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<document id="2BBEA51F9B2B69C74F94223F3B0CFEB5" ID-DOI="10.21248/contrib.entomol.67.2.275-318" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742497" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638268338673" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2017" docId="03F6C753FFFDFFBAFC83D826FD79FAE4" docLanguage="en" docName="ContribEntomol.67.2.275-318.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Eleusis peruviana Irmler 2017, spec. nov." docType="treatment" docUuid="819FB601-081E-4EEF-BAD3-2363C6D299CF" docUuidSource="ZooBank" docVersion="5" lastPageNumber="300" masterDocId="FFCFBF2BFFE5FFA3FFFFD92CFFDAFF9C" masterDocTitle="The Neotropical species of the genus Eleusis LAPORTE, 1835 with description of the new genus Eleusima (Coleoptera: Staphylinidae: Osoriinae)" masterLastPageNumber="318" masterPageNumber="275" pageNumber="299" updateTime="1699437037312" updateUser="plazi">
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<mods:title id="A96BE3DED86C3C65CA1734A1D087000B">The Neotropical species of the genus Eleusis LAPORTE, 1835 with description of the new genus Eleusima (Coleoptera: Staphylinidae: Osoriinae)</mods:title>
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<mods:namePart id="749B0CFE40E8239786D5017CDAB40E0F">Irmler, Ulrich</mods:namePart>
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<mods:typeOfResource id="7B8FCA62A082DC26567BF7BF8348701B">text</mods:typeOfResource>
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<mods:title id="F75BA91551FCE8092CFF5F5F30678263">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="7BD28946F9CDE500F4C0651D55E9194B">
<mods:date id="AF92BF75B6B9C19BEAF4B0CBB70DA97E">2017</mods:date>
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<mods:number id="436DFD4796E68D67AC4F0AD93B3EECD4">67</mods:number>
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<mods:identifier id="ABC59A768A88672773796047532D0328" type="DOI">10.21248/contrib.entomol.67.2.275-318</mods:identifier>
<mods:identifier id="1BFA237B12C3B7954BC3C9CB3D134E90" type="ISSN">0005-805X</mods:identifier>
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<treatment id="03F6C753FFFDFFBAFC83D826FD79FAE4" ID-DOI="http://doi.org/10.5281/zenodo.5742698" ID-Zenodo-Dep="5742698" LSID="urn:lsid:zoobank.org:act:819FB601-081E-4EEF-BAD3-2363C6D299CF" httpUri="http://treatment.plazi.org/id/03F6C753FFFDFFBAFC83D826FD79FAE4" lastPageId="25" lastPageNumber="300" pageId="24" pageNumber="299">
<subSubSection id="C34525CEFFFDFFBBFC83D826FA5BFEDA" pageId="24" pageNumber="299" type="nomenclature">
<paragraph id="8BE07645FFFDFFBBFC83D826FB45FEB9" blockId="24.[892,1409,266,358]" box="[892,1183,266,293]" pageId="24" pageNumber="299">
<heading id="D0A8C129FFFDFFBBFC83D826FB45FEB9" box="[892,1183,266,293]" fontSize="12" level="2" pageId="24" pageNumber="299" reason="2">
<taxonomicName id="4C5F0DC6FFFDFFBBFC83D826FBF5FEB9" ID-CoL="398R7" authority="Irmler, 2017" authorityName="Irmler" authorityYear="2017" box="[892,1071,266,293]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="peruviana" status="sp. nov.">
<emphasis id="B92BAA57FFFDFFBBFC83D826FBF5FEB9" box="[892,1071,266,293]" italics="true" pageId="24" pageNumber="299">Eleusis peruviana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A218172CFFFDFFBBFBC9D826FB45FEB9" box="[1078,1183,266,293]" pageId="24" pageNumber="299" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BE07645FFFDFFBBFC83D81EFA5BFEDA" blockId="24.[892,1409,266,358]" box="[892,1409,306,326]" pageId="24" pageNumber="299">
<uri id="FFCE7A47FFFDFFBBFC83D81EFA5BFEDA" box="[892,1409,306,326]" pageId="24" pageNumber="299">
urn:lsid:zoobank.org: act:
<uuid id="FFF94C90FFFDFFBBFBD5D81EFA5BFEDA" box="[1066,1409,306,326]" pageId="24" pageNumber="299">819FB601-081E-4EEF-BAD3-2363C6D299CF</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFFDFFBBFC83D87EFC2FFEFA" box="[892,1013,338,358]" pageId="24" pageNumber="299" type="description">
<paragraph id="8BE07645FFFDFFBBFC83D87EFC2FFEFA" blockId="24.[892,1409,266,358]" box="[892,1013,338,358]" pageId="24" pageNumber="299">
<figureCitation id="13646AC0FFFDFFBBFC83D87EFC68FEFA" box="[892,946,338,358]" captionStart="Fig" captionStartId="35.[174,207,2051,2073]" captionTargetBox="[184,1398,974,2044]" captionTargetId="figure-101@35.[174,1421,950,2050]" captionTargetPageId="35" captionText="Fig. 2427: E. peruviana (24), E. panamaensis (25); E. frater (26), E. elongata (27): aedeagus in lateral (A) and ventral (B) view," figureDoi="http://doi.org/10.5281/zenodo.5742511" httpUri="https://zenodo.org/record/5742511/files/figure.png" pageId="24" pageNumber="299">Figs 24</figureCitation>
A-C, 30G
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFFDFFBBFCA9D8ACFC63FD84" pageId="24" pageNumber="299" type="materials_examined">
<paragraph id="8BE07645FFFDFFBBFCA9D8ACFC63FD84" blockId="24.[854,1460,383,536]" pageId="24" pageNumber="299">
<emphasis id="B92BAA57FFFDFFBBFCA9D8ACFC35FE0B" bold="true" box="[854,1007,383,408]" pageId="24" pageNumber="299">Type material</emphasis>
:
<materialsCitation id="3B377C18FFFDFFBBFC02D8ACFC6FFD84" collectingDate="1948-03" collectionCode="AMNH, IRSNB" collectorName="Prov. Tama &amp; Dept. Of &amp; Utcuyacu &amp; Agua Dulce &amp; F. Woytkovski" country="Peru" location="Junin" pageId="24" pageNumber="299" specimenCount="2" stateProvince="Junin" typeStatus="holotype">
male,
<typeStatus id="54E4C8E7FFFDFFBBFBC0D8ACFB7FFE04" box="[1087,1189,384,408]" pageId="24" pageNumber="299" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34836D5FFFDFFBBFB4ED8ACFB3CFE0B" box="[1201,1254,384,407]" name="Peru" pageId="24" pageNumber="299">Peru</collectingCountry>
:
<collectorName id="26AA1393FFFDFFBBFB0ED8ACFAB6FE0B" box="[1265,1388,384,407]" pageId="24" pageNumber="299">Prov. Tama</collectorName>
,
<collectorName id="26AA1393FFFDFFBBFA87D8ACFCA8FE24" pageId="24" pageNumber="299">Dept. Of</collectorName>
<collectingRegion id="499BB8A7FFFDFFBBFC87D88CFC6EFE24" box="[888,948,416,440]" country="Peru" name="Junin" pageId="24" pageNumber="299">Junin</collectingRegion>
,
<collectorName id="26AA1393FFFDFFBBFC40D88CFBFFFE24" box="[959,1061,416,440]" pageId="24" pageNumber="299">Utcuyacu</collectorName>
&amp;
<collectorName id="26AA1393FFFDFFBBFBBBD88CFB19FE24" box="[1092,1219,416,440]" pageId="24" pageNumber="299">Agua Dulce</collectorName>
,
<date id="FFE15085FFFDFFBBFB31D88CFAFEFE24" box="[1230,1316,416,440]" pageId="24" pageNumber="299" value="1948-03">
<collectingDate id="EFA5A96DFFFDFFBBFB31D88CFAFEFE24" box="[1230,1316,416,440]" pageId="24" pageNumber="299" value="1948-03">III 1948</collectingDate>
</date>
, leg.
<collectorName id="26AA1393FFFDFFBBFAA6D88CFC41FE44" pageId="24" pageNumber="299">F. Woytkovski</collectorName>
(
<collectionCode id="ED4EEE80FFFDFFBBFC4BD8ECFBD7FE44" box="[948,1037,448,472]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="299">AMNH</collectionCode>
).
<typeStatus id="54E4C8E7FFFDFFBBFBD7D8ECFB52FE44" box="[1064,1160,448,472]" pageId="24" pageNumber="299" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="F34836D5FFFDFFBBFB63D8ECFB30FE44" box="[1180,1258,448,472]" name="Bolivia" pageId="24" pageNumber="299">Bolivia</collectingCountry>
: male, Yuracaris, corticale saillant a la base (bark not attached at base) (
<collectionCode id="ED4EEE80FFFDFFBBFCA2DB2CFC77FD84" box="[861,941,512,536]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="24" pageNumber="299">IRSNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFFDFFBBFCA9DB13FC3CFBAB" pageId="24" pageNumber="299" type="diagnosis">
<paragraph id="8BE07645FFFDFFBBFCA9DB13FC3CFBAB" blockId="24.[854,1459,575,1080]" pageId="24" pageNumber="299">
<emphasis id="B92BAA57FFFDFFBBFCA9DB13FC19FDCB" bold="true" box="[854,963,575,599]" pageId="24" pageNumber="299">Diagnosis</emphasis>
: The species is closely related to
<taxonomicName id="4C5F0DC6FFFDFFBBFAAEDB6CFA69FDC4" authorityName="ERICHSON" authorityYear="1840" box="[1361,1459,576,600]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B92BAA57FFFDFFBBFAAEDB6CFA69FDC4" box="[1361,1459,576,600]" italics="true" pageId="24" pageNumber="299">E. bicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F0DC6FFFDFFBBFC78DB4CFC2EFDEB" authorityName="Irmler" authorityYear="2017" box="[903,1012,608,631]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="B92BAA57FFFDFFBBFC78DB4CFC2EFDEB" box="[903,1012,608,631]" italics="true" pageId="24" pageNumber="299">E. struyvei</emphasis>
</taxonomicName>
regarding colouration. Because only
<specimenCount id="9D59BDCCFFFDFFBBFA74DB4CFC1FFD04" pageId="24" pageNumber="299" type="generic">two specimens</specimenCount>
are known, the variation of colouration is unknown. However, the different colour of the abdomen between
<typeStatus id="54E4C8E7FFFDFFBBFC44DBECFBC3FD44" box="[955,1049,704,728]" pageId="24" pageNumber="299" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E4C8E7FFFDFFBBFBA6DBECFB6CFD44" box="[1113,1206,704,728]" pageId="24" pageNumber="299" type="paratype">paratype</typeStatus>
may indicate that the variation of the colour might be as large as in the other two species. The species can be separated from
<taxonomicName id="4C5F0DC6FFFDFFBBFAAEDA2CFA69FC84" authorityName="ERICHSON" authorityYear="1840" box="[1361,1459,768,792]" class="Insecta" family="Staphylinidae" genus="Isomalus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B92BAA57FFFDFFBBFAAEDA2CFA69FC84" box="[1361,1459,768,792]" italics="true" pageId="24" pageNumber="299">E. bicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F0DC6FFFDFFBBFC76DA0CFC2CFCAB" authorityName="Irmler" authorityYear="2017" box="[905,1014,800,823]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="B92BAA57FFFDFFBBFC76DA0CFC2CFCAB" box="[905,1014,800,823]" italics="true" pageId="24" pageNumber="299">E. struyvei</emphasis>
</taxonomicName>
by the uniform punctation of the head and the dense microsculpture that is still denser than in
<taxonomicName id="4C5F0DC6FFFDFFBBFCA9DA4CFC19FCEB" authorityName="Irmler" authorityYear="2017" box="[854,963,864,887]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="B92BAA57FFFDFFBBFCA9DA4CFC19FCEB" box="[854,963,864,887]" italics="true" pageId="24" pageNumber="299">E. struyvei</emphasis>
</taxonomicName>
. The aedeagus resembles that of
<taxonomicName id="4C5F0DC6FFFDFFBBFABADA4CFA69FCEB" authorityName="Irmler" authorityYear="2017" box="[1349,1459,864,887]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="B92BAA57FFFDFFBBFABADA4CFA69FCEB" box="[1349,1459,864,887]" italics="true" pageId="24" pageNumber="299">E. struyvei</emphasis>
</taxonomicName>
by its longitudinal shape. It is still more elongate than in
<taxonomicName id="4C5F0DC6FFFDFFBBFCA9DA8CFC19FC2B" authorityName="Irmler" authorityYear="2017" box="[854,963,928,951]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="B92BAA57FFFDFFBBFCA9DA8CFC19FC2B" box="[854,963,928,951]" italics="true" pageId="24" pageNumber="299">E. struyvei</emphasis>
</taxonomicName>
. The apex is slightly sinuated in lateral aspect, whereas it is straight in
<taxonomicName id="4C5F0DC6FFFDFFBBFBABDAECFB1BFC4B" authorityName="Irmler" authorityYear="2017" box="[1108,1217,960,983]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="B92BAA57FFFDFFBBFBABDAECFB1BFC4B" box="[1108,1217,960,983]" italics="true" pageId="24" pageNumber="299">E. struyvei</emphasis>
</taxonomicName>
. Moreover, the central groove of central lobe margined by weak ridges is absent in
<taxonomicName id="4C5F0DC6FFFDFFBBFC8BDD2CFC3BFB8B" authorityName="Irmler" authorityYear="2017" box="[884,993,1024,1047]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="B92BAA57FFFDFFBBFC8BDD2CFC3BFB8B" box="[884,993,1024,1047]" italics="true" pageId="24" pageNumber="299">E. struyvei</emphasis>
</taxonomicName>
and the parameres are more elongate than in
<taxonomicName id="4C5F0DC6FFFDFFBBFC8DDD0CFC05FBAB" authorityName="Irmler" authorityYear="2017" box="[882,991,1056,1079]" class="Insecta" family="Staphylinidae" genus="Eleusis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="299" phylum="Arthropoda" rank="species" species="struyvei">
<emphasis id="B92BAA57FFFDFFBBFC8DDD0CFC05FBAB" box="[882,991,1056,1079]" italics="true" pageId="24" pageNumber="299">E. struyvei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFFDFFBAFCA9DD73FED9FA84" lastPageId="25" lastPageNumber="300" pageId="24" pageNumber="299" type="description">
<paragraph id="8BE07645FFFDFFBBFCA9DD73FC1BFA84" blockId="24.[854,1459,1119,2104]" pageId="24" pageNumber="299">
<emphasis id="B92BAA57FFFDFFBBFCA9DD73FC02FBEB" bold="true" box="[854,984,1119,1143]" pageId="24" pageNumber="299">Description</emphasis>
: Length: 11.0 13.0 mm. Colouration: head and pronotum black; elytra orange with blackish posterior margin; size of blackish spot slightly varying; abdomen orange with posterior abdominal segments dark brown or abdomen totally dark brown; legs light brown; antennae black.
</paragraph>
<paragraph id="8BE07645FFFDFFBBFCA9DC0CFC1FF94B" blockId="24.[854,1459,1119,2104]" pageId="24" pageNumber="299">
Head:
<quantity id="4CA7DBA0FFFDFFBBFC5BDC0CFBD8FAAB" box="[932,1026,1312,1336]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="24" pageNumber="299" unit="mm" value="1.29">1.29 mm</quantity>
long,
<quantity id="4CA7DBA0FFFDFFBBFBABDC0CFB68FAAB" box="[1108,1202,1312,1336]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="24" pageNumber="299" unit="mm" value="1.14">1.14 mm</quantity>
wide; eyes short; not prominent; temples more than twice as long as eyes; supraocular groove reaching from anterior edge of eyes to posterior edge of temples; preocular sides shortly parallel to base of antennae; front margin of clypeus slightly sinuate; central part prominent; punctation moderately deep and dense; size of puncture not varying; interstices between punctures between twice as wide as to four times as wide as diameter of punctures; at each end of supraocular groove with setiferous puncture; another pair of setiferous punctures at each side of prominent part of clypeus; elongate microsculpture dense, but not deep; on clypeus and vertex longitudinally striate; surface moderately matt.
</paragraph>
<paragraph id="8BE07645FFFDFFBBFCA9DFCCFB9DF82B" blockId="24.[854,1459,1119,2104]" pageId="24" pageNumber="299">Antennae as long as head and half of pronotum combined; first antennomere slightly thicker than following antennomeres; second antennomere short, approximately half as long as third; following antennomeres slightly decreasing in length; forth slightly longer than wide; tenth approximately quadrate; fourth to eleventh antennomere with long yellow setae.</paragraph>
<paragraph id="8BE07645FFFDFFBAFCA9DEECFEFAFDAB" blockId="24.[854,1459,1119,2104]" lastBlockId="25.[174,779,256,1304]" lastPageId="25" lastPageNumber="300" pageId="24" pageNumber="299">
Pronotum:
<quantity id="4CA7DBA0FFFDFFBBFC27DEECFBE2F84B" box="[984,1080,1984,2008]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="24" pageNumber="299" unit="mm" value="1.07">1.07 mm</quantity>
long,
<quantity id="4CA7DBA0FFFDFFBBFB78DEECFB3DF84B" box="[1159,1255,1984,2008]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="24" pageNumber="299" unit="mm" value="1.19">1.19 mm</quantity>
wide; in anterior half, sides parallel; in posterior half, sides smoothly narrowed in convex curve to posterior angles; posterior margin only half as wide as anterior margin; slightly prominent; in front of posterior margin, with pair of parallel short grooves bordering prominent central part; anterior angles at grooves shortly emarginate; anterior margin sinuate; punctation as dense and deep as on head; two setiferous punctures in anterior grooves; several setiferous punctures along lateral margin; two pairs of setiferous punctures at anterior margin close to anterior angles; elongate microsculpture as dense and deep as on head; longitudinally striate; surface moderately matt.
</paragraph>
<paragraph id="8BE07645FFFCFFBAFF51DB6CFE52FCAB" blockId="25.[174,779,256,1304]" pageId="25" pageNumber="300">
Elytra:
<quantity id="4CA7DBA0FFFCFFBAFF05DB6CFE83FDCB" box="[250,345,576,600]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="25" pageNumber="300" unit="mm" value="1.32">1.32 mm</quantity>
long,
<quantity id="4CA7DBA0FFFCFFBAFE65DB6CFE22FDCB" box="[410,504,576,600]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="25" pageNumber="300" unit="mm" value="1.26">1.26 mm</quantity>
wide; sides slightly divergent posteriad; humeral angles and posterior angles obtusely angled; punctation as dense and deep as on pronotum; several setiferous punctures along lateral margin; one setiferous puncture in central position on each side of suture; dense elongate microsculpture weaker as on head and pronotum; surface shiner than on head and pronotum.
</paragraph>
<paragraph id="8BE07645FFFCFFBAFF51DA6CFF29FC24" blockId="25.[174,779,256,1304]" pageId="25" pageNumber="300">Abdomen with weak punctation; few setiferous punctures in transverse line of each abdominal segment; microsculpture weak; surface still shinier than on elytra.</paragraph>
<paragraph id="8BE07645FFFCFFBAFF51DAECFED9FA84" blockId="25.[174,779,256,1304]" pageId="25" pageNumber="300">Aedeagus elongate; moderately thick; in lateral aspect, with broad base and shortly acute apex; ventral side in front of apex slightly concave; sclerotised ventral plate of central lobe with several sensillae; in ventral aspect, slightly widened in upper central part; in apical opening with oval lobe; along midline with indistinct groove margined by weak ridges; on each side of groove with numerous sensillae; parameres slender; nearly as long as central lobe; transparent widening at apex short and narrow; with few micro-sensillae and one setiferous sensilla.</paragraph>
</subSubSection>
<subSubSection id="C34525CEFFFCFFBAFF51DC6CFD79FAE4" pageId="25" pageNumber="300" type="etymology">
<paragraph id="8BE07645FFFCFFBAFF51DC6CFD79FAE4" blockId="25.[174,779,1344,1400]" pageId="25" pageNumber="300">
<emphasis id="B92BAA57FFFCFFBAFF51DC6CFEFFFAC4" bold="true" box="[174,293,1344,1368]" pageId="25" pageNumber="300">Etymology</emphasis>
: The species is named according to the countrys name, where the
<typeStatus id="54E4C8E7FFFCFFBAFE34DC4CFDF3FAE4" box="[459,553,1376,1400]" pageId="25" pageNumber="300" type="holotype">holotype</typeStatus>
was found.
</paragraph>
</subSubSection>
</treatment>
</document>