC42FD8AD6BC" bold="true" box="[607,630,947,968]" pageId="8" pageNumber="7">sp</emphasis>
. 
<emphasis id="D3A9EA9402098F4AFD70FC42FD73D6BC" bold="true" box="[642,655,947,968]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
, male paratype, right chelicera dorsal (35) and ventral (36) views.
</paragraph>
</caption>
<paragraph id="E162368602098F4AFF32FBBBFE84D06D" blockId="8.[192,793,1006,1305]" pageId="8" pageNumber="7">
LEGS (
<figureCitation id="79E62A0302098F4AFEFAFBBBFE81D115" box="[264,381,1098,1121]" captionStart="Figures 5–12" captionStartId="5.[192,278,1754,1777]" captionTargetBox="[191,1440,191,1716]" captionTargetId="figure-6@5.[192,1439,192,1715]" captionTargetPageId="5" captionText="Figures 5–12: Chaerilus majkusi sp. n. Figures 5, 7, 9–12. Holotype male, carapace and tergites I–IV (5), sternopectinal region and sternites (7), left legs I–IV, retrolateral aspect (9–12). Figures 6, 8. Paratype female, carapace and tergites I–III (6), sternopectinal region and sternites III–V (8)." pageId="8" pageNumber="7">Figs. 9–12</figureCitation>
). The legs are sparsely hirsute, without bristlecombs and carinae. The femora and patellae are granulated dorsally, with other surfaces smooth. The tarsomeres bear four rows of spiniform setae. Spiniform setal formula of inner rows is 7–8/7–8: 10– 12/10–11: 9–12/9–11: 10–11/10–12; of outer rows is 5– 8/5–8 on all legs.
</paragraph>
<paragraph id="E162368602098F4AFF32FAB2FEB5D09B" blockId="8.[192,793,1343,1519]" pageId="8" pageNumber="7">
HEMISPERMATOPHORE (
<figureCitation id="79E62A0302098F4AFE30FACEFDDBD022" box="[450,551,1343,1366]" captionStart="Figures 108–114" captionStartId="23.[192,278,1730,1753]" captionTargetBox="[191,1440,191,1701]" captionTargetId="figure-6@23.[192,1439,192,1700]" captionTargetPageId="23" captionText="Figures 108–114: Hemispermatophores of Chaerilus species. Figure 108. C.cimrmani. Figure 109. C.majkusi sp. n., male holotype. Figure 110. C.tichyi. Figures 111–112. C.stockmannorum sp. n., male holotype. Figures 113–114. C.hofereki, samples from two different individuals, capsule region and distal lamina (113) and whole hemispermatophore (114, reproduced from Kovařík et al., 2015). Convex views shown in all figures except Fig 112 (anterior view). Left hemispermatophore shown in all figures except for Fig. 108, which is right hemispermatophore shown in mirror image for comparison. Scale bars: 500 µm (108–109, 113–114), 1 mm (110–112)." pageId="8" pageNumber="7">Fig. 109</figureCitation>
). Fusiform, with relatively large capsule region (ca. 650 µm), and relatively short distal lamina (ca. 370 µm) angled posteriorly and apically truncated. Sperm hemiduct of capsule weakly sclerotized, with long posterior carina and shorter anterior carina.
</paragraph>
<paragraph id="E162368602098F4AFF32F9E7FC6BD157" blockId="8.[192,793,1558,1918]" lastBlockId="8.[840,1440,1006,1059]" pageId="8" pageNumber="7">
KARYOTYPE (
<figureCitation id="79E62A0302098F4AFEA5F9E7FE09D359" box="[343,501,1558,1581]" captionStart="Figures 115–123" captionStartId="24.[192,278,1474,1497]" captionTargetBox="[191,1437,191,1445]" captionTargetId="figure-383@24.[192,1437,192,1444]" captionTargetPageId="24" captionText="Figures 115–123: The chromosomes of Chaerilus species. Figures 115–116. C. majkusi sp. n., mitotic metaphase (115) and meiotic metaphase I (116). Figures 117–118. C. neradorum sp. n., mitotic metaphase (117) and meiotic metaphase I (118). Figures 119–120. C. stockmannorum sp. n., mitotic metaphase (119) and meiotic metaphase I (120), arrows show chromosomes in pentavalent and arrowheads show chromosomes in trivalent. Figure 121. Mitotic metaphase of C. cimrmani. Figure 122. Meiotic metaphase I of C. sejnai. Figure 123. Mitotic metaphase of C. tichyi. Scale bar: 10 µm." pageId="8" pageNumber="7">Figs. 115–116</figureCitation>
). We analyzed the karyotypes of the male 
<typeStatus id="3E66882402098F4AFE67F9C5FE0FD33F" box="[405,499,1588,1611]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
and the 
<specimenCount id="F7DBFD0F02098F4AFDADF9C5FD29D33F" box="[607,725,1588,1611]" pageId="8" pageNumber="7" type="male">three male</specimenCount>
<typeStatus id="3E66882402098F4AFD13F9C5FF00D31E" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
. The diploid complement of all specimens is 76 chromosomes (
<figureCitation id="79E62A0302098F4AFE9AF983FE39D3FD" box="[360,453,1650,1673]" captionStart="Figures 115–123" captionStartId="24.[192,278,1474,1497]" captionTargetBox="[191,1437,191,1445]" captionTargetId="figure-383@24.[192,1437,192,1444]" captionTargetPageId="24" captionText="Figures 115–123: The chromosomes of Chaerilus species. Figures 115–116. C. majkusi sp. n., mitotic metaphase (115) and meiotic metaphase I (116). Figures 117–118. C. neradorum sp. n., mitotic metaphase (117) and meiotic metaphase I (118). Figures 119–120. C. stockmannorum sp. n., mitotic metaphase (119) and meiotic metaphase I (120), arrows show chromosomes in pentavalent and arrowheads show chromosomes in trivalent. Figure 121. Mitotic metaphase of C. cimrmani. Figure 122. Meiotic metaphase I of C. sejnai. Figure 123. Mitotic metaphase of C. tichyi. Scale bar: 10 µm." pageId="8" pageNumber="7">Fig. 115</figureCitation>
). The chromosomes are small and their morphology is not clearly visible during mitotic metaphase. However, the conspicuous constrictions on chromosomes during postpachytene and metaphase I indicate the existence and predominance of bi-armed chromosomes in this species (
<figureCitation id="79E62A0302098F4AFD85F8FAFD25D256" box="[631,729,1803,1826]" captionStart="Figures 115–123" captionStartId="24.[192,278,1474,1497]" captionTargetBox="[191,1437,191,1445]" captionTargetId="figure-383@24.[192,1437,192,1444]" captionTargetPageId="24" captionText="Figures 115–123: The chromosomes of Chaerilus species. Figures 115–116. C. majkusi sp. n., mitotic metaphase (115) and meiotic metaphase I (116). Figures 117–118. C. neradorum sp. n., mitotic metaphase (117) and meiotic metaphase I (118). Figures 119–120. C. stockmannorum sp. n., mitotic metaphase (119) and meiotic metaphase I (120), arrows show chromosomes in pentavalent and arrowheads show chromosomes in trivalent. Figure 121. Mitotic metaphase of C. cimrmani. Figure 122. Meiotic metaphase I of C. sejnai. Figure 123. Mitotic metaphase of C. tichyi. Scale bar: 10 µm." pageId="8" pageNumber="7">Fig. 116</figureCitation>
). The chromosomes gradually decrease in length from 3.38% to 1.65% of the haploid set. We did not identify any heteromorphic bivalent during postpachytene and metaphase I, and we did not observe chiasmata during these phases.
</paragraph>
<paragraph id="E162368602098F4AFCBAFBB5FB81D024" blockId="8.[840,1441,1092,1666]" pageId="8" pageNumber="7">
A FFINITIES. 
<taxonomicName id="26DD4D0502098F4AFC20FBB5FB60D12F" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[978,1180,1092,1115]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
<emphasis id="D3A9EA9402098F4AFC20FBB5FB60D12F" box="[978,1180,1092,1115]" italics="true" pageId="8" pageNumber="7">Chaerilus majkusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C89A57EF02098F4AFB5BFBB5FB11D12F" box="[1193,1261,1092,1115]" pageId="8" pageNumber="7" rank="species">
<emphasis id="D3A9EA9402098F4AFB5BFBB5FB3ED12F" bold="true" box="[1193,1218,1092,1115]" pageId="8" pageNumber="7">sp</emphasis>
. 
<emphasis id="D3A9EA9402098F4AFB25FBB5FB1AD12F" bold="true" box="[1239,1254,1092,1115]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
is reliably distinguished from all other 
<taxonomicName id="26DD4D0502098F4AFB8CFB93FAB8D10D" box="[1150,1348,1122,1145]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D3A9EA9402098F4AFB8CFB93FB1BD10D" box="[1150,1255,1122,1145]" italics="true" pageId="8" pageNumber="7">Chaerilus</emphasis>
species
</taxonomicName>
by the following unique combination of characters: total length 
<quantity id="26259B6302098F4AFC66FB51FBFAD1C3" box="[916,1030,1184,1207]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="8" pageNumber="7" unit="mm" value="27.0" valueMax="29.0" valueMin="25.0">25–29 mm</quantity>
; movable fingers of pedipalp with 16 rows of granules; pedipalp chela wide, stout with swollen manus in the male; pedipalp fingers straight in both sexes; metasomal segments I–III ventrally smooth without carinae and granules; tarsomeres of legs bear four rows of spiniform setae.
</paragraph>
<paragraph id="E162368602098F4AFC8AFAA9FB36D3F6" blockId="8.[840,1441,1092,1666]" pageId="8" pageNumber="7">
<taxonomicName id="26DD4D0502098F4AFC8AFAA9FBBFD01B" authority="Kovařík &amp; Lowe &amp; Šťáhlavský, 2018" authorityName="Kovařík &amp; Lowe &amp; Šťáhlavský" authorityYear="2018" box="[888,1091,1368,1391]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="majkusi" status="sp. nov.">
<emphasis id="D3A9EA9402098F4AFC8AFAA9FBBFD01B" box="[888,1091,1368,1391]" italics="true" pageId="8" pageNumber="7">Chaerilus majkusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C89A57EF02098F4AFBA0FAA9FB6BD01B" box="[1106,1175,1368,1391]" pageId="8" pageNumber="7" rank="species">
<emphasis id="D3A9EA9402098F4AFBA0FAA9FB97D01B" bold="true" box="[1106,1131,1368,1391]" pageId="8" pageNumber="7">sp</emphasis>
. 
<emphasis id="D3A9EA9402098F4AFB73FAA9FB6CD01B" bold="true" box="[1153,1168,1368,1391]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
is the third 
<taxonomicName id="26DD4D0502098F4AFAC5FAA9FC69D0F9" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D3A9EA9402098F4AFAC5FAA9FA5CD01B" box="[1335,1440,1368,1391]" italics="true" pageId="8" pageNumber="7">Chaerilus</emphasis>
species
</taxonomicName>
recorded from Tioman Island. This island is of special interest because it is inhabited both by 
<taxonomicName id="26DD4D0502098F4AFAADFA64FB8DD0BF" authority="Kovarik, 2000" authorityName="Kovarik" authorityYear="2000" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="tichyi">
<emphasis id="D3A9EA9402098F4AFAADFA64FC3ED0BF" italics="true" pageId="8" pageNumber="7">Chaerilus tichyi</emphasis>
Kovařík, 2000
</taxonomicName>
(
<figureCitation id="79E62A0302098F4AFB7AFA45FAF3D0BF" box="[1160,1295,1460,1483]" captionStart="Figures 41–42" captionStartId="11.[192,278,1764,1787]" captionTargetBox="[191,1440,191,1734]" captionTargetId="figure-125@11.[192,1439,192,1733]" captionTargetPageId="11" captionText="Figures 41–42: Chaerilus tichyi Kovařík, 2000, male (41) and female (42) topotypes in vivo habitus." pageId="8" pageNumber="7">Figs. 41–42</figureCitation>
), the largest known member of the family 
<taxonomicName id="26DD4D0502098F4AFB7DFA23FAF6D09D" authorityName="Pocock" authorityYear="1893" box="[1167,1290,1490,1513]" class="Arachnida" family="Chaerilidae" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="family">Chaerilidae</taxonomicName>
(
<quantity id="26259B6302098F4AFAEEFA23FA6CD09D" box="[1308,1424,1490,1513]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="8" pageNumber="7" unit="mm" value="75.0" valueMax="80.0" valueMin="70.0">70–80 mm</quantity>
), and by 
<taxonomicName id="26DD4D0502098F4AFC6AFA00FB15D37C" authority="Kovarik, 2005" authorityName="Kovarik" authorityYear="2005" box="[920,1257,1521,1544]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="sejnai">
<emphasis id="D3A9EA9402098F4AFC6AFA00FBB4D37C" box="[920,1096,1521,1544]" italics="true" pageId="8" pageNumber="7">Chaerilus sejnai</emphasis>
Kovařík, 2005
</taxonomicName>
(
<figureCitation id="79E62A0302098F4AFB0AFA00FAB3D37C" box="[1272,1359,1521,1544]" captionStart="Figures 37–38" captionStartId="9.[192,278,1830,1853]" captionTargetBox="[191,1440,212,1801]" captionTargetId="figure-42@9.[192,1439,213,1800]" captionTargetPageId="9" captionText="Figures 37–38: Chaerilus majkusi sp. n., female paratype (37) and Chaerilus sejnai Kovařík, 2005, male (38) in vivo habitus." pageId="8" pageNumber="7">Figs. 38</figureCitation>
) which may be smallest known member of the family (
<quantity id="26259B6302098F4AFAAFF9FEFC8FD331" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="8" pageNumber="7" unit="mm" value="19.0" valueMax="22.0" valueMin="16.0">16–22 mm</quantity>
) if we exclude species that were described from juveniles (
<taxonomicName id="26DD4D0502098F4AFC4DF9BCFA97D310" authority="Lourenco, 2012" authorityName="Lourenco" authorityYear="2012" box="[959,1387,1613,1636]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="kampuchea">
<emphasis id="D3A9EA9402098F4AFC4DF9BCFB52D310" box="[959,1198,1613,1636]" italics="true" pageId="8" pageNumber="7">Chaerilus kampuchea</emphasis>
Lourenço, 2012
</taxonomicName>
and 
<taxonomicName id="26DD4D0502098F4AFCBAF99AFB46D3F6" authority="Lourenco, 1995" authorityName="Lourenco" authorityYear="1995" box="[840,1210,1643,1666]" class="Arachnida" family="Chaerilidae" genus="Chaerilus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="sabinae">
<emphasis id="D3A9EA9402098F4AFCBAF99AFBF7D3F6" box="[840,1035,1643,1666]" italics="true" pageId="8" pageNumber="7">Chaerilus sabinae</emphasis>
Lourenço, 1995
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>