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<document ID-DOI="10.5733/afin.054.0213" ID-ISSN="2305-2562" ID-Zenodo-Dep="7918661" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583268648" checkinUser="felipe" docAuthor="Niekerk, P. van &amp; Dippenaar-Schoeman, A. S." docDate="2013" docId="03D8D254FFE3A92621BAFCE1FDCFFAE0" docLanguage="en" docName="AfrInvertebr.54.2.447-476.pdf" docOrigin="African Invertebrates 54 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.054.0213" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Heriaeus antoni Niekerk &amp; Dippenaar-Schoeman 2013, sp. n." docType="treatment" docVersion="3" lastPageNumber="456" masterDocId="FFE1AA2CFFEBA92F2055FF99FFE4FFE6" masterDocTitle="A revision of the crab spider genus Heriaeus Simon, 1875 (Araneae: Thomisidae) in the Afrotropical Region" masterLastPageNumber="447" masterPageNumber="447" pageNumber="455" updateTime="1683701492856" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>A revision of the crab spider genus Heriaeus Simon, 1875 (Araneae: Thomisidae) in the Afrotropical Region</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Niekerk, P. van</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dippenaar-Schoeman, A. S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-11-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>447</mods:start>
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<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.054.0213</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.054.0213</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7918661</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03D8D254FFE3A92621BAFCE1FDCFFAE0" httpUri="http://treatment.plazi.org/id/03D8D254FFE3A92621BAFCE1FDCFFAE0" lastPageId="9" lastPageNumber="456" pageId="8" pageNumber="455">
<subSubSection box="[495,752,888,914]" pageId="8" pageNumber="455" type="nomenclature">
<paragraph blockId="8.[495,752,888,956]" box="[495,752,888,914]" pageId="8" pageNumber="455">
<heading box="[495,752,888,914]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="455" reason="2">
<taxonomicName ID-CoL="6M45C" authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[495,685,888,914]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="455" phylum="Arthropoda" rank="species" species="antoni" status="sp. n.">
<emphasis bold="true" box="[495,685,888,914]" pageId="8" pageNumber="455">Heriaeus antoni</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[692,752,888,914]" pageId="8" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="456" pageId="8" pageNumber="455" type="description">
<paragraph blockId="8.[495,752,888,956]" box="[524,723,929,956]" pageId="8" pageNumber="455">
<figureCitation box="[524,595,929,955]" captionStart="Figs 19" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 19. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="8" pageNumber="455">Figs 3</figureCitation>
,
<figureCitation box="[608,683,929,955]" captionStart="Figs 1421" captionStartId="6.[144,183,1606,1628]" captionTargetBox="[163,1081,584,1580]" captionTargetId="figure-165@6.[97,1155,568,1587]" captionTargetPageId="6" captionText="Figs 1421. (1417) Heriaeus allenjonesi sp. n.: (14) male palp, ventral view; (15) male palp, lateral view; (16) epigyne,ventral view;(17) epigyne, dorsal view;(1821) Heriaeus antonisp.n.: (18) male palp, ventral view; (19) male palp, lateral view; (20) epigyne, ventral view; (21) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918667" httpUri="https://zenodo.org/record/7918667/files/figure.png" pageId="8" pageNumber="455">1821</figureCitation>
,
<figureCitation box="[695,723,929,956]" captionStart="Figs 5759" captionStartId="27.[347,386,1654,1676]" captionTargetBox="[181,1141,609,1633]" captionTargetId="figure-176@27.[132,1190,602,1636]" captionTargetPageId="27" captionText="Figs 5759. Distribution of Heriaeus spp. in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.7918679" httpUri="https://zenodo.org/record/7918679/files/figure.png" pageId="8" pageNumber="455">57</figureCitation>
</paragraph>
<paragraph blockId="8.[144,1104,972,1030]" pageId="8" pageNumber="455">
Etymology: Named for Antonius van Harten, the collector of the
<typeStatus box="[864,960,972,998]" pageId="8" pageNumber="455">paratype</typeStatus>
material, for his contribution in documenting the arachnid fauna of
<collectingCountry box="[752,834,1004,1030]" name="Yemen" pageId="8" pageNumber="455">Yemen</collectingCountry>
.
</paragraph>
<paragraph blockId="8.[144,1104,1047,1233]" pageId="8" pageNumber="455">
Diagnosis: The species is recognized by the numerous very short, blunt abdominal setae and the epigyne, which is very small with a small knob-like hood (
<figureCitation box="[953,1039,1079,1105]" captionStart="Figs 1421" captionStartId="6.[144,183,1606,1628]" captionTargetBox="[163,1081,584,1580]" captionTargetId="figure-165@6.[97,1155,568,1587]" captionTargetPageId="6" captionText="Figs 1421. (1417) Heriaeus allenjonesi sp. n.: (14) male palp, ventral view; (15) male palp, lateral view; (16) epigyne,ventral view;(17) epigyne, dorsal view;(1821) Heriaeus antonisp.n.: (18) male palp, ventral view; (19) male palp, lateral view; (20) epigyne, ventral view; (21) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918667" httpUri="https://zenodo.org/record/7918667/files/figure.png" pageId="8" pageNumber="455">Fig. 20</figureCitation>
). The male palp is distinct with its embolus having a U-shaped tip and a ventral lobe on the RTA (
<figureCitation box="[208,336,1143,1169]" captionStart="Figs 1421" captionStartId="6.[144,183,1606,1628]" captionTargetBox="[163,1081,584,1580]" captionTargetId="figure-165@6.[97,1155,568,1587]" captionTargetPageId="6" captionText="Figs 1421. (1417) Heriaeus allenjonesi sp. n.: (14) male palp, ventral view; (15) male palp, lateral view; (16) epigyne,ventral view;(17) epigyne, dorsal view;(1821) Heriaeus antonisp.n.: (18) male palp, ventral view; (19) male palp, lateral view; (20) epigyne, ventral view; (21) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918667" httpUri="https://zenodo.org/record/7918667/files/figure.png" pageId="8" pageNumber="455">Figs 18, 19</figureCitation>
). The embolus tip of the male palp resembles that of
<taxonomicName authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[920,1039,1143,1169]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="455" phylum="Arthropoda" rank="species" species="xanderi" status="sp. n.">
<emphasis box="[920,1039,1143,1169]" italics="true" pageId="8" pageNumber="455">H. xanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1045,1104,1143,1169]" pageId="8" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation box="[152,234,1174,1201]" captionStart="Figs 4552" captionStartId="23.[181,220,1202,1224]" captionTargetBox="[193,1135,210,1171]" captionTargetId="figure-202@23.[181,1141,204,1180]" captionTargetPageId="23" captionText="Figs 4552. (4548) Heriaeus transvaalicus Simon, 1875: (45) male palp, ventral view; (46) male palp, lateral view; (47) epigyne, ventral view; (48) epigyne, dorsal view; (4952) Heriaeus xanderi sp. n.: (49) male palp, ventral view; (50) male palp, lateral, view; (51) epigyne, ventral view; (52) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918675" httpUri="https://zenodo.org/record/7918675/files/figure.png" pageId="8" pageNumber="455">Fig. 50</figureCitation>
), but the much longer RTA of this species has a ventral lobe, which
<taxonomicName authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[985,1104,1175,1201]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="455" phylum="Arthropoda" rank="species" species="xanderi" status="sp. n.">
<emphasis box="[985,1104,1175,1201]" italics="true" pageId="8" pageNumber="455">H. xanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[144,204,1207,1233]" pageId="8" pageNumber="455" rank="species">sp. n.</taxonomicNameLabel>
lacks.
</paragraph>
<paragraph blockId="8.[144,1104,1249,1676]" box="[144,283,1249,1275]" pageId="8" pageNumber="455">Description:</paragraph>
<paragraph blockId="8.[144,1104,1249,1676]" box="[144,351,1289,1315]" pageId="8" pageNumber="455">
<emphasis box="[144,228,1289,1315]" italics="true" pageId="8" pageNumber="455">Female</emphasis>
(
<typeStatus box="[243,339,1289,1315]" pageId="8" pageNumber="455">allotype</typeStatus>
).
</paragraph>
<paragraph blockId="8.[144,1104,1249,1676]" lastBlockId="9.[181,1142,203,390]" lastPageId="9" lastPageNumber="456" pageId="8" pageNumber="455">
<emphasis box="[144,189,1330,1356]" italics="true" pageId="8" pageNumber="455">Size</emphasis>
: TL 4.80, CL 1.47, CW 1.52.
<emphasis box="[516,595,1329,1355]" italics="true" pageId="8" pageNumber="455">Colour</emphasis>
: Carapace with two broad longitudinal brown stripes; lateral edge with white line; setae brown; chelicerae yellow-brown; clypeus with white line on anterior edge; eye region white or with white markings between median eyes. Abdomen cream, slightly mottled with brown and white; specimen from
<collectingCountry name="Senegal" pageId="8" pageNumber="455">Senegal</collectingCountry>
differs in having six round marks arranged in two longitudinal lines centrally (
<figureCitation captionStart="Figs 19" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 19. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="8" pageNumber="455">Fig. 3</figureCitation>
) venter white, may be mottled with white or have a few brown spots laterally. Legs cream, dorsally with brown marks on femora II and III, tarsus I and metatarsi IIV; slight variations in
<collectingCountry box="[287,376,1553,1579]" name="Senegal" pageId="8" pageNumber="455">Senegal</collectingCountry>
specimens, markings absent in
<collectingCountry box="[725,803,1554,1580]" name="Yemen" pageId="8" pageNumber="455">Yemen</collectingCountry>
specimens.
<emphasis box="[938,1048,1554,1580]" italics="true" pageId="8" pageNumber="455">Carapace</emphasis>
:
<emphasis italics="true" pageId="8" pageNumber="455">Eye measurements</emphasis>
: AMEAME 0.17, ALEAME 0.11, PMEPME 0.18, PLEPME 0.20, AMEPME 0.16, MOQL AMEPME 0.23, MOQAW AMEAME 0.18, MOQPW PMEPME 0.23.
<emphasis box="[343,549,1650,1675]" italics="true" pageId="8" pageNumber="455">Leg measurements</emphasis>
: leg I Fe 1.42, Pat 0.76, Tib 1.17, Mt 1.07, Ta 0.52, total 4.94; leg II Fe 1.39, Pat 0.48, Tib 1.02, Mt 0.70, Ta 0.57, total 4.16; leg III Fe 0.84, Pat 0.42, Tib 0.60, Mt 0.52, Ta 0.43, total 2.81; leg IV Fe 1.18, Pat 0.32, Tib 0.60, Mt 0.57, Ta 0.41, total 3.08.
<emphasis box="[467,573,268,294]" italics="true" pageId="9" pageNumber="456">Abdomen</emphasis>
: Round, bearing numerous very short blunt setae; venter clothed with fine hairs.
<emphasis box="[525,617,300,325]" italics="true" pageId="9" pageNumber="456">Epigyne</emphasis>
: Very small with an unsclerotized small knob-like hood; two dark brown spots posteriorly indicating position of spermathecae (
<figureCitation captionStart="Figs 1421" captionStartId="6.[144,183,1606,1628]" captionTargetBox="[163,1081,584,1580]" captionTargetId="figure-165@6.[97,1155,568,1587]" captionTargetPageId="6" captionText="Figs 1421. (1417) Heriaeus allenjonesi sp. n.: (14) male palp, ventral view; (15) male palp, lateral view; (16) epigyne,ventral view;(17) epigyne, dorsal view;(1821) Heriaeus antonisp.n.: (18) male palp, ventral view; (19) male palp, lateral view; (20) epigyne, ventral view; (21) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918667" httpUri="https://zenodo.org/record/7918667/files/figure.png" pageId="9" pageNumber="456">Fig. 20</figureCitation>
); copulatory ducts two S-shaped coiled tubes (
<figureCitation box="[735,818,364,390]" captionStart="Figs 1421" captionStartId="6.[144,183,1606,1628]" captionTargetBox="[163,1081,584,1580]" captionTargetId="figure-165@6.[97,1155,568,1587]" captionTargetPageId="6" captionText="Figs 1421. (1417) Heriaeus allenjonesi sp. n.: (14) male palp, ventral view; (15) male palp, lateral view; (16) epigyne,ventral view;(17) epigyne, dorsal view;(1821) Heriaeus antonisp.n.: (18) male palp, ventral view; (19) male palp, lateral view; (20) epigyne, ventral view; (21) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918667" httpUri="https://zenodo.org/record/7918667/files/figure.png" pageId="9" pageNumber="456">Fig. 21</figureCitation>
).
</paragraph>
<paragraph blockId="9.[181,1149,406,1168]" box="[181,370,406,432]" pageId="9" pageNumber="456">
<emphasis box="[181,239,406,432]" italics="true" pageId="9" pageNumber="456">Male</emphasis>
(
<typeStatus box="[254,359,406,432]" pageId="9" pageNumber="456">holotype</typeStatus>
).
</paragraph>
<paragraph blockId="9.[181,1149,406,1168]" pageId="9" pageNumber="456">
<emphasis box="[181,226,447,473]" italics="true" pageId="9" pageNumber="456">Size</emphasis>
: TL 2.60, CL 1.23, CW 1.31.
<emphasis box="[566,645,446,472]" italics="true" pageId="9" pageNumber="456">Colour</emphasis>
: Carapace with two very broad longitudinal brown stripes extending from PEs and towards carapace sides, becoming dark brown at posterior end; setae brown; chelicerae yellow-brown; clypeus yellow-brown with brown marks; eye region yellow-brown; eye tubercles grey. Abdomen yellow-brown, lateral thirds dark brown to yellow-brown with brown and white markings; venter white; setae brown. Legs cream with brown marks on distal part of tibia and metatarsus I.
<emphasis box="[1024,1134,607,633]" italics="true" pageId="9" pageNumber="456">Carapace</emphasis>
:
<emphasis box="[181,391,639,664]" italics="true" pageId="9" pageNumber="456">Eye measurements</emphasis>
: AMEAME 0.12, ALEAME 0.8, PMEPME 0.15, PLEPME 0.19,AMEPME 0.13, MOQL AMEPME 0.20, MOQAW AMEAME 0.18, MOQPW PMEPME 0.23.
<emphasis box="[380,587,703,728]" italics="true" pageId="9" pageNumber="456">Leg measurements</emphasis>
: leg I Fe 1.54, Pat 0.71, Tib 1.45, Mt 1.33, Ta 0.71, total 5.74; leg II Fe 1.52, Pat 0.54, Tib 1.10, Mt 1.15, Ta 0.68, total 4.99; leg III Fe 0.91, Pat 0.35, Tib 0.62, Mt 0.59, Ta 0.38, total 2.85; leg IV Fe 1.05, Pat 0.31, Tib 0.72, Mt 0.66, Ta 0.40, total 3.14.
<emphasis box="[461,567,798,824]" italics="true" pageId="9" pageNumber="456">Abdomen</emphasis>
: Oval, truncated anteriorly; bearing numerous very short, blunt setae; venter with fine hairs.
<emphasis box="[621,732,830,856]" italics="true" pageId="9" pageNumber="456">Male palp</emphasis>
: Embolus long, winding once around tegulum (
<figureCitation box="[287,368,862,888]" captionStart="Figs 1421" captionStartId="6.[144,183,1606,1628]" captionTargetBox="[163,1081,584,1580]" captionTargetId="figure-165@6.[97,1155,568,1587]" captionTargetPageId="6" captionText="Figs 1421. (1417) Heriaeus allenjonesi sp. n.: (14) male palp, ventral view; (15) male palp, lateral view; (16) epigyne,ventral view;(17) epigyne, dorsal view;(1821) Heriaeus antonisp.n.: (18) male palp, ventral view; (19) male palp, lateral view; (20) epigyne, ventral view; (21) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918667" httpUri="https://zenodo.org/record/7918667/files/figure.png" pageId="9" pageNumber="456">Fig. 18</figureCitation>
); embolus tip slender with U-shaped curve; VTA small and two-lobed, RTA with ventral lobe and long distal tooth (
<figureCitation box="[681,764,894,920]" captionStart="Figs 1421" captionStartId="6.[144,183,1606,1628]" captionTargetBox="[163,1081,584,1580]" captionTargetId="figure-165@6.[97,1155,568,1587]" captionTargetPageId="6" captionText="Figs 1421. (1417) Heriaeus allenjonesi sp. n.: (14) male palp, ventral view; (15) male palp, lateral view; (16) epigyne,ventral view;(17) epigyne, dorsal view;(1821) Heriaeus antonisp.n.: (18) male palp, ventral view; (19) male palp, lateral view; (20) epigyne, ventral view; (21) epigyne, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7918667" httpUri="https://zenodo.org/record/7918667/files/figure.png" pageId="9" pageNumber="456">Fig. 19</figureCitation>
).
</paragraph>
<paragraph blockId="9.[181,1149,406,1168]" pageId="9" pageNumber="456">
<materialsCitation collectingDate="1989-08" collectorName="J. Everts" country="Senegal" latitude="16.33" location="Richard Toll" longLatPrecision="769" longitude="15.5" pageId="9" pageNumber="456" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[181,269,933,955]" pageId="9" pageNumber="456">Holotype</typeStatus>
:
<specimenCount box="[280,297,933,954]" pageId="9" pageNumber="456" type="male"></specimenCount>
<collectingCountry box="[304,411,933,954]" name="Senegal" pageId="9" pageNumber="456">SENEGAL</collectingCountry>
:
<quantity box="[421,505,932,955]" metricMagnitude="3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="9" pageNumber="456" unit="km" value="7.5" valueMax="10.0" valueMin="5.0">510 km</quantity>
S of
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE3A92621BAFCE1FDCFFAE0:8EAE3599FFE2A926227AFC3CFD46FC5D" box="[559,674,933,955]" country="Senegal" latitude="16.33" longLatPrecision="769" longitude="15.5" name="Richard Toll" pageId="9" pageNumber="456">Richard Toll</location>
(
<geoCoordinate box="[688,766,932,954]" degrees="16.33" direction="north" orientation="latitude" pageId="9" pageNumber="456" precision="555" value="16.33">16.33°N</geoCoordinate>
<geoCoordinate box="[773,850,932,954]" degrees="15.50" direction="east" orientation="longitude" pageId="9" pageNumber="456" precision="555" value="15.5">15.50°E</geoCoordinate>
),
<date box="[868,952,933,955]" pageId="9" pageNumber="456" value="1989-08">
<collectingDate box="[868,952,933,955]" pageId="9" pageNumber="456" value="1989-08">viii.1989</collectingDate>
</date>
,
<collectorName box="[963,1045,933,954]" pageId="9" pageNumber="456">J. Everts</collectorName>
, semi­arid thorn bush (MRAC 172.021a).
</materialsCitation>
</paragraph>
<paragraph blockId="9.[181,1149,406,1168]" box="[181,691,991,1013]" pageId="9" pageNumber="456">
<materialsCitation box="[181,691,991,1013]" collectingDate="1989-08" collectorName="J. Everts" country="Senegal" latitude="16.33" location="Richard Toll" longLatPrecision="769" longitude="15.5" pageId="9" pageNumber="456" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
<typeStatus box="[181,264,991,1013]" pageId="9" pageNumber="456">Allotype</typeStatus>
:
<specimenCount box="[273,301,991,1012]" pageId="9" pageNumber="456" type="female">1♀</specimenCount>
same data as holotype (MRAC 172.021b).
</materialsCitation>
</paragraph>
<paragraph blockId="9.[181,1149,406,1168]" pageId="9" pageNumber="456">
<materialsCitation box="[181,923,1026,1048]" collectingDate="1989-08" collectionCode="MRAC" collectorName="J. Everts" country="Senegal" latitude="16.33" location="Richard Toll" longLatPrecision="769" longitude="15.5" pageId="9" pageNumber="456" specimenCode="MRAC 172.021" specimenCount="23" specimenCount-female="6" specimenCount-male="3" typeStatus="paratype">
<typeStatus box="[181,273,1026,1047]" pageId="9" pageNumber="456">Paratypes</typeStatus>
:
<collectingCountry box="[282,389,1026,1047]" name="Senegal" pageId="9" pageNumber="456">SENEGAL</collectingCountry>
:
<specimenCount box="[398,426,1026,1047]" pageId="9" pageNumber="456" type="male">3♂</specimenCount>
<specimenCount box="[433,461,1026,1048]" pageId="9" pageNumber="456" type="female">6♀</specimenCount>
<specimenCount box="[467,545,1026,1048]" pageId="9" pageNumber="456" type="generic">14 imm.</specimenCount>
same data as holotype (
<specimenCode box="[763,915,1026,1047]" collectionCode="MRAC" country="Belgium" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="456" type="Museum">MRAC 172.021</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1989-09" collectionCode="MRAC" collectorName="J. Everts" country="Senegal" latitude="16.33" location="Richard Toll" longLatPrecision="769" longitude="15.5" pageId="9" pageNumber="456" specimenCount="25" specimenCount-female="12" specimenCount-male="12" typeStatus="paratype">
<specimenCount box="[932,972,1026,1047]" pageId="9" pageNumber="456" type="male">12♂</specimenCount>
<specimenCount box="[978,1017,1026,1047]" pageId="9" pageNumber="456" type="female">12♀</specimenCount>
<specimenCount box="[1024,1090,1026,1048]" pageId="9" pageNumber="456" type="generic">1 imm.</specimenCount>
same data but
<date box="[258,326,1050,1072]" pageId="9" pageNumber="456" value="1989-09">
<collectingDate box="[258,326,1050,1072]" pageId="9" pageNumber="456" value="1989-09">ix.1989</collectingDate>
</date>
(MRAC 171.999, 172.005)
</materialsCitation>
;
<materialsCitation box="[587,1026,1050,1072]" collectingDate="1989-08" collectingDateMax="1989-11-31" collectingDateMin="1989-08" collectionCode="MRAC" collectorName="J. Everts" country="Senegal" latitude="16.33" location="Richard Toll" longLatPrecision="769" longitude="15.5" pageId="9" pageNumber="456" specimenCode="MRAC 174.239" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[587,615,1050,1071]" pageId="9" pageNumber="456" type="male">1♂</specimenCount>
same data but
<date box="[748,858,1050,1072]" pageId="9" pageNumber="456" value="1989-08" valueMax="1989-11-31" valueMin="1989-08-00">
<collectingDate box="[748,858,1050,1072]" pageId="9" pageNumber="456" value="1989-08" valueMax="1989-11-31" valueMin="1989-08-00">viiixi.1989</collectingDate>
</date>
(
<specimenCode box="[869,1020,1050,1071]" collectionCode="MRAC" country="Belgium" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="456" type="Museum">MRAC 174.239</specimenCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1991-09-18" collectionCode="MRAC" collectorName="H. van der Valk" country="Senegal" latitude="16.33" location="Richard Toll" longLatPrecision="769" longitude="15.5" pageId="9" pageNumber="456" specimenCode="MRAC 200.583" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount box="[1034,1063,1050,1071]" pageId="9" pageNumber="456" type="female">1♀</specimenCount>
<quantity box="[1068,1124,1050,1072]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="456" unit="km" value="20.0">20 km</quantity>
S of
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE3A92621BAFCE1FDCFFAE0:8EAE3599FFE2A926209BFBABFEA4FBAE" box="[206,320,1074,1096]" country="Senegal" latitude="16.33" longLatPrecision="769" longitude="15.5" name="Richard Toll" pageId="9" pageNumber="456">Richard Toll</location>
(
<geoCoordinate box="[332,410,1074,1096]" degrees="16.33" direction="north" orientation="latitude" pageId="9" pageNumber="456" precision="555" value="16.33">16.33°N</geoCoordinate>
<geoCoordinate box="[416,493,1074,1096]" degrees="15.50" direction="east" orientation="longitude" pageId="9" pageNumber="456" precision="555" value="15.5">15.50°E</geoCoordinate>
),
<date box="[509,607,1074,1096]" pageId="9" pageNumber="456" value="1991-09-18">
<collectingDate box="[509,607,1074,1096]" pageId="9" pageNumber="456" value="1991-09-18">18.ix.1991</collectingDate>
</date>
,
<collectorName box="[618,763,1074,1096]" pageId="9" pageNumber="456">H. van der Valk</collectorName>
,
<collectingMethod box="[772,876,1074,1096]" pageId="9" pageNumber="456">pitfall traps</collectingMethod>
(
<specimenCode box="[888,1041,1074,1096]" collectionCode="MRAC" country="Belgium" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="456" type="Museum">MRAC 200.583</specimenCode>
)
</materialsCitation>
.
<materialsCitation collectingDate="1998-05-15" collectionCode="NCA" collectorName="A. van Harten" country="Yemen" latitude="15.35" location="Sana'a" longLatPrecision="771" longitude="44.2" pageId="9" pageNumber="456" specimenCode="NCA 2012" specimenCount="1" specimenCount-female="1" stateProvince="San'a'" typeStatus="paratype">
<collectingCountry box="[1055,1139,1074,1095]" name="Yemen" pageId="9" pageNumber="456">YEMEN</collectingCountry>
:
<specimenCount box="[181,210,1098,1119]" pageId="9" pageNumber="456" type="female">1♀</specimenCount>
near
<collectingRegion box="[257,319,1098,1119]" country="Yemen" name="San'a'" pageId="9" pageNumber="456">Sanaa</collectingRegion>
(
<geoCoordinate box="[329,407,1098,1120]" degrees="15.35" direction="north" orientation="latitude" pageId="9" pageNumber="456" precision="555" value="15.35">15.35°N</geoCoordinate>
<geoCoordinate box="[411,488,1098,1119]" degrees="44.20" direction="east" orientation="longitude" pageId="9" pageNumber="456" precision="555" value="44.2">44.20°E</geoCoordinate>
),
<date box="[502,593,1098,1120]" pageId="9" pageNumber="456" value="1998-05-15">
<collectingDate box="[502,593,1098,1120]" pageId="9" pageNumber="456" value="1998-05-15">15.v.1998</collectingDate>
</date>
,
<collectorName box="[600,725,1098,1119]" pageId="9" pageNumber="456">A. van Harten</collectorName>
(
<specimenCode box="[734,834,1098,1120]" collectionCode="NCA" pageId="9" pageNumber="456">NCA 2012</specimenCode>
/2056)
</materialsCitation>
;
<materialsCitation collectingDate="1998-07" collectionCode="NCA" collectorName="A. van Harten" country="Yemen" latitude="15.35" location="Sana'a" longLatPrecision="771" longitude="44.2" pageId="9" pageNumber="456" specimenCode="NCA 2012" specimenCount="1" specimenCount-female="1" stateProvince="San'a'" typeStatus="paratype">
<specimenCount box="[902,930,1098,1119]" pageId="9" pageNumber="456" type="female">1♀</specimenCount>
same locality,
<date box="[1061,1138,1098,1120]" pageId="9" pageNumber="456" value="1998-07">
<collectingDate box="[1061,1138,1098,1120]" pageId="9" pageNumber="456" value="1998-07">vii.1998</collectingDate>
</date>
,
<collectorName box="[181,310,1122,1143]" pageId="9" pageNumber="456">A. van Harten</collectorName>
(
<specimenCode box="[321,423,1122,1144]" collectionCode="NCA" pageId="9" pageNumber="456">NCA 2012</specimenCode>
/2058)
</materialsCitation>
;
<materialsCitation collectingDate="1998-04-20" collectionCode="NCA" collectorName="A. van Harten" country="Yemen" latitude="16.0" location="Abs" longLatPrecision="770" longitude="43.2" pageId="9" pageNumber="456" specimenCode="NCA 2012" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount box="[492,521,1122,1143]" pageId="9" pageNumber="456" type="female">2♀</specimenCount>
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE3A92621BAFCE1FDCFFAE0:8EAE3599FFE2A9262258FBFBFDD6FB9E" box="[525,562,1122,1144]" country="Yemen" latitude="16.0" longLatPrecision="770" longitude="43.2" name="Abs" pageId="9" pageNumber="456">Abs</location>
(
<geoCoordinate box="[574,651,1122,1144]" degrees="16.00" direction="north" orientation="latitude" pageId="9" pageNumber="456" precision="555" value="16.0">16.00°N</geoCoordinate>
<geoCoordinate box="[658,735,1122,1143]" degrees="43.20" direction="east" orientation="longitude" pageId="9" pageNumber="456" precision="555" value="43.2">43.20°E</geoCoordinate>
),
<date box="[751,848,1122,1144]" pageId="9" pageNumber="456" value="1998-04-20">
<collectingDate box="[751,848,1122,1144]" pageId="9" pageNumber="456" value="1998-04-20">20.iv.1998</collectingDate>
</date>
,
<collectorName box="[856,987,1122,1143]" pageId="9" pageNumber="456">A. van Harten</collectorName>
, in garden (
<specimenCode collectionCode="NCA" pageId="9" pageNumber="456">NCA 2012</specimenCode>
/2057)
</materialsCitation>
;
<materialsCitation box="[298,1065,1146,1168]" collectingDate="1999-12-17" collectionCode="NCA" collectorName="A. van Harten" country="Yemen" latitude="13.4" location="Wadi Warazan" longLatPrecision="774" longitude="44.23" pageId="9" pageNumber="456" specimenCode="NCA 2012" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount box="[298,326,1146,1167]" pageId="9" pageNumber="456" type="female">1♀</specimenCount>
<location LSID="urn:lsid:plazi:treatment:03D8D254FFE3A92621BAFCE1FDCFFAE0:8EAE3599FFE2A9262119FBE3FE2BFB69" box="[332,463,1146,1168]" country="Yemen" latitude="13.4" longLatPrecision="774" longitude="44.23" name="Wadi Warazan" pageId="9" pageNumber="456">Wadi Warazan</location>
(
<geoCoordinate box="[475,552,1146,1167]" degrees="13.40" direction="north" orientation="latitude" pageId="9" pageNumber="456" precision="555" value="13.4">13.40°N</geoCoordinate>
<geoCoordinate box="[558,635,1146,1167]" degrees="44.23" direction="east" orientation="longitude" pageId="9" pageNumber="456" precision="555" value="44.23">44.23°E</geoCoordinate>
),
<date box="[651,756,1146,1168]" pageId="9" pageNumber="456" value="1999-12-17">
<collectingDate box="[651,756,1146,1168]" pageId="9" pageNumber="456" value="1999-12-17">17.xii.1999</collectingDate>
</date>
,
<collectorName box="[764,892,1146,1167]" pageId="9" pageNumber="456">A. van Harten</collectorName>
(
<specimenCode box="[904,1005,1146,1168]" collectionCode="NCA" pageId="9" pageNumber="456">NCA 2012</specimenCode>
/2059)
</materialsCitation>
.
</paragraph>
<paragraph blockId="9.[181,664,1185,1212]" box="[181,664,1185,1212]" pageId="9" pageNumber="456">
Distribution:
<collectingCountry box="[332,421,1185,1211]" name="Senegal" pageId="9" pageNumber="456">Senegal</collectingCountry>
and
<collectingCountry box="[474,552,1185,1211]" name="Yemen" pageId="9" pageNumber="456">Yemen</collectingCountry>
(
<figureCitation box="[567,651,1185,1212]" captionStart="Figs 5759" captionStartId="27.[347,386,1654,1676]" captionTargetBox="[181,1141,609,1633]" captionTargetId="figure-176@27.[132,1190,602,1636]" captionTargetPageId="27" captionText="Figs 5759. Distribution of Heriaeus spp. in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.7918679" httpUri="https://zenodo.org/record/7918679/files/figure.png" pageId="9" pageNumber="456">Fig. 57</figureCitation>
).
</paragraph>
<paragraph blockId="9.[181,1141,1228,1286]" pageId="9" pageNumber="456">Natural history: Females collected during April, May, June and December and adult males from August to November.</paragraph>
</subSubSection>
</treatment>
</document>