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<document id="B4525A9047707B31095CEE4663C3095E" ID-CLB-Dataset="29186" ID-DOI="10.11646/zootaxa.4466.1.6" ID-GBIF-Dataset="202f1fea-aa6f-4fc7-bfad-5a1a10d28703" ID-ISSN="1175-5326" ID-Zenodo-Dep="1454373" ID-ZooBank="39B8D898-27B4-4BCE-A800-E729652C619B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539175046679" checkinUser="plazi" docAuthor="Van Soest, Rob W. M." docDate="2018" docId="04138630FF9CFF884AB7C8DEFD2D0E6F" docLanguage="en" docName="zootaxa.4466.1.6.pdf" docOrigin="Zootaxa 4466 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mycale (Oxymycale) klausjanusorum Van Soest 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="53" masterDocId="F82AFE48FF9DFF8C4A20CE44FF980B3A" masterDocTitle="Mycale (Oxymycale) klausjanusorum sp. nov. (Porifera: Demospongiae: Poecilosclerida) from Flores Sea deep water, Indonesia" masterLastPageNumber="60" masterPageNumber="49" pageNumber="50" updateTime="1698571152293" updateUser="ExternalLinkService">
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<mods:title id="DAADBF675ACFD7750472173F55AD584A">Mycale (Oxymycale) klausjanusorum sp. nov. (Porifera: Demospongiae: Poecilosclerida) from Flores Sea deep water, Indonesia</mods:title>
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<mods:namePart id="5E0AF1C29A339C58E783112B367FEDA2">Van Soest, Rob W. M.</mods:namePart>
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<treatment id="04138630FF9CFF884AB7C8DEFD2D0E6F" ID-DOI="http://doi.org/10.5281/zenodo.5951176" ID-GBIF-Taxon="148673245" ID-Zenodo-Dep="5951176" LSID="urn:lsid:plazi:treatment:04138630FF9CFF884AB7C8DEFD2D0E6F" httpUri="http://treatment.plazi.org/id/04138630FF9CFF884AB7C8DEFD2D0E6F" lastPageId="4" lastPageNumber="53" pageId="1" pageNumber="50">
<subSubSection id="C4A064ADFF9CFF8D4AB7C8DEFE1F0DEC" pageId="1" pageNumber="50" type="nomenclature">
<paragraph id="8C053726FF9CFF8D4AB7C8DEFD580D8F" blockId="1.[151,704,1690,1750]" box="[151,704,1690,1717]" pageId="1" pageNumber="50">
<heading id="D74D804AFF9CFF8D4AB7C8DEFD580D8F" bold="true" box="[151,704,1690,1717]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
<emphasis id="BECEEB34FF9CFF8D4AB7C8DEFD580D8F" bold="true" box="[151,704,1690,1717]" pageId="1" pageNumber="50">
<taxonomicName id="4BBA4CA5FF9CFF8D4AB7C8DEFDC40D8E" ID-CoL="44RZ8" authority="Van Soest, 2018" authorityName="Van Soest" authorityYear="2018" box="[151,604,1690,1716]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="1" pageNumber="50" phylum="Porifera" rank="species" species="klausjanusorum" status="sp. nov." subGenus="Oxymycale">
<emphasis id="BECEEB34FF9CFF8D4AB7C8DEFDC40D8E" bold="true" box="[151,604,1690,1716]" italics="true" pageId="1" pageNumber="50">Mycale (Oxymycale) klausjanusorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5FD564FFF9CFF8D4843C8DFFD580D8F" box="[611,704,1691,1717]" pageId="1" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8C053726FF9CFF8D4AB7C8F9FE1F0DEC" blockId="1.[151,704,1690,1750]" box="[151,391,1725,1750]" pageId="1" pageNumber="50">
<figureCitation id="14812BA3FF9CFF8D4AB7C8F9FE9F0DEC" box="[151,263,1725,1750]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="1" pageNumber="50">Figs 1a–d</figureCitation>
,
<figureCitation id="14812BA3FF9CFF8D4B33C8FAFED20DEC" box="[275,330,1726,1750]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="1" pageNumber="50">2a–c</figureCitation>
,
<figureCitation id="14812BA3FF9CFF8D4B76C8F9FE1F0DEC" box="[342,391,1725,1750]" captionStart="FIGURE 3" captionStartId="6.[151,250,1894,1916]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@6.[151,1436,210,1873]" captionTargetPageId="6" captionText="FIGURE 3. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of the microsclere complement, a, anisochelae I shown at three different angles, b, anisochelae II shown at two different angles, c, anisochelae III, shown at three different angles, d-e, sigmas I and II, f, larger partial trichodragma (incomplete as most trichodragmas fell apart in the process of spicule dissociation)." httpUri="https://zenodo.org/record/1454379/files/figure.png" pageId="1" pageNumber="50">3a–f</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C4A064ADFF9CFF8D4AB7C941FC610CB0" pageId="1" pageNumber="50" type="materials_examined">
<paragraph id="8C053726FF9CFF8D4AB7C941FEA00C5C" blockId="1.[151,1436,1797,2038]" pageId="1" pageNumber="50">
<emphasis id="BECEEB34FF9CFF8D4AB7C941FE1C0C24" bold="true" box="[151,388,1797,1822]" pageId="1" pageNumber="50">Material examined.</emphasis>
<typeStatus id="53018984FF9CFF8D4BAEC941FE6E0C24" box="[398,502,1797,1822]" pageId="1" pageNumber="50" type="holotype">Holotype</typeStatus>
<collectionCode id="EAABAFE3FF9CFF8D4821C942FDA70C24" box="[513,575,1798,1822]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="1" pageNumber="50">ZMA</collectionCode>
Por. 0 9452,
<collectingCountry id="F4AD77B6FF9CFF8D48F8C941FCD00C24" box="[728,840,1797,1822]" name="Indonesia" pageId="1" pageNumber="50">Indonesia</collectingCountry>
,
<collectingCountry id="F4AD77B6FF9CFF8D4977C941FC050C24" box="[855,925,1797,1822]" name="Indonesia" pageId="1" pageNumber="50">Flores</collectingCountry>
Sea, SW Salayar, E of Bahuluang,
<geoCoordinate id="E98E51E1FF9CFF8D4F62C941FA040C24" box="[1346,1436,1797,1822]" direction="south" orientation="latitude" pageId="1" pageNumber="50" precision="55" value="-6.485">6.485°S</geoCoordinate>
<geoCoordinate id="E98E51E1FF9CFF8D4AB7C96DFE9A0C78" box="[151,258,1833,1858]" direction="east" orientation="longitude" pageId="1" pageNumber="50" precision="555" value="120.45">120.45°E</geoCoordinate>
, depth
<quantity id="4B429AC3FF9CFF8D4B73C96DFE4B0C7B" box="[339,467,1833,1858]" metricMagnitude="2" metricUnit="m" metricValue="2.975" metricValueMax="3.0" metricValueMin="2.95" pageId="1" pageNumber="50" unit="m" value="297.5" valueMax="300.0" valueMin="295.0">295–300 m</quantity>
, rectangular dredge, coll. J. van der Land, RV
<taxonomicName id="4BBA4CA5FF9CFF8D49F9C96EFBF30C78" authority="Snellius" authorityName="Snellius" box="[985,1131,1833,1858]" class="Malacostraca" family="Scinidae" genus="Tyro" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="BECEEB34FF9CFF8D49F9C96EFB910C7B" box="[985,1033,1834,1857]" italics="true" pageId="1" pageNumber="50">Tyro</emphasis>
Snellius
</taxonomicName>
II Expedition, stat. 217,
<date id="F80411E6FF9CFF8D4FA0C96EFEAC0C5C" pageId="1" pageNumber="50" value="1984-10-12">12 October 1984</date>
.
</paragraph>
<paragraph id="8C053726FF9CFF8D4AE7C936FC610CB0" blockId="1.[151,1436,1797,2038]" box="[199,1017,1905,1930]" pageId="1" pageNumber="50">
<materialsCitation id="3CD23D7BFF9CFF8D4AE7C936FC610CB0" ID-GBIF-Occurrence="1915499073" box="[199,1017,1905,1930]" collectionCode="ZMA" pageId="1" pageNumber="50" specimenCount="1" typeStatus="paratype">
<typeStatus id="53018984FF9CFF8D4AE7C936FEAC0CB0" box="[199,308,1906,1930]" pageId="1" pageNumber="50" type="paratype">Paratypes</typeStatus>
(detached fragments)
<collectionCode id="EAABAFE3FF9CFF8D480FC936FDF50CB0" box="[559,621,1906,1930]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="1" pageNumber="50">ZMA</collectionCode>
Por. 0 9459, same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C4A064ADFF9CFF8E4AE7C9D1FE4C0E25" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50" type="description">
<paragraph id="8C053726FF9CFF8D4AE7C9D1FCE20CCC" blockId="1.[151,1436,1797,2038]" pageId="1" pageNumber="50">
<emphasis id="BECEEB34FF9CFF8D4AE7C9D1FEC90C94" bold="true" box="[199,337,1941,1966]" pageId="1" pageNumber="50">Description</emphasis>
. (
<figureCitation id="14812BA3FF9CFF8D4B4CC9D1FE260C94" box="[364,446,1941,1966]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="1" pageNumber="50">Fig. 1a</figureCitation>
) Semiglobular massive encrustations on calcareous rubble, whitish with pale orange patches (in alcohol). Size of
<typeStatus id="53018984FF9CFF8D4BF6C9FDFDA00CE8" box="[470,568,1977,2002]" pageId="1" pageNumber="50" type="holotype">holotype</typeStatus>
2
<date id="F80411E6FF9CFF8D4874C9FEFD380CE8" box="[596,672,1978,2002]" pageId="1" pageNumber="50">
x 1 x
<quantity id="4B429AC3FF9CFF8D48B2C9FEFD380CE8" box="[658,672,1978,2002]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="50" unit="cm" value="1.0">1</quantity>
</date>
cm, of
<typeStatus id="53018984FF9CFF8D48D5C9FEFCCE0CE8" box="[757,854,1978,2002]" pageId="1" pageNumber="50" type="paratype">paratype</typeStatus>
detached fragments the largest of which is 3.5
<date id="F80411E6FF9CFF8D4F44C9FEFA1F0CE8" box="[1380,1415,1978,2002]" pageId="1" pageNumber="50">x 1</date>
x
<quantity id="4B429AC3FF9CFF8D4AB7C999FF7F0CCF" box="[151,231,2013,2038]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="50" unit="cm" value="0.5">0.5 cm</quantity>
. Surface markedly hispid. Consistency compressible, firm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CED3FB8B0963" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
<emphasis id="BECEEB34FF9FFF8E4AE7CED3FEAD0B8A" bold="true" box="[199,309,151,176]" pageId="2" pageNumber="51">Skeleton.</emphasis>
(
<figureCitation id="14812BA3FF9FFF8E4B65CED3FE210B8A" box="[325,441,151,176]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="2" pageNumber="51">Figs 1b–d</figureCitation>
,
<figureCitation id="14812BA3FF9FFF8E4BE6CEDCFE7A0B8A" box="[454,482,152,176]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">2a</figureCitation>
) The choanosomal skeleton is strongly plumose, with thick spicule tracts present in the interior region (visible in
<figureCitation id="14812BA3FF9FFF8E4BFECEFFFDB50BEE" box="[478,557,187,212]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="2" pageNumber="51">Fig. 1d</figureCitation>
) about 500–600 µm in diameter, which towards the surface divide into thinner tracts of about 200 µm (
<figureCitation id="14812BA3FF9FFF8E4B86CE9AFE650BCD" box="[422,509,222,247]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="2" pageNumber="51">Figs 1b</figureCitation>
,
<figureCitation id="14812BA3FF9FFF8E482ACE9BFDB00BCD" box="[522,552,223,247]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">2a</figureCitation>
). The subectosomal tracts partially carry a tangential ectosomal skeleton of 80 µm in thickness (visible in
<figureCitation id="14812BA3FF9FFF8E4BE5CF46FDAC0A21" box="[453,564,258,283]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="2" pageNumber="51">Figs 1b–c</figureCitation>
,
<figureCitation id="14812BA3FF9FFF8E4860CF47FDC30A21" box="[576,603,259,283]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">2a</figureCitation>
) consisting predominantly of oxeas, but the tracts are also fanning out into bouquets of individual megascleres (
<figureCitation id="14812BA3FF9FFF8E481ACF61FD350A04" box="[570,685,293,318]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="2" pageNumber="51">Figs 1b–c</figureCitation>
,
<figureCitation id="14812BA3FF9FFF8E489BCF62FD400A04" box="[699,728,294,318]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">2a</figureCitation>
), that protrude beyond the tangential ectosomal skeleton. The bouquets consist of longer style-like megascleres, differing in length and shape to the oxea-like megascleres from the tangential ectosomal skeleton. The choanosomal tracts appear to consist of a mixture of the two megasclere
<typeStatus id="53018984FF9FFF8E4AB7CFD5FF4E0A93" box="[151,214,401,425]" pageId="2" pageNumber="51">types</typeStatus>
. This bouquet-type of surface skeleton is unusual for
<taxonomicName id="4BBA4CA5FF9FFF8E497CCFD4FC370A93" authorityName="Gray" authorityYear="1867" box="[860,943,400,425]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF9FFF8E497CCFD4FC370A93" box="[860,943,400,425]" italics="true" pageId="2" pageNumber="51">Mycale</emphasis>
</taxonomicName>
species. Microscleres are confined to the choanosomal and tangential ectosomal skeleton, and are absent in the surface bouquets. Anisochelae I are predominantly arranged in rosettes (
<figureCitation id="14812BA3FF9FFF8E4864CF93FD030AD5" box="[580,667,471,496]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">Fig. 2a</figureCitation>
), with diameters of 300-400 µm, anisochelae II are individually distributed, relatively scarce, anisochelae III are crowded in irregular groups among the spicule tracts. Sigmas I are evenly and densely distributed, sigmas II are rather rare, and are individually distributed. Trichodragmas are densely distributed, the larger category most abundant, the smaller quite scarce.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CC21FE03099B" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
<emphasis id="BECEEB34FF9FFF8E4AE7CC21FEA80944" bold="true" box="[199,304,613,638]" pageId="2" pageNumber="51">Spicules.</emphasis>
(
<figureCitation id="14812BA3FF9FFF8E4B61CC21FE2D0944" box="[321,437,613,638]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">Figs 2b–c</figureCitation>
,
<figureCitation id="14812BA3FF9FFF8E4BE3CC21FE6E0944" box="[451,502,613,638]" captionStart="FIGURE 3" captionStartId="6.[151,250,1894,1916]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@6.[151,1436,210,1873]" captionTargetPageId="6" captionText="FIGURE 3. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of the microsclere complement, a, anisochelae I shown at three different angles, b, anisochelae II shown at two different angles, c, anisochelae III, shown at three different angles, d-e, sigmas I and II, f, larger partial trichodragma (incomplete as most trichodragmas fell apart in the process of spicule dissociation)." httpUri="https://zenodo.org/record/1454379/files/figure.png" pageId="2" pageNumber="51">3a–f</figureCitation>
) Oxeote and stylote anisoxeas, three categories of anisochelae, two categories of sigmas, trichodragmas.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CCE8FE560836" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
Oxeote megascleres (
<figureCitation id="14812BA3FF9FFF8E4B98CCE8FDB009FF" box="[440,552,684,709]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">Figs 2b,b</figureCitation>
<figureCitation id="14812BA3FF9FFF8E4808CCE8FDAA09FF" box="[552,562,684,709]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="2" pageNumber="51">1</figureCitation>
), at lower magnification optically symmetrical (
<figureCitation id="14812BA3FF9FFF8E4E71CCE8FB3209FF" box="[1105,1194,684,709]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">Figs 2b</figureCitation>
), but magnified (
<figureCitation id="14812BA3FF9FFF8E4F4ACCE8FF2009D2" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">Figs 2b</figureCitation>
<figureCitation id="14812BA3FF9FFF8E4A97CC8BFF5909D2" box="[183,193,719,744]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="2" pageNumber="51">1</figureCitation>
) they are slightly asymmetric with opposite more gradually pointed and more mucronated blunt endings, 522–
<emphasis id="BECEEB34FF9FFF8E4AB7CCB7FF590836" box="[151,193,755,780]" italics="true" pageId="2" pageNumber="51">656</emphasis>
–732
<date id="F80411E6FF9FFF8E4B20CCB0FEB70831" box="[256,303,755,779]" pageId="2" pageNumber="51" value="2014-10">x 14</date>
–
<emphasis id="BECEEB34FF9FFF8E4B1FCCB7FEE80836" box="[319,368,755,780]" italics="true" pageId="2" pageNumber="51">16.8</emphasis>
–20 µm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CD52FC8D0868" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
Stylote megascleres (
<figureCitation id="14812BA3FF9FFF8E4B95CD52FD930815" box="[437,523,790,815]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">Figs 2c</figureCitation>
), with opposite gradually pointed and bluntly rounded end (
<figureCitation id="14812BA3FF9FFF8E4E90CD52FA910815" box="[1200,1289,790,815]" captionStart="FIGURE 2" captionStartId="5.[151,250,1878,1900]" captionTargetBox="[151,1436,195,1856]" captionTargetId="figure@5.[151,1436,195,1856]" captionTargetPageId="5" captionText="FIGURE 2. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of skeleton and megascleres, a, cross section of peripheral skeleton showing bouquets of larger styles and tangential ectosomal skeleton of oxeas, carried by subectosomal spicule tracts. In the subectosomal spaces several anisochelae I rosettes are visible. b, oxeote megascleres, b1, magnified details of apices and shaft of the oxeote megascleres, c, stylote megascleres, c1, details of apices and shaft of stylote megascleres." httpUri="https://zenodo.org/record/1454377/files/figure.png" pageId="2" pageNumber="51">Figs 2c</figureCitation>
<figureCitation id="14812BA3FF9FFF8E4F28CD53FA8B0815" box="[1288,1299,791,815]" captionStart="FIGURE 1" captionStartId="3.[151,247,1618,1640]" captionTargetBox="[151,1435,193,1597]" captionTargetId="figure@3.[151,1436,193,1597]" captionTargetPageId="3" captionText="FIGURE 1. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts." httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="2" pageNumber="51">1</figureCitation>
), often with slightly narrow neck, 992–
<emphasis id="BECEEB34FF9FFF8E4BE3CD7EFE630868" box="[451,507,826,850]" italics="true" pageId="2" pageNumber="51">1082</emphasis>
–
<date id="F80411E6FF9FFF8E482ACD7DFDEE0868" box="[522,630,825,850]" pageId="2" pageNumber="51" value="1175-10-19">1175 x 19</date>
–
<emphasis id="BECEEB34FF9FFF8E48A6CD7EFD2F0868" box="[646,695,826,850]" italics="true" pageId="2" pageNumber="51">20.7</emphasis>
–22 µm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CD19FEE208A3" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
Anisochela I (
<figureCitation id="14812BA3FF9FFF8E4B46CD19FE25084C" box="[358,445,861,886]" captionStart="FIGURE 3" captionStartId="6.[151,250,1894,1916]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@6.[151,1436,210,1873]" captionTargetPageId="6" captionText="FIGURE 3. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of the microsclere complement, a, anisochelae I shown at three different angles, b, anisochelae II shown at two different angles, c, anisochelae III, shown at three different angles, d-e, sigmas I and II, f, larger partial trichodragma (incomplete as most trichodragmas fell apart in the process of spicule dissociation)." httpUri="https://zenodo.org/record/1454379/files/figure.png" pageId="2" pageNumber="51">Figs 3a</figureCitation>
), robust, with flaring upper alae, naked shaft about one third of the total spicule length, 157–
<emphasis id="BECEEB34FF9FFF8E4AEFCDC4FE9608A3" box="[207,270,896,921]" italics="true" pageId="2" pageNumber="51">172.6</emphasis>
–189 µm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CDE0FC0108DB" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
Anisochela II (
<figureCitation id="14812BA3FF9FFF8E4B56CDE0FE490887" box="[374,465,932,957]" captionStart="FIGURE 3" captionStartId="6.[151,250,1894,1916]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@6.[151,1436,210,1873]" captionTargetPageId="6" captionText="FIGURE 3. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of the microsclere complement, a, anisochelae I shown at three different angles, b, anisochelae II shown at two different angles, c, anisochelae III, shown at three different angles, d-e, sigmas I and II, f, larger partial trichodragma (incomplete as most trichodragmas fell apart in the process of spicule dissociation)." httpUri="https://zenodo.org/record/1454379/files/figure.png" pageId="2" pageNumber="51">Figs 3b</figureCitation>
), rather similar in shape to
<taxonomicName id="4BBA4CA5FF9FFF8E493FCDE0FC0D0887" authorityName="Levi" authorityYear="1963" box="[799,917,932,957]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="species" species="anisochela">anisochela</taxonomicName>
I but with upper alae less flaring and shaft occupying only a quarter of the total spicule length, 33–
<emphasis id="BECEEB34FF9FFF8E492ACD8CFCA308DA" box="[778,827,968,992]" italics="true" pageId="2" pageNumber="51">44.9</emphasis>
–57 µm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CDAFFC120F1D" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
Anisochela III (
<figureCitation id="14812BA3FF9FFF8E4B5CCDAFFE4C0F39" box="[380,468,1003,1028]" captionStart="FIGURE 3" captionStartId="6.[151,250,1894,1916]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@6.[151,1436,210,1873]" captionTargetPageId="6" captionText="FIGURE 3. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of the microsclere complement, a, anisochelae I shown at three different angles, b, anisochelae II shown at two different angles, c, anisochelae III, shown at three different angles, d-e, sigmas I and II, f, larger partial trichodragma (incomplete as most trichodragmas fell apart in the process of spicule dissociation)." httpUri="https://zenodo.org/record/1454379/files/figure.png" pageId="2" pageNumber="51">Figs 3c</figureCitation>
), with upper alae close to the shaft and occupying more than half the length of the spicule, lower part reduced to a single spurred ala, 21–
<emphasis id="BECEEB34FF9FFF8E48DBCA4BFCB40F1D" box="[763,812,1039,1063]" italics="true" pageId="2" pageNumber="51">24.0</emphasis>
–29 µm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CA76FB2A0F71" blockId="2.[151,1438,151,2021]" box="[199,1202,1074,1099]" pageId="2" pageNumber="51">
Sigma I (
<figureCitation id="14812BA3FF9FFF8E4B0ECA76FE190F71" box="[302,385,1074,1099]" captionStart="FIGURE 3" captionStartId="6.[151,250,1894,1916]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@6.[151,1436,210,1873]" captionTargetPageId="6" captionText="FIGURE 3. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of the microsclere complement, a, anisochelae I shown at three different angles, b, anisochelae II shown at two different angles, c, anisochelae III, shown at three different angles, d-e, sigmas I and II, f, larger partial trichodragma (incomplete as most trichodragmas fell apart in the process of spicule dissociation)." httpUri="https://zenodo.org/record/1454379/files/figure.png" pageId="2" pageNumber="51">Fig. 3d</figureCitation>
), predominantly strongly curved with thin sharp apices, 37–
<emphasis id="BECEEB34FF9FFF8E4E03CA76FBCC0F70" box="[1059,1108,1074,1098]" italics="true" pageId="2" pageNumber="51">44.7</emphasis>
–54 µm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CA11FAF90F54" blockId="2.[151,1438,151,2021]" box="[199,1377,1109,1134]" pageId="2" pageNumber="51">
Sigma II (
<figureCitation id="14812BA3FF9FFF8E4B17CA11FE100F54" box="[311,392,1109,1134]" captionStart="FIGURE 3" captionStartId="6.[151,250,1894,1916]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@6.[151,1436,210,1873]" captionTargetPageId="6" captionText="FIGURE 3. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of the microsclere complement, a, anisochelae I shown at three different angles, b, anisochelae II shown at two different angles, c, anisochelae III, shown at three different angles, d-e, sigmas I and II, f, larger partial trichodragma (incomplete as most trichodragmas fell apart in the process of spicule dissociation)." httpUri="https://zenodo.org/record/1454379/files/figure.png" pageId="2" pageNumber="51">Fig. 3e</figureCitation>
), similar in shape to the larger sigmas and close to overlapping in size, 16–
<emphasis id="BECEEB34FF9FFF8E4EF2CA12FA9B0F54" box="[1234,1283,1110,1134]" italics="true" pageId="2" pageNumber="51">20.3</emphasis>
–27 µm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CA3DFCD40FE3" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
Trichodragmas (
<figureCitation id="14812BA3FF9FFF8E4BA1CA3DFE4B0FA8" box="[385,467,1145,1170]" captionStart="FIGURE 3" captionStartId="6.[151,250,1894,1916]" captionTargetBox="[151,1436,210,1873]" captionTargetId="figure@6.[151,1436,210,1873]" captionTargetPageId="6" captionText="FIGURE 3. Mycale (Oxymycale) klausjanusorum sp. nov., holotype ZMA Por. 09452, SEM images of the microsclere complement, a, anisochelae I shown at three different angles, b, anisochelae II shown at two different angles, c, anisochelae III, shown at three different angles, d-e, sigmas I and II, f, larger partial trichodragma (incomplete as most trichodragmas fell apart in the process of spicule dissociation)." httpUri="https://zenodo.org/record/1454379/files/figure.png" pageId="2" pageNumber="51">Fig. 3f</figureCitation>
), straight, possibly divisible in two size categories, but overlapping, the larger, very common, 78–
<emphasis id="BECEEB34FF9FFF8E4B19CAD9FEF20F8F" box="[313,362,1181,1205]" italics="true" pageId="2" pageNumber="51">85.8</emphasis>
–107
<date id="F80411E6FF9FFF8E4B90CAD9FE7F0F8F" box="[432,487,1181,1205]" pageId="2" pageNumber="51" value="2014-10">x 14</date>
–
<emphasis id="BECEEB34FF9FFF8E4BD7CAD8FDB00F8F" box="[503,552,1180,1205]" italics="true" pageId="2" pageNumber="51">20.6</emphasis>
–28 µm, the smaller, rare (only a few (n=4) were observed in the light microscopic slides, not shown), 24–
<emphasis id="BECEEB34FF9FFF8E480ACA84FDC30FE2" box="[554,603,1216,1240]" italics="true" pageId="2" pageNumber="51">35.5</emphasis>
–
<date id="F80411E6FF9FFF8E4849CAFBFD360FE2" box="[617,686,1215,1240]" pageId="2" pageNumber="51" value="1954-10-09">54 x 9</date>
–
<emphasis id="BECEEB34FF9FFF8E489CCAFBFD750FE2" box="[700,749,1215,1240]" italics="true" pageId="2" pageNumber="51">10.6</emphasis>
–12 µm.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7CAA7FE4C0E25" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
<emphasis id="BECEEB34FF9FFF8E4AE7CAA7FE680FC6" bold="true" box="[199,496,1251,1276]" pageId="2" pageNumber="51">Occurrence and ecology.</emphasis>
<collectingCountry id="F4AD77B6FF9FFF8E4BD8CAA7FDF00FC6" box="[504,616,1251,1276]" name="Indonesia" pageId="2" pageNumber="51">Indonesia</collectingCountry>
, between the islands of Salayar and Bahuluang,
<collectingCountry id="F4AD77B6FF9FFF8E4EB4CAA7FB420FC6" box="[1172,1242,1251,1276]" name="Indonesia" pageId="2" pageNumber="51">Flores</collectingCountry>
Sea, on gravelly bottom at
<quantity id="4B429AC3FF9FFF8E4B28CB42FE1F0E25" box="[264,391,1286,1311]" metricMagnitude="2" metricUnit="m" metricValue="2.975" metricValueMax="3.0" metricValueMin="2.95" pageId="2" pageNumber="51" unit="m" value="297.5" valueMax="300.0" valueMin="295.0">295–300 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="C4A064ADFF9FFF8E4AE7CB6EFB9B0EB0" pageId="2" pageNumber="51" type="etymology">
<paragraph id="8C053726FF9FFF8E4AE7CB6EFB9B0EB0" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
<emphasis id="BECEEB34FF9FFF8E4AE7CB6EFED60E79" bold="true" box="[199,334,1322,1347]" pageId="2" pageNumber="51">Etymology.</emphasis>
It is my pleasure to name the new species combinedly to Klaus Rützler and Jean (‘Janus’) Vacelet in recognition of their life-long dedication to the science of sponges. Both were supportive and formative during my early sponge activities and over the years showed much valued friendship.
</paragraph>
</subSubSection>
<subSubSection id="C4A064ADFF9FFF884AE7CBD0FD5508D7" lastPageId="4" lastPageNumber="53" pageId="2" pageNumber="51" type="discussion">
<paragraph id="8C053726FF9FFF8E4AE7CBD0FD730ECE" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
<emphasis id="BECEEB34FF9FFF8E4AE7CBD0FEA30E97" bold="true" box="[199,315,1428,1453]" pageId="2" pageNumber="51">Remarks.</emphasis>
The
<typeStatus id="53018984FF9FFF8E4B5DCBD0FE470E97" box="[381,479,1428,1453]" pageId="2" pageNumber="51" type="holotype">holotype</typeStatus>
and the
<typeStatus id="53018984FF9FFF8E486BCBD1FD330E97" box="[587,683,1429,1453]" pageId="2" pageNumber="51" type="paratype">paratype</typeStatus>
fragments are possibly part of a compound specimen, of which fragments became detached during the dredging operation. The uncertainty is the reason for separately registering two samples, distinguished as
<typeStatus id="53018984FF9FFF8E4BCBCB9FFDD50ECE" box="[491,589,1499,1524]" pageId="2" pageNumber="51" type="holotype">holotype</typeStatus>
and
<typeStatus id="53018984FF9FFF8E48A3CB98FD7F0ECE" box="[643,743,1500,1524]" pageId="2" pageNumber="51" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7C844FAF90DD6" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
I assembled the main features of all
<taxonomicName id="4BBA4CA5FF9FFF8E4840CBBBFCDD0D22" authority="(Oxymycale)" authorityName="Gray" authorityYear="1867" baseAuthorityName="Oxymycale" box="[608,837,1535,1560]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF9FFF8E4840CBBBFCDD0D22" box="[608,837,1535,1560]" italics="true" pageId="2" pageNumber="51">Mycale (Oxymycale)</emphasis>
</taxonomicName>
species in
<tableCitation id="C138029DFF9FFF8E49E4CBBBFB1A0D22" box="[964,1154,1535,1560]" captionStart="TABLE 1" captionStartId="7.[147,223,586,607]" captionText="TABLE 1a. Macroscopical characters, skeletal features and ecogeographical data of species assigned to Mycale (Oxymycale), including source references. Species are listed alphabetically. For spicule categories and sizes cf. Table 1b." pageId="2" pageNumber="51">Tables 1a and 1b</tableCitation>
[one species,
<taxonomicName id="4BBA4CA5FF9FFF8E4F02CBBBFE660D01" authority="Pulitzer-Finali, 1986" authorityName="Pulitzer-Finali" authorityYear="1986" class="Demospongiae" family="Mycalidae" genus="Oxymycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="species" species="strongylata">
<emphasis id="BECEEB34FF9FFF8E4F02CBBBFE8B0D01" italics="true" pageId="2" pageNumber="51">Oxymycale strongylata</emphasis>
<bibRefCitation id="E82B4AD7FF9FFF8E4B3BC866FE660D01" box="[283,510,1570,1595]" pageId="2" pageNumber="51" refId="ref6602" refString="Pulitzer-Finali, G. (1986) A collection of West Indian Demospongiae (Porifera). In: appendix, a list of the Demospongiae hitherto recorded from the West Indies. Annali del Museo civico di storia naturale Giacomo Doria, 86, 65 - 216. Available from: https: // www. biodiversitylibrary. org / item / 106570 # page / 75 / mode / 1 up (accessed 30 August 2018)" type="journal article">Pulitzer-Finali, 1986</bibRefCitation>
</taxonomicName>
is excluded because Hajdu &amp; Desqeyroux-Faúndez (1994, p. 578) found it to be a junior synonym of
<taxonomicName id="4BBA4CA5FF9FFF8E4B4CC802FC800D65" authority="(Carter, 1882)" authorityName="Carter" authorityYear="1882" baseAuthorityName="Carter" baseAuthorityYear="1882" box="[364,792,1606,1631]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="species" species="laevis" subGenus="Mycale">
<emphasis id="BECEEB34FF9FFF8E4B4CC802FDF70D65" box="[364,623,1606,1631]" italics="true" pageId="2" pageNumber="51">Mycale (Mycale) laevis</emphasis>
(
<bibRefCitation id="E82B4AD7FF9FFF8E485FC803FC880D65" box="[639,784,1607,1631]" pageId="2" pageNumber="51" refId="ref5150" refString="Carter, H. J. (1882) Some Sponges from the West Indies and Acapulco in the Liverpool Free Museum described, with general and classificatory Remarks. Annals and Magazine of Natural History, Series 5, 9 (52), 266 - 301, 346 - 368. Available from: https: // www. biodiversitylibrary. org / item / 84520 # page / 280 / mode / 1 up (accessed 30 August 2018)" type="journal article">Carter, 1882</bibRefCitation>
)
</taxonomicName>
]. From this table it can be concluded that the new species differs from all other
<taxonomicName id="4BBA4CA5FF9FFF8E4BA2C82DFDFE0DB8" authority="(Oxymycale)" authorityName="Gray" authorityYear="1867" baseAuthorityName="Oxymycale" box="[386,614,1641,1666]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF9FFF8E4BA2C82DFDFE0DB8" box="[386,614,1641,1666]" italics="true" pageId="2" pageNumber="51">Mycale (Oxymycale)</emphasis>
</taxonomicName>
in (1) having two distinct
<typeStatus id="53018984FF9FFF8E49A9C82EFC5B0DB8" box="[905,963,1642,1666]" pageId="2" pageNumber="51">types</typeStatus>
of megascleres differing also in length, (2) in the greater length of the megascleres (up to 1175 µm, whereas the greatest length of the other species is 850 µm), (3) the ectosomal bouquets of single megascleres protruding beyond the tangential ectosomal skeleton, (4) the larger anisochelae I (up to 189 µm, whereas the largest anisochelae of the other species reach only up to 105–110 µm).
</paragraph>
<paragraph id="8C053726FF9FFF8E4AE7C8B3FA060CDF" blockId="2.[151,1438,151,2021]" pageId="2" pageNumber="51">
The closest species to the new species appears to be the one
<bibRefCitation id="E82B4AD7FF9FFF8E4957C8B3FBD60C2A" box="[887,1102,1783,1808]" pageId="2" pageNumber="51" refId="ref6984" refString="Sim, C. J. &amp; Kim, M. H. (1988) A systematic study on the marine sponges in Korea. 7. Demospongiae and Hexactinellida. Korean Journal of Systematic Zoology, 4 (1), 21 - 42. Available from: http: // www. dbpia. co. kr / Journal / ArticleDetail / NODE 01801424 (accessed 30 August 2018)" type="journal article">Sim &amp; Kim (1988)</bibRefCitation>
(p. 25) described from
<collectingCountry id="F4AD77B6FF9FFF8E4F7BC8B3FF430C09" name="South Korea" pageId="2" pageNumber="51">South Korea</collectingCountry>
as
<emphasis id="BECEEB34FF9FFF8E4B22C95FFEEA0C0E" box="[258,370,1819,1844]" italics="true" pageId="2" pageNumber="51">Oxymcale</emphasis>
<taxonomicName id="4BBA4CA5FF9FFF8E4B5AC95FFD440C0E" authority="De Laubenfels, 1935" authorityName="De Laubenfels" authorityYear="1935" box="[378,732,1819,1844]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="species" species="paradoxa">
<emphasis id="BECEEB34FF9FFF8E4B5AC95FFE7E0C0E" box="[378,486,1819,1844]" italics="true" pageId="2" pageNumber="51">paradoxa</emphasis>
<bibRefCitation id="E82B4AD7FF9FFF8E4BCDC958FD440C0E" box="[493,732,1819,1844]" pageId="2" pageNumber="51" refId="ref5233" refString="De Laubenfels, M. W. (1935). Some Sponges of Lower California (Mexico). American Museum Novitates, 779, 1 - 14. Available from: https: // digitallibrary. amnh. org / bitstream / handle / 2246 / 4560 / v 2 / dspace / ingest / pdfSource / nov / N 0779. pdf? sequence = 1 &amp; isAllowed = y (accessed 30 August 2018)" type="journal article">De Laubenfels, 1935</bibRefCitation>
</taxonomicName>
. It shares the presence of oxea-like and style-like megascleres and overall habitus, but differs in spicule sizes (the two megasclere forms are not different in size, the larger anisochelae are only up to 115 µm). Details on the skeleton and the surface are not given. It is not conspecific with the
<collectingRegion id="4E7EF9C4FF9FFF8E4AE4C9C1FEAB0CA4" box="[196,307,1925,1950]" country="United States of America" name="California" pageId="2" pageNumber="51">California</collectingRegion>
species, which is regarded as chimeric and not a
<taxonomicName id="4BBA4CA5FF9FFF8E495EC9C1FC570CA4" authorityName="Gray" authorityYear="1867" box="[894,975,1925,1950]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF9FFF8E495EC9C1FC570CA4" box="[894,975,1925,1950]" italics="true" pageId="2" pageNumber="51">Mycale</emphasis>
</taxonomicName>
species by Hajdu (1994). Apparently it concerns a
<emphasis id="BECEEB34FF9FFF8E4B36C9EEFEE70CFB" box="[278,383,1962,1985]" italics="true" pageId="2" pageNumber="51">Topsentia</emphasis>
(
<taxonomicName id="4BBA4CA5FF9FFF8E4BB0C9ECFDDC0CFB" box="[400,580,1960,1985]" class="Demospongiae" family="Halichondriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Halichondrida" pageId="2" pageNumber="51" phylum="Porifera" rank="family">Halichondriidae</taxonomicName>
) or a
<taxonomicName id="4BBA4CA5FF9FFF8E48A8C9ECFC740CFB" authorityName="Ehlers" authorityYear="1870" box="[648,1004,1960,1985]" class="Demospongiae" family="Heteroxyidae" genus="Myrmekioderma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Halichondrida" pageId="2" pageNumber="51" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF9FFF8E48A8C9ECFCA50CFB" box="[648,829,1960,1985]" italics="true" pageId="2" pageNumber="51">Myrmekioderma</emphasis>
(Heteroxyidae)
</taxonomicName>
contaminated with spicules of
<taxonomicName id="4BBA4CA5FF9FFF8E4F6BC9ECFDDD0CDF" authority="(Lambe, 1905)" authorityName="Lambe" authorityYear="1905" baseAuthorityName="Lambe" baseAuthorityYear="1905" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="2" pageNumber="51" phylum="Porifera" rank="species" species="bellabellensis" subGenus="Carmia">
<emphasis id="BECEEB34FF9FFF8E4F6BC9ECFE010CDF" italics="true" pageId="2" pageNumber="51">Mycale (Carmia) bellabellensis</emphasis>
(
<bibRefCitation id="E82B4AD7FF9FFF8E4B87C988FDA50CDF" box="[423,573,1996,2021]" pageId="2" pageNumber="51" refId="ref6296" refString="Lambe, L. M. (1905) A new recent marine sponge (Esperella Bellabellensis) from the Pacific Coast of Canada. Ottawa Naturalist, 19 (1), 14 - 15. Available from: https: // www. biodiversitylibrary. org / item / 95226 # page / 20 / mode / 1 up (accessed 30 August 2018)" type="journal article">Lambe, 1905</bibRefCitation>
)
</taxonomicName>
. It is likely that a new name for the Korean specimen will have to be proposed.
</paragraph>
<caption id="D8C567AEFF9EFF8F4AB7C816FF4A0DD8" httpUri="https://zenodo.org/record/1454375/files/figure.png" pageId="3" pageNumber="52" startId="3.[151,247,1618,1640]" targetBox="[151,1435,193,1597]" targetPageId="3">
<paragraph id="8C053726FF9EFF8F4AB7C816FF4A0DD8" blockId="3.[151,1436,1618,1762]" pageId="3" pageNumber="52">
<emphasis id="BECEEB34FF9EFF8F4AB7C816FE890D5D" bold="true" box="[151,273,1618,1640]" pageId="3" pageNumber="52">FIGURE 1.</emphasis>
<taxonomicName id="4BBA4CA5FF9EFF8F4B38C816FD110D52" authority="Van Soest, 2018" authorityName="Van Soest" authorityYear="2018" box="[280,649,1618,1640]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="3" pageNumber="52" phylum="Porifera" rank="species" species="klausjanusorum" status="sp. nov." subGenus="Oxymycale">
<emphasis id="BECEEB34FF9EFF8F4B38C816FD110D52" box="[280,649,1618,1640]" italics="true" pageId="3" pageNumber="52">Mycale (Oxymycale) klausjanusorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5FD564FFF9EFF8F48B0C816FD470D5D" box="[656,735,1618,1639]" pageId="3" pageNumber="52" rank="species">
<emphasis id="BECEEB34FF9EFF8F48B0C816FD470D5D" bold="true" box="[656,735,1618,1639]" pageId="3" pageNumber="52">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype ZMA Por. 0 9452, a, habitus encrusting a piece of limestone rubble (scale bar = 1 cm), b–c, light microscope cross sections of peripheral region (b, overview, c, detailed view), showing bouquets of mycalostyles resting with their base on an ectosomal tangential skeleton consisting of smaller oxeas, carried by subsectosomal spicule tracts, d, cross section of deeper choanosomal skeleton showing thick spicule tracts dividing into thinner tracts.
</paragraph>
</caption>
<paragraph id="8C053726FF9EFF8F4AE7C954FD910CE6" blockId="3.[151,1437,1807,2012]" pageId="3" pageNumber="52">
Two other
<taxonomicName id="4BBA4CA5FF9EFF8F4B65C94BFE5A0C12" authorityName="Hentschel" authorityYear="1929" box="[325,450,1807,1832]" class="Demospongiae" family="Mycalidae" genus="Oxymycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="3" pageNumber="52" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF9EFF8F4B65C94BFE5A0C12" box="[325,450,1807,1832]" italics="true" pageId="3" pageNumber="52">Oxymycale</emphasis>
</taxonomicName>
species have been described from
<collectingCountry id="F4AD77B6FF9EFF8F4977C94BFC700C1D" box="[855,1000,1807,1832]" name="South Korea" pageId="3" pageNumber="52">South Korea</collectingCountry>
,
<taxonomicName id="4BBA4CA5FF9EFF8F49D6C954FADA0C12" authority="(Sim, 1982)" authorityName="Sim" authorityYear="1982" baseAuthorityName="Sim" baseAuthorityYear="1982" box="[1014,1346,1807,1832]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="3" pageNumber="52" phylum="Porifera" rank="species" species="koreana" subGenus="Oxymycale">
<emphasis id="BECEEB34FF9EFF8F49D6C954FB280C12" box="[1014,1200,1807,1832]" italics="true" pageId="3" pageNumber="52">M. (O.) koreana</emphasis>
(
<bibRefCitation id="E82B4AD7FF9EFF8F4EE2C94BFAA20C12" box="[1218,1338,1807,1832]" pageId="3" pageNumber="52" refId="ref6905" refString="Sim, C. J. (1982) A systematic study on the marine sponges from Jeju Island. Soong Jun University Essays Papers, 12, 187 - 210." type="journal article">Sim, 1982</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4BBA4CA5FF9EFF8F4F5DC954FE750C76" authority="(Sim &amp; Lee, 1998)" authorityName="Sim &amp; Lee" authorityYear="1998" baseAuthorityName="Sim &amp; Lee" baseAuthorityYear="1998" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="3" pageNumber="52" phylum="Porifera" rank="species" species="rhoi" subGenus="Oxymycale">
<emphasis id="BECEEB34FF9EFF8F4F5DC954FF660C76" italics="true" pageId="3" pageNumber="52">M. (O.) rhoi</emphasis>
(
<bibRefCitation id="E82B4AD7FF9EFF8F4B31C977FE7D0C76" box="[273,485,1843,1868]" pageId="3" pageNumber="52" refId="ref6937" refString="Sim, C. J. &amp; Lee, K. J. (1998) Three new species of Poecilosclerid sponge from Korea. Korean Journal of Biological Sciences, 2, 21 - 26. https: // doi. org / 10.1080 / 12265071.1998.9647386" type="journal article">Sim &amp; Lee, 1998</bibRefCitation>
)
</taxonomicName>
. The first is described as erect, and with skeleton
<taxonomicName id="4BBA4CA5FF9EFF8F4E66C970FB030C71" authorityName="Gray" authorityYear="1867" box="[1094,1179,1844,1867]" class="Demospongiae" family="Mycalidae" genus="Carmia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="3" pageNumber="52" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF9EFF8F4E66C970FB030C71" box="[1094,1179,1844,1867]" italics="true" pageId="3" pageNumber="52">Carmia</emphasis>
</taxonomicName>
-like without a proper ectosomal skeleton, megascleres are oxeas only, up to 539 µm, and anisochelae up to 88 µm, clearly unlike the new species. The latter Korean species is described as having a tangential ectosomal skeleton, oxea megascleres only, up to 660 µm, and anisochelae up to 110 µm, and thus similar to the new species. Nevertheless, the overall suite of characters is sufficiently distinct.
</paragraph>
<paragraph id="8C053726FF99FF884AE7CEDCFB7B0AB2" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="53">
From the
<typeStatus id="53018984FF99FF884B11CEDCFEF80B8A" box="[305,352,152,176]" pageId="4" pageNumber="53">type</typeStatus>
species, Arctic
<taxonomicName id="4BBA4CA5FF99FF88482ECEDDFC0E0B8A" authority="(Schmidt, 1874)" authorityName="Schmidt" authorityYear="1874" baseAuthorityName="Schmidt" baseAuthorityYear="1874" box="[526,918,151,176]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="intermedia" subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF88482ECEDDFD450B8A" box="[526,733,151,176]" italics="true" pageId="4" pageNumber="53">M. (O.) intermedia</emphasis>
(
<bibRefCitation id="E82B4AD7FF99FF8848CBCED3FC160B8A" box="[747,910,151,176]" pageId="4" pageNumber="53" refId="ref6873" refString="Schmidt, O. (1874) Kieselspongien. Die Zweite deutsche Nordpolfahrt in 1869 - 70 (' Germania' Expedition), 2 (13), 429 - 433." type="journal article">Schmidt, 1874</bibRefCitation>
)
</taxonomicName>
(p. 433, fig. 10) as redescribed by
<bibRefCitation id="E82B4AD7FF99FF884F33CEDCFE800BEE" pageId="4" pageNumber="53" refId="ref7228" refString="Van Soest, R. W. M. &amp; Hajdu, E. (2002) Family Mycalidae Lundbeck, 1905. In: Hooper, J. N. A. &amp; Van Soest, R. W. M. (Eds.), Systema Porifera. A guide to the classification of sponges 1. Kluwer Academic / Plenum Publishers, New York, Boston, Dordrecht, London and Moscow, pp. 669 - 690. https: // doi. org / 10.1007 / 978 - 1 - 4615 - 0747 - 5 _ 73" type="book chapter">Van Soest &amp; Hajdu 2002</bibRefCitation>
(based mostly on Thiele’s 1903b description of a slide from the Weltner collection, and on Koltun’s 1959 descriptions of
<specimenCount id="9ABCFCAFFF99FF884B1ECE9BFE5F0BC2" box="[318,455,223,248]" pageId="4" pageNumber="53" type="generic">5 specimens</specimenCount>
, including the junior synonym
<taxonomicName id="4BBA4CA5FF99FF884939CE9BFBE30BC2" authority="Koltun, 1952" authorityName="Koltun" authorityYear="1952" box="[793,1147,223,248]" class="Demospongiae" family="Mycalidae" genus="Oxymycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="wagini">
<emphasis id="BECEEB34FF99FF884939CE9BFC7B0BC2" box="[793,995,223,248]" italics="true" pageId="4" pageNumber="53">Oxymycale wagini</emphasis>
<bibRefCitation id="E82B4AD7FF99FF8849C9CE9BFBE30BC2" box="[1001,1147,223,248]" pageId="4" pageNumber="53" refId="ref6056" refString="Koltun, V. M. (1952) [New sponges from the northern seas]. Uchenye Zapiski Leningradskogo Ordena Lenina Gosudarstvennogo, 31 (145), 125 - 129. [in Russian, title translated in English]" type="journal article">Koltun, 1952</bibRefCitation>
</taxonomicName>
) the new species differs in the weak development of the skeleton of Schmidt’s species of which the ectosomal part is
<taxonomicName id="4BBA4CA5FF99FF884E57CF41FB530A26" authorityName="Gray" authorityYear="1867" box="[1143,1227,261,284]" class="Demospongiae" family="Mycalidae" genus="Carmia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF99FF884E57CF41FB530A26" box="[1143,1227,261,284]" italics="true" pageId="4" pageNumber="53">Carmia</emphasis>
</taxonomicName>
-like in having thin choanosomal tracts and no mention of a tangential spicule arrangement, and in the apparent lack of sigmas and trichodragmas. Megascleres and anisochelae I are about half as large as those of
<taxonomicName id="4BBA4CA5FF99FF884E21CF09FA9F0A5F" authority="Van Soest, 2018" authorityName="Van Soest" authorityYear="2018" box="[1025,1287,332,357]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="klausjanusorum" status="sp. nov." subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF884E21CF09FA9F0A5F" box="[1025,1287,332,357]" italics="true" pageId="4" pageNumber="53">M. (O.) klausjanusorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5FD564FFF99FF884F2DCF08FAFC0A5F" box="[1293,1380,332,357]" pageId="4" pageNumber="53" rank="species">
<emphasis id="BECEEB34FF99FF884F2DCF08FAFC0A5F" bold="true" box="[1293,1380,332,357]" pageId="4" pageNumber="53">sp. nov.</emphasis>
</taxonomicNameLabel>
Like the latter,
<taxonomicName id="4BBA4CA5FF99FF884B23CF35FE4A0AB2" baseAuthorityName="Schmidt" baseAuthorityYear="1874" box="[259,466,367,392]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="intermedia" subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF884B23CF35FE4A0AB2" box="[259,466,367,392]" italics="true" pageId="4" pageNumber="53">M. (O.) intermedia</emphasis>
</taxonomicName>
is reported from depths down to
<specimenCount id="9ABCFCAFFF99FF88491CCF2BFC110ABD" box="[828,905,367,392]" pageId="4" pageNumber="53" type="generic">
<quantity id="4B429AC3FF99FF88491CCF2BFC1D0ABD" box="[828,901,367,392]" metricMagnitude="2" metricUnit="m" metricValue="3.25" pageId="4" pageNumber="53" unit="m" value="325.0">325 m</quantity>
.
</specimenCount>
The species is not well-known.
</paragraph>
<paragraph id="8C053726FF99FF884AE7CFD0FB2209F6" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="53">
Further species assigned to
<taxonomicName id="4BBA4CA5FF99FF884BDCCFD0FD7C0A97" authority="(Oxymycale)" authorityName="Gray" authorityYear="1867" baseAuthorityName="Oxymycale" box="[508,740,404,429]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF99FF884BDCCFD0FDD60A97" box="[508,590,404,429]" italics="true" pageId="4" pageNumber="53">Mycale</emphasis>
(
<emphasis id="BECEEB34FF99FF88487FCFD0FD7C0A97" box="[607,740,404,429]" italics="true" pageId="4" pageNumber="53">Oxymycale)</emphasis>
</taxonomicName>
are progressively more unlike the new species: South African
<taxonomicName id="4BBA4CA5FF99FF884AB7CFFDFD140AEA" authority="Samaai &amp; Gibbons, 2005" authorityName="Samaai &amp; Gibbons" authorityYear="2005" box="[151,652,439,464]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="stephensae" subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF884AB7CFFDFEF30AEA" box="[151,363,439,464]" italics="true" pageId="4" pageNumber="53">M. (O.) stephensae</emphasis>
<bibRefCitation id="E82B4AD7FF99FF884B52CFF3FD140AEA" box="[370,652,439,464]" pageId="4" pageNumber="53" refId="ref6775" refString="Samaai, T. &amp; Gibbons, M. J. (2005) Demospongiae taxonomy and biodiversity of the Benguela region on the west coast of South Africa. African Natural History, 1, 1 - 96. Available from: https: // journals. co. za / content / afnathist / 1 / 1 / EJC 17340 (Accessed 30 Aug. 2018)" type="journal article">Samaai &amp; Gibbons, 2005</bibRefCitation>
</taxonomicName>
lacks trichodragmas. Antarctic
<taxonomicName id="4BBA4CA5FF99FF8849D0CFFDFAF50AEA" authority="Kirkpatrick, 1907" authorityName="Kirkpatrick" authorityYear="1907" box="[1008,1389,439,464]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="acerata" subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF8849D0CFFDFB380AEA" box="[1008,1184,440,464]" italics="true" pageId="4" pageNumber="53">M. (O.) acerata</emphasis>
<bibRefCitation id="E82B4AD7FF99FF884E86CFF3FAF50AEA" box="[1190,1389,439,464]" pageId="4" pageNumber="53" refId="ref6009" refString="Kirkpatrick, R. (1907) Preliminary Report on the Monaxonellida of the National Antarctic Expedition. Annals and Magazine of Natural History, Series 7, 20 (117), 271 - 291. https: // doi. org / 10.1080 / 00222930709487333" type="journal article">Kirkpatrick, 1907</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4BBA4CA5FF99FF884AB7CF99FD940ACF" authority="Koltun, 1964" authorityName="Koltun" authorityYear="1964" box="[151,524,476,501]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="tylotornota" subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF884AB7CF99FEE80ACF" box="[151,368,476,501]" italics="true" pageId="4" pageNumber="53">M. (O.) tylotornota</emphasis>
<bibRefCitation id="E82B4AD7FF99FF884B57CF98FD940ACF" box="[375,524,476,501]" pageId="4" pageNumber="53" refId="ref6213" refString="Koltun, V. M. (1964) Sponges of the Antarctic. 1 Tetraxonida and Cornacuspongida. In: Pavlovskii, E. P., Andriyashev, A. P. &amp; Ushakov, P. V. (Eds.), Biological Reports of the Soviet Antarctic Expedition (1955 - 1958). Akademya Nauk SSSR, 6 - 133, 443 - 448. [English translation, 1966, Israel Program for Scientific Translation]" type="journal article">Koltun, 1964</bibRefCitation>
</taxonomicName>
lack sigmas, Western Pacific
<taxonomicName id="4BBA4CA5FF99FF884940CF99FAF70ACF" authority="(De Laubenfels, 1954)" authorityName="De Laubenfels" authorityYear="1954" baseAuthorityName="De Laubenfels" baseAuthorityYear="1954" box="[864,1391,476,501]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="strongylophora" subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF884940CF99FBF00ACF" box="[864,1128,476,501]" italics="true" pageId="4" pageNumber="53">M. (O.) strongylophora</emphasis>
(
<bibRefCitation id="E82B4AD7FF99FF884E58CF98FAFF0ACF" box="[1144,1383,476,501]" pageId="4" pageNumber="53" refId="ref5312" refString="De Laubenfels, M. W. (1954) The Sponges of the West-Central Pacific. Oregon State Monographs, Studies in Zoology, 7, i - x, 1 - 306. Available from: https: // www. biodiversitylibrary. org / item / 28986 # page / 3 / mode / 1 up (accessed 30 August 2018)" type="book chapter">De Laubenfels, 1954</bibRefCitation>
)
</taxonomicName>
has strongyles for megascleres. Western Pacific
<taxonomicName id="4BBA4CA5FF99FF8848ABCC45FBC50922" authority="(De Laubenfels, 1954)" authorityName="De Laubenfels" authorityYear="1954" baseAuthorityName="De Laubenfels" baseAuthorityYear="1954" box="[651,1117,511,536]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="stecarmia" subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF8848ABCC45FCCD0922" box="[651,853,512,536]" italics="true" pageId="4" pageNumber="53">M. (O.) stecarmia</emphasis>
(
<bibRefCitation id="E82B4AD7FF99FF884946CC44FBCD0922" box="[870,1109,511,536]" pageId="4" pageNumber="53" refId="ref5312" refString="De Laubenfels, M. W. (1954) The Sponges of the West-Central Pacific. Oregon State Monographs, Studies in Zoology, 7, i - x, 1 - 306. Available from: https: // www. biodiversitylibrary. org / item / 28986 # page / 3 / mode / 1 up (accessed 30 August 2018)" type="book chapter">De Laubenfels, 1954</bibRefCitation>
)
</taxonomicName>
has very small oxeas (up to 154 µm) for megascleres, the anisochelae are only up to 42 µm, it may possess toxas (not considered proper), together making it a very unlikely relative of the new species. Finally,
<collectingRegion id="4E7EF9C4FF99FF88498FCC03FB98095A" box="[943,1024,583,608]" country="United States of America" name="Florida" pageId="4" pageNumber="53">Florida</collectingRegion>
<taxonomicName id="4BBA4CA5FF99FF884E26CC0DFA04095A" authority="(Schmidt, 1870)" authorityName="Schmidt" authorityYear="1870" baseAuthorityName="Schmidt" baseAuthorityYear="1870" box="[1030,1436,583,608]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="renieroides" subGenus="Oxymycale">
<emphasis id="BECEEB34FF99FF884E26CC0DFB46095A" box="[1030,1246,583,608]" italics="true" pageId="4" pageNumber="53">M. (O.) renieroides</emphasis>
(
<bibRefCitation id="E82B4AD7FF99FF884ECDCC03FA0C095A" box="[1261,1428,583,608]" pageId="4" pageNumber="53" refId="ref6844" refString="Schmidt, O. (1870) Grundzuge einer Spongien-Fauna des atlantischen Gebietes. Wilhelm Engelmann, Leipzig, 2 (iii - iv) + 88 pp." type="book chapter">Schmidt, 1870</bibRefCitation>
)
</taxonomicName>
cannot be properly assessed as a
<taxonomicName id="4BBA4CA5FF99FF884831CC28FDFC09BF" authorityName="Gray" authorityYear="1867" box="[529,612,620,645]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF99FF884831CC28FDFC09BF" box="[529,612,620,645]" italics="true" pageId="4" pageNumber="53">Mycale</emphasis>
</taxonomicName>
according to
<bibRefCitation id="E82B4AD7FF99FF884921CC28FB8A09BE" box="[769,1042,620,645]" pageId="4" pageNumber="53" refId="ref5621" refString="Hajdu, E. &amp; Rutzler, K. (1998) Sponges, genus Mycale (Poecilosclerida: Demospongiae: Porifera), from a Caribbean mangrove and comments on subgeneric classification. Proceedings of the Biological Socety of Washington, 111 (4), 737 - 773. Available from: https: // www. biodiversitylibrary. org / item / 107585 # page / 275 / mode / 1 up (accessed 30 August 2018)" type="journal article">Hajdu &amp; Rützler (1998)</bibRefCitation>
after examination of a slide in the Natural History Museum made by Schmidt. The latter species is apparently still represented by
<typeStatus id="53018984FF99FF884EE9CCD4FB610992" box="[1225,1273,656,680]" pageId="4" pageNumber="53">type</typeStatus>
material, both dried and in alcohol, in the collection of the Museum of Comparative Zoology, Harvard,
<collectingCountry id="F4AD77B6FF99FF884E5BCCF0FB2F09F6" box="[1147,1207,692,716]" name="United States of America" pageId="4" pageNumber="53">USA</collectingCountry>
.
</paragraph>
<paragraph id="8C053726FF99FF884AE7CC93FD40089F" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="53">
Similarities with the new species are evident in
<taxonomicName id="4BBA4CA5FF99FF8848D5CC93FB4809CA" authority="Koltun, 1958" authorityName="Koltun" authorityYear="1958" box="[757,1232,727,752]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="longistyla" subGenus="Mycale">
<emphasis id="BECEEB34FF99FF8848D5CC93FBA809CA" box="[757,1072,727,752]" italics="true" pageId="4" pageNumber="53">Mycale (Mycale) longistyla</emphasis>
<bibRefCitation id="E82B4AD7FF99FF884E19CC93FB4809CA" box="[1081,1232,727,752]" pageId="4" pageNumber="53" refId="ref6099" refString="Koltun, V. M. (1958) [Cornacuspongia of sea waters washing the South Sakhalin and the South Kurile Island region]. Issledovaniya dal'nevostochnykh morei SSR, 5, 42 - 77. [in Russian]" type="journal article">Koltun, 1958</bibRefCitation>
</taxonomicName>
from
<quantity id="4B429AC3FF99FF884F3ACC9CFA0409D5" box="[1306,1436,728,752]" metricMagnitude="2" metricUnit="m" metricValue="2.01" metricValueMax="2.7" metricValueMin="1.3199999999999998" pageId="4" pageNumber="53" unit="m" value="201.0" valueMax="270.0" valueMin="132.0">132–270 m</quantity>
depth in the North Pacific, where the megascleres protrude far beyond the tangential ectosomal skeleton. These megascleres consist of three categories, large styles of up to
<quantity id="4B429AC3FF99FF884963CD5BFC02080D" box="[835,922,799,824]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="53" unit="mm" value="3.5">3.5 mm</quantity>
<date id="F80411E6FF99FF884981CD64FC4B0802" box="[929,979,800,824]" pageId="4" pageNumber="53" value="1932-10">x 32</date>
µm, intermediate styles of 1144–
<date id="F80411E6FF99FF884F6FCD5BFF2A0866" pageId="4" pageNumber="53" value="1768-10-23">1768 x 23</date>
–32 µm, and small strongyles of 416–530
<date id="F80411E6FF99FF8848AACD01FD4E0866" box="[650,726,836,860]" pageId="4" pageNumber="53" value="2011-10-09">x 9–11</date>
µm, differentiated in a way reminding of the megascleres of the new species, but upper sizes are clearly in excess of the latter. Assignment of this species to the subgenus
<taxonomicName id="4BBA4CA5FF99FF884F69CD23FEBE089F" authority="(Oxymycale)" authorityName="Gray" authorityYear="1867" baseAuthorityName="Oxymycale" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF99FF884F69CD23FEBE089F" italics="true" pageId="4" pageNumber="53">Mycale (Oxymycale)</emphasis>
</taxonomicName>
may be contemplated (but see below).
</paragraph>
<paragraph id="8C053726FF99FF884AE7CDEBFD5508D7" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="53">
Comparison is also warranted with several species not currently assigned to the subgenus
<taxonomicName id="4BBA4CA5FF99FF884E94CDEBFA0408F2" authority="(Oxymycale)" authorityName="Gray" authorityYear="1867" baseAuthorityName="Oxymycale" box="[1204,1436,943,968]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF99FF884E94CDEBFA0408F2" box="[1204,1436,943,968]" italics="true" pageId="4" pageNumber="53">Mycale (Oxymycale)</emphasis>
</taxonomicName>
showing skeletal similarities with the new species.
</paragraph>
</subSubSection>
<subSubSection id="C4A064ADFF99FF884AE7CDB3FD2D0E6F" pageId="4" pageNumber="53" type="reference_group">
<paragraph id="8C053726FF99FF884AE7CDB3FB240FFF" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="53">
<taxonomicName id="4BBA4CA5FF99FF884AE7CDB3FDF50F2A" ID-CoL="44RYH" authority="Burton, 1959" authorityName="Burton" authorityYear="1959" box="[199,621,1015,1040]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="topsenti" subGenus="Mycale">
<emphasis id="BECEEB34FF99FF884AE7CDB3FE410F2A" box="[199,473,1015,1040]" italics="true" pageId="4" pageNumber="53">Mycale (Mycale) topsenti</emphasis>
<bibRefCitation id="E82B4AD7FF99FF884BFFCDBCFDF50F2A" box="[479,621,1015,1040]" pageId="4" pageNumber="53" refId="ref4926" refString="Burton, M. (1959) Sponges. In: Scientific Reports. John Murray Expedition 1933 - 34. 10 (5). British Museum (Natural History), London, pp. 151 - 281. Available from: https: // www. biodiversitylibrary. org / item / 195785 # page / 189 / mode / 1 up (accessed 30 August 2018)" type="book chapter">Burton, 1959</bibRefCitation>
</taxonomicName>
from
<quantity id="4B429AC3FF99FF88488ECDB3FC980F35" box="[686,768,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="1.0459999999999998" pageId="4" pageNumber="53" unit="m" value="1046.0">1046 m</quantity>
depth off the coast of
<collectingCountry id="F4AD77B6FF99FF8849CECDBCFBA90F2A" box="[1006,1073,1016,1040]" name="Oman" pageId="4" pageNumber="53">Oman</collectingCountry>
shares the structure of the surface skeleton with the new species, with ‘diffuse brushes’ of megascleres projecting beyond the tangential ectosomal skeleton. Megascleres are clearly different in consisting of a single size of subtylostyles of 900
<date id="F80411E6FF99FF884EB6CA04FB5D0F62" box="[1174,1221,1088,1112]" pageId="4" pageNumber="53" value="1918-10">x 18</date>
µm, and among the microscleres the largest anisochelae are up to 88 µm. The skeletal properties described for New Caledonian
<taxonomicName id="4BBA4CA5FF99FF884F6CCA20FDC80F9A" ID-CoL="44RY4" authority="Levi &amp; Levi, 1983" authorityName="Levi &amp; Levi" authorityYear="1983" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="myriasclera" subGenus="Mycale">
<emphasis id="BECEEB34FF99FF884F6CCA20FE180F9A" italics="true" pageId="4" pageNumber="53">Mycale (Mycale) myriasclera</emphasis>
<bibRefCitation id="E82B4AD7FF99FF884BA6CAC3FDC80F9A" box="[390,592,1159,1184]" pageId="4" pageNumber="53" refId="ref6408" refString="Levi, C. &amp; Levi, P. (1983) Demosponges bathyales recoltees par le N / O'Vauban' au sud de la Nouvelle-Caledonie. Bulletin du Museum National d'Histoire Naturelle, 4 A, 5 (4), 931 - 997." type="journal article">Lévi &amp; Lévi, 1983</bibRefCitation>
</taxonomicName>
, from
<quantity id="4B429AC3FF99FF8848B6CACCFD420F9A" box="[662,730,1160,1184]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="53" unit="m" value="300.0">300 m</quantity>
depth, include also megascleres protruding beyond the tangential ectosomal skeleton, but a clear difference is the absence of sigmas in the New Caledonian species.
</paragraph>
<paragraph id="8C053726FF99FF884AE7CA8BFD2D0E6F" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="53">
Several
<taxonomicName id="4BBA4CA5FF99FF884B02CA8BFEED0FD2" ID-CoL="84QN6" authorityName="Gray" authorityYear="1867" box="[290,373,1231,1256]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="genus">
<emphasis id="BECEEB34FF99FF884B02CA8BFEED0FD2" box="[290,373,1231,1256]" italics="true" pageId="4" pageNumber="53">Mycale</emphasis>
</taxonomicName>
species with anisochelae I of the same size or larger as those of the new species reported from the Indo-West Pacific region were also reviewed, e.g.
<taxonomicName id="4BBA4CA5FF99FF884937CAB0FB520E37" ID-CoL="73QRS" authority="Gray, 1867" authorityName="Gray" authorityYear="1867" box="[791,1226,1268,1293]" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="grandis" subGenus="Mycale">
<emphasis id="BECEEB34FF99FF884937CAB0FBA20E37" box="[791,1082,1268,1293]" italics="true" pageId="4" pageNumber="53">Mycale (Mycale) grandis</emphasis>
<bibRefCitation id="E82B4AD7FF99FF884E66CAB0FB520E37" box="[1094,1226,1268,1293]" pageId="4" pageNumber="53" refId="ref5377" refString="Gray, J. E. (1867) Notes on the arrangement of sponges, with the descriptions of some new genera. Proceedings of the Zoological Society of London, 1867 (2), 492 - 558. Available from: https: // www. biodiversitylibrary. org / item / 93424 # page / 514 / mode / 1 up (accessed 30 August 2018)" type="journal article">Gray, 1867</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4BBA4CA5FF99FF884EF9CAB0FE0E0E0A" ID-CoL="44RWW" authority="Levi, 1963" authorityName="Levi" authorityYear="1963" class="Demospongiae" family="Mycalidae" genus="Mycale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Poecilosclerida" pageId="4" pageNumber="53" phylum="Porifera" rank="species" species="anisochela" subGenus="Mycale">
<emphasis id="BECEEB34FF99FF884EF9CAB0FE890E0A" italics="true" pageId="4" pageNumber="53">Mycale (Mycale) anisochela</emphasis>
<bibRefCitation id="E82B4AD7FF99FF884B39CB53FE0E0E0A" box="[281,406,1303,1328]" pageId="4" pageNumber="53" refId="ref6363" refString="Levi, C. (1963) Spongiaires d'Afrique du Sud. (1) Poecilosclerides. Transactions of the Royal Society of South Africa, 37 (1), 1 - 72. https: // doi. org / 10.1080 / 00359196309519054" type="journal article">Lévi, 1963</bibRefCitation>
</taxonomicName>
. However, these species differ clearly in the structure of the surface skeleton, the shape of the megascleres and the additional microscleres.
</paragraph>
</subSubSection>
</treatment>
</document>
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