312 lines
33 KiB
XML
312 lines
33 KiB
XML
<document id="235FE1A42C05946E4DC569CDDBE6E8BB" ID-DOI="10.11646/zootaxa.3904.1.10" ID-GBIF-Dataset="4cb136fd-d20e-42c2-a6f9-5abf5246b422" ID-ISSN="1175-5326" ID-Zenodo-Dep="244415" ID-ZooBank="60435E31-9E0F-449F-9DE2-09C32EDFA583" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460885254563" checkinUser="plazi" docAuthor="Antić, Dragan Ž., Tomić, Vladimir T., Rađa, Tonći, Lučić, Luka R., Dudić, Boris D. & Makarov, Slobodan E." docDate="2015" docId="03D8917DA7096C24FF12FA653BFAFF34" docLanguage="en" docName="zt03904p150.pdf" docOrigin="Zootaxa 3904 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heterolatzelia karlstrasseri Antic and Makarov, new species" docType="treatment" docVersion="8" lastPageNumber="149" masterDocId="FFE1E905A7096C27FF85FFB53F66FFE5" masterDocTitle="A new species of the family Heterolatzeliidae from the Balkan Peninsula (Diplopoda, Chordeumatida)" masterLastPageNumber="150" masterPageNumber="147" pageNumber="147" updateTime="1698358935506" updateUser="plazi">
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<mods:title id="0424FD15730C957C7DEE3A24A6B93A83">A new species of the family Heterolatzeliidae from the Balkan Peninsula (Diplopoda, Chordeumatida)</mods:title>
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</mods:titleInfo>
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<mods:name id="0F42C23C5DD4CB9FCCBE282C3E306CCD" type="personal">
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<mods:roleTerm id="F0C8EF0F55092E0CE5A170994D3410D3">Author</mods:roleTerm>
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<mods:namePart id="362D40FC7209D99CAE1A18C4CDAAADCD">Antić, Dragan Ž.</mods:namePart>
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</mods:name>
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<mods:roleTerm id="C0231DF06CB62654D3A7E2C8BEA98D17">Author</mods:roleTerm>
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<mods:namePart id="182769735A3C9227939736561A0D478E">Tomić, Vladimir T.</mods:namePart>
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</mods:name>
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<mods:name id="537A892ECB2D0DA202D6BBA52D4FCF0D" type="personal">
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<mods:role id="B348E37921FC0269084F8A62839D93D0">
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<mods:roleTerm id="AB4DF5E04402F95298EF61C1DF98A874">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8A7246801E26EEF8A04F580AEE720057">Rađa, Tonći</mods:namePart>
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</mods:name>
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<mods:name id="05132F09C79B6A0C900D5E1CF39E44F8" type="personal">
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<mods:roleTerm id="B257EFED494F961407FD39D1669C053F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="16D59B6AAFBAE7EE1EA9F38DFABCBA97">Lučić, Luka R.</mods:namePart>
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</mods:name>
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<mods:name id="282D45439BFD216E071C58B0F9441D20" type="personal">
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<mods:roleTerm id="1339D964F76762A47510EB18140864BB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F078B829C1ECA0AEF728ADA04C45E07D">Dudić, Boris D.</mods:namePart>
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</mods:name>
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<mods:name id="BD4DF488BB38A5326C4F501B9057DD6F" type="personal">
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<mods:roleTerm id="519439FF44C11A8F3F7D80DF8BD1ABBE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="78DEB3DC173AD5664B0289AABB0F9ADB">Makarov, Slobodan E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="CC5A6B6FEC686AE4CF9A5E49AA67870F">text</mods:typeOfResource>
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<mods:title id="C9DFC836DD31D08B63BD4A4733E72106">Zootaxa</mods:title>
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<mods:part id="73D7762ACDA31832D56B33287DE04090">
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<mods:date id="7F3AAAE2ECF6F3A7DA61ADC7B7B52AA8">2015</mods:date>
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<mods:detail id="42BE0FED183D7C274C4441C422F4A055" type="volume">
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<mods:number id="C4A24DF394FF69A277C582DA5F95A1BA">3904</mods:number>
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<mods:classification id="6A158E5919695AA9099C0D75E75FBDA5">journal article</mods:classification>
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<mods:identifier id="943E2ABCF91BA37275055B13F00D481D" type="DOI">10.11646/zootaxa.3904.1.10</mods:identifier>
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<mods:identifier id="1D6BFD5BCF1ECC757A92FDD2945DC6F6" type="GBIF-Dataset">4cb136fd-d20e-42c2-a6f9-5abf5246b422</mods:identifier>
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<mods:identifier id="CCD8D0E3FDF9F0B7DB7A90ABB7F5AFA6" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="DCE49BD204553E529FAFECA687C672BC" type="Zenodo-Dep">244415</mods:identifier>
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<treatment id="03D8917DA7096C24FF12FA653BFAFF34" ID-DOI="http://doi.org/10.5281/zenodo.6112712" ID-GBIF-Taxon="119586291" ID-Zenodo-Dep="6112712" LSID="urn:lsid:plazi:treatment:03D8917DA7096C24FF12FA653BFAFF34" httpUri="http://treatment.plazi.org/id/03D8917DA7096C24FF12FA653BFAFF34" lastPageId="3" lastPageNumber="149" pageId="0" pageNumber="147">
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<subSubSection id="C36B73E0A7096C27FF12FA653F9CF9EF" pageId="0" pageNumber="147" type="nomenclature">
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<paragraph id="8BCE206BA7096C27FF12FA653C11FA0F" blockId="0.[151,887,1488,1546]" box="[151,887,1488,1514]" pageId="0" pageNumber="147">
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<heading id="D0869707A7096C27FF12FA653C11FA0F" bold="true" box="[151,887,1488,1514]" fontSize="11" level="1" pageId="0" pageNumber="147" reason="1">
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<emphasis id="B905FC79A7096C27FF12FA653C11FA0F" bold="true" box="[151,887,1488,1514]" pageId="0" pageNumber="147">
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<taxonomicName id="4C715BE8A7096C27FF12FA653DBCFA0F" ID-CoL="3L9YC" authority="Antic and Makarov" authorityName="Antic and Makarov" box="[151,730,1488,1514]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="karlstrasseri" status="sp. nov.">
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<emphasis id="B905FC79A7096C27FF12FA653EBBFA0F" bold="true" box="[151,477,1488,1514]" italics="true" pageId="0" pageNumber="147">Heterolatzelia karlstrasseri</emphasis>
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Antić and Makarov
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2364102A7096C27FD62FA653C11FA0F" box="[743,887,1488,1514]" pageId="0" pageNumber="147" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCE206BA7096C27FF12FA463F9CF9EF" blockId="0.[151,887,1488,1546]" box="[151,250,1523,1546]" pageId="0" pageNumber="147">
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<figureCitation id="134A3CEEA7096C27FF12FA463F9CF9EF" box="[151,250,1523,1546]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,269,1875,1898]" captionTargetBox="[152,1411,703,1874]" captionTargetId="figure@1.[151,1436,678,1874]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Heterolatzelia karlstrasseri sp. n., holotype male gonopods. 1. anterior gonopods, oral view; 2. anterior gonopods, caudal view; 3. anterior gonopods, lateral view; 4. posterior gonopods, oral view. Designations: a: angiocoxite; b: comb-like process; c: colpocoxite; d: coxal cavity; e: wide subquadrangular part; f: teeth of comb-like process; l: connecting lamella between colpocoxites; t: rudimentary telopodites; cb: caudal bulges of angiocoxite. Scale line: 0.4 mm." httpUri="https://zenodo.org/record/244416/files/figure.png" pageId="0" pageNumber="147">Figs. 1–4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B73E0A7096C27FF12F98D3A3EF952" pageId="0" pageNumber="147" type="materials_examined">
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<paragraph id="8BCE206BA7096C27FF12F98D3A3EF952" blockId="0.[151,1436,1592,2031]" pageId="0" pageNumber="147">
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<emphasis id="B905FC79A7096C27FF12F98D3E1EF9AA" bold="true" box="[151,376,1592,1615]" pageId="0" pageNumber="147">Material examined.</emphasis>
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<typeStatus id="54CA9EC9A7096C27FE04F98D3E8EF9AA" box="[385,488,1592,1615]" pageId="0" pageNumber="147" type="holotype">Holotype</typeStatus>
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: male from the Đatlo Cave (
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<geoCoordinate id="EE4546ACA7096C27FCA3F98D3CAEF9AA" box="[806,968,1592,1615]" direction="north" orientation="latitude" pageId="0" pageNumber="147" precision="1" value="43.06167">43°03'42.01"N</geoCoordinate>
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,
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<geoCoordinate id="EE4546ACA7096C27FC53F98D3B13F9AA" box="[982,1141,1592,1615]" direction="east" orientation="longitude" pageId="0" pageNumber="147" precision="1" value="18.494686">18°29'40.87"E</geoCoordinate>
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), Kobilja Glava, village of Korita, boundary of the municipalities of Bileća and Gacko,
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<collectingCountry id="F36660FBA7096C27FCADF9EE3B55F997" box="[808,1075,1627,1650]" name="Bosnia and Herzegovina" pageId="0" pageNumber="147">Bosnia and Herzegovina</collectingCountry>
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, collected on
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<date id="FFCF06ABA7096C27FB4CF9EE3A3DF997" box="[1225,1371,1627,1650]" pageId="0" pageNumber="147" value="2013-07-14">July 14, 2013</date>
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by T. Rađa (IZB, HTL
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<collectingCountry id="F36660FBA7096C27FED7F9C83E1EF971" box="[338,376,1661,1684]" name="Hong Kong" pageId="0" pageNumber="147">HK</collectingCountry>
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100-1).
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<typeStatus id="54CA9EC9A7096C27FE51F9C83D26F971" box="[468,576,1661,1684]" pageId="0" pageNumber="147" type="paratype">Paratypes</typeStatus>
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: four juveniles, same data as
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<typeStatus id="54CA9EC9A7096C27FCFEF9C83CBEF971" box="[891,984,1661,1684]" pageId="0" pageNumber="147" type="holotype">holotype</typeStatus>
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(IZB, HTL
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<collectingCountry id="F36660FBA7096C27FBD9F9C83BE4F971" box="[1116,1154,1661,1684]" name="Hong Kong" pageId="0" pageNumber="147">HK</collectingCountry>
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100-2). The
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<typeStatus id="54CA9EC9A7096C27FA8AF9C83A5BF971" box="[1295,1341,1661,1684]" pageId="0" pageNumber="147">type</typeStatus>
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material is deposited in the collection of the Institute of Zoology, Faculty of Biology, University of Belgrade,
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<collectingCountry id="F36660FBA7096C27FB4FF9153A68F952" box="[1226,1294,1696,1719]" name="Serbia" pageId="0" pageNumber="147">Serbia</collectingCountry>
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(IZB).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B73E0A7096C27FF42F9763C4CF919" pageId="0" pageNumber="147" type="etymology">
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<paragraph id="8BCE206BA7096C27FF42F9763C4CF919" blockId="0.[151,1436,1592,2031]" pageId="0" pageNumber="147">
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<emphasis id="B905FC79A7096C27FF42F9763E26F93F" bold="true" box="[199,320,1731,1754]" pageId="0" pageNumber="147">Etymology</emphasis>
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. In honor of the late Dr. Karl Strasser, one of the greatest European diplopodologists, whose contribution to our knowledge of the Balkan millipede fauna is invaluable.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B73E0A7096C24FF42F8BD3BFAFF34" lastPageId="3" lastPageNumber="150" pageId="0" pageNumber="147" type="description">
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<paragraph id="8BCE206BA7096C27FF42F8BD3FBDF862" blockId="0.[151,1436,1592,2031]" pageId="0" pageNumber="147">
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<emphasis id="B905FC79A7096C27FF42F8BD3E2DF8FA" bold="true" box="[199,331,1800,1823]" pageId="0" pageNumber="147">Description</emphasis>
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. Body with 30 segments (including telson) in
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<typeStatus id="54CA9EC9A7096C27FCCBF8BD3CCDF8FA" box="[846,939,1800,1823]" pageId="0" pageNumber="147" type="holotype">holotype</typeStatus>
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male. Juveniles with 28 segments (including telson). Dorsal and ventrolateral side of metazonites brownish, prozonites, lateral keels, and anterior part of metazonites yellowish. Body length
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<quantity id="4C898D8EA7096C27FE1BF8F83D6BF881" box="[414,525,1869,1892]" metricMagnitude="-2" metricUnit="m" metricValue="1.264" pageId="0" pageNumber="147" unit="mm" value="12.64">12.64 mm</quantity>
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(
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<typeStatus id="54CA9EC9A7096C27FD98F8F83D1AF881" box="[541,636,1869,1892]" pageId="0" pageNumber="147" type="holotype">holotype</typeStatus>
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male). Vertical diameter of the largest pleurotergites
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<quantity id="4C898D8EA7096C27FB4FF8F83A4DF881" box="[1226,1323,1869,1892]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="0" pageNumber="147" unit="mm" value="0.92">0.92 mm</quantity>
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(
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<typeStatus id="54CA9EC9A7096C27FAB9F8F83AFDF881" box="[1340,1435,1869,1892]" pageId="0" pageNumber="147" type="holotype">holotype</typeStatus>
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male).
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</paragraph>
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<paragraph id="8BCE206BA7096C26FF42F8263CC5FEF2" blockId="0.[151,1436,1592,2031]" lastBlockId="1.[151,1436,152,660]" lastPageId="1" lastPageNumber="148" pageId="0" pageNumber="147">
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<emphasis id="B905FC79A7096C27FF42F8263E67F84F" box="[199,257,1939,1962]" italics="true" pageId="0" pageNumber="147">Head</emphasis>
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(
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<typeStatus id="54CA9EC9A7096C27FE8BF8263E0BF84F" box="[270,365,1939,1962]" pageId="0" pageNumber="147" type="holotype">holotype</typeStatus>
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male): With five rows of 22 ocellae. Occipital axial suture short and clear. Labrum with three labral teeth, and with 5+6 labral and 2+2 supralabral setae. Ventral margins of stipes densely denticulated. Each stipe with one prominent anterior tooth with two setae. Gnathochilarium without any peculiarities. Promentum triangular; stipites with 4+4 long marginal setae, 1+1 long apical setae, and 7+7 basal microsetae; lingual plates with two rows of 5+7 setae (2+2 inner setae, and 3+5 outer setae). Antennal length
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<quantity id="4C898D8EA7086C26FD3BFF0F3C73FF34" box="[702,789,186,209]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="148" unit="mm" value="2.1">2.1 mm</quantity>
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. In length: antennomeres I <VII <VI <II ~ IV <V ~ III. Length/breadth ratios of antennomeres I–VII: 1 (I), 2.27 (II), 8.14 (III), 3.57 (IV), 4.90 (V), 2.09 (VI), and 1.88 (VII), respectively. Antennomeres IV, V and VI with a few long sensitive setae.
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</paragraph>
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<paragraph id="8BCE206BA7086C26FF42FE973D9CFEDC" blockId="1.[151,1436,152,660]" box="[199,762,290,313]" pageId="1" pageNumber="148">
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<emphasis id="B905FC79A7086C26FF42FE973E73FEDC" box="[199,277,290,313]" italics="true" pageId="1" pageNumber="148">Collum</emphasis>
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: Narrower than head, with six macrochaetae.
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</paragraph>
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<paragraph id="8BCE206BA7086C26FF42FEF03D16FDAA" blockId="1.[151,1436,152,660]" pageId="1" pageNumber="148">
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<emphasis id="B905FC79A7086C26FF42FEF03E3BFEB9" box="[199,349,325,348]" italics="true" pageId="1" pageNumber="148">Pleurotergites</emphasis>
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: Lateral keels well developed, with two rounded lobes, becoming larger from pleurotergite V, poorly developed on pleurotergites XXVI and XXVII, disappearing on pleurotergites XXVIII and XXIX. Macrochaetae short, trichoid. Macrochaetal index CIX (pleurotergite 15),
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<emphasis id="B905FC79A7086C26FD5DFE3F3D9FFE44" box="[728,761,394,417]" italics="true" pageId="1" pageNumber="148">i.e.</emphasis>
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, distance between exterior and median macrochaetae/distance between interior and median macrochaetae = 0.58. Median index MIX (pleurotergite 15)
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<emphasis id="B905FC79A7086C26FBDBFE183B1EFE21" box="[1118,1144,429,452]" italics="true" pageId="1" pageNumber="148">i.e</emphasis>
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., distance between interior macrochaetae and axial suture/distance between interior and median macrochaetae = 0.74. Paratergal index PIX (pleurotergite 15),
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<emphasis id="B905FC79A7086C26FEE7FE473EEFFDEC" box="[354,393,498,521]" italics="true" pageId="1" pageNumber="148">i.e.,</emphasis>
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(width of metazonite - width of prozonite)/(
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<date id="FFCF06ABA7086C26FCE9FE473CEBFDEC" box="[876,909,498,521]" pageId="1" pageNumber="148">2 x</date>
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length of paratergum) = 0.95. The macrochaetal angle between the arm created by the median and exterior macrochaetae and the arm formed by the median and interior macrochaetae, MA (pleurotergite 15) ~120˚.
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</paragraph>
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<paragraph id="8BCE206BA7086C26FF42FDEF3CAFFD71" blockId="1.[151,1436,152,660]" pageId="1" pageNumber="148">
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<emphasis id="B905FC79A7086C26FF42FDEF3E6AFD94" box="[199,268,602,625]" italics="true" pageId="1" pageNumber="148">Telson</emphasis>
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: Epiproct with expanded apical part (dorsal view), and with pair of spinnerets. Each paraproct with three short marginal setae. Hypoproct subquadrangular with two long apical setae.
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</paragraph>
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<caption id="DF0E70E3A7086C26FF12F8E63E40F810" httpUri="https://zenodo.org/record/244416/files/figure.png" pageId="1" pageNumber="148" targetBox="[152,1411,703,1874]" targetPageId="1">
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<paragraph id="8BCE206BA7086C26FF12F8E63E40F810" blockId="1.[151,1436,1875,2037]" pageId="1" pageNumber="148">
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<emphasis id="B905FC79A7086C26FF12F8E63E24F88F" bold="true" box="[151,322,1875,1898]" pageId="1" pageNumber="148">FIGURES 1–4.</emphasis>
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<taxonomicName id="4C715BE8A7086C26FECCF8E63D08F88F" box="[329,622,1875,1898]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="karlstrasseri" status="sp. nov.">
|
||
<emphasis id="B905FC79A7086C26FECCF8E63D08F88F" box="[329,622,1875,1898]" italics="true" pageId="1" pageNumber="148">Heterolatzelia karlstrasseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2364102A7086C26FDF0F8E63DD7F88F" box="[629,689,1875,1898]" pageId="1" pageNumber="148" rank="species">
|
||
<emphasis id="B905FC79A7086C26FDF0F8E63DD7F88F" bold="true" box="[629,689,1875,1898]" pageId="1" pageNumber="148">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male gonopods. 1. anterior gonopods, oral view; 2. anterior gonopods, caudal view; 3. anterior gonopods, lateral view; 4. posterior gonopods, oral view. Designations:
|
||
<emphasis id="B905FC79A7086C26FA02F8C33AF2F868" bold="true" box="[1415,1428,1910,1933]" pageId="1" pageNumber="148">a</emphasis>
|
||
: angiocoxite;
|
||
<emphasis id="B905FC79A7086C26FEA6F82D3E54F84A" bold="true" box="[291,306,1944,1967]" pageId="1" pageNumber="148">b</emphasis>
|
||
: comb-like process;
|
||
<emphasis id="B905FC79A7086C26FD93F82D3D44F84A" bold="true" box="[534,546,1944,1967]" pageId="1" pageNumber="148">c</emphasis>
|
||
: colpocoxite;
|
||
<emphasis id="B905FC79A7086C26FD3BF82D3DABF84A" bold="true" box="[702,717,1944,1967]" pageId="1" pageNumber="148">d</emphasis>
|
||
: coxal cavity;
|
||
<emphasis id="B905FC79A7086C26FCF5F82D3C1AF84A" bold="true" box="[880,892,1944,1967]" pageId="1" pageNumber="148">e</emphasis>
|
||
: wide subquadrangular part;
|
||
<emphasis id="B905FC79A7086C26FB39F82D3BA3F84A" bold="true" box="[1212,1221,1944,1967]" pageId="1" pageNumber="148">f</emphasis>
|
||
: teeth of comb-like process;
|
||
<emphasis id="B905FC79A7086C26FF70F80E3F9AF837" bold="true" box="[245,252,1979,2002]" pageId="1" pageNumber="148">l</emphasis>
|
||
: connecting lamella between colpocoxites;
|
||
<emphasis id="B905FC79A7086C26FD56F80E3DBAF837" bold="true" box="[723,732,1979,2002]" pageId="1" pageNumber="148">t</emphasis>
|
||
: rudimentary telopodites;
|
||
<emphasis id="B905FC79A7086C26FC72F80E3B74F837" bold="true" box="[1015,1042,1979,2002]" pageId="1" pageNumber="148">cb</emphasis>
|
||
: caudal bulges of angiocoxite. Scale line: 0.4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCE206BA70B6C25FF42FF2D3AFAFF34" blockId="2.[151,1436,152,764]" pageId="2" pageNumber="149">
|
||
<emphasis id="B905FC79A70B6C25FF42FF2D3E37FF4A" box="[199,337,152,175]" italics="true" pageId="2" pageNumber="149">Walking legs</emphasis>
|
||
(
|
||
<typeStatus id="54CA9EC9A70B6C25FEE7FF2D3EA7FF4A" box="[354,449,152,175]" pageId="2" pageNumber="149" type="holotype">holotype</typeStatus>
|
||
male): Coxae and prefemores with denticles. Tarsi with papillae, and with two additional tarsal claws (one long and thin subapical accessory claw and one rudimentary claw situated at the base of the main claw).
|
||
</paragraph>
|
||
<paragraph id="8BCE206BA70B6C25FF42FF683C77FEDC" blockId="2.[151,1436,152,764]" pageId="2" pageNumber="149">
|
||
<emphasis id="B905FC79A70B6C25FF42FF683EA1FF11" box="[199,455,221,244]" italics="true" pageId="2" pageNumber="149">Male sexual characters</emphasis>
|
||
: Head with concave frontal side. Leg-pairs 1 and 2 slightly reduced, with tarsal brushes ventrally. Leg-pairs 3–7 somewhat enlarged, with tarsal papillae, and with more or less swollen podomeres. Leg-pairs 10 and 11 with coxal glands, without any additional processes.
|
||
</paragraph>
|
||
<paragraph id="8BCE206BA70B6C25FF42FEF03DF5FD71" blockId="2.[151,1436,152,764]" pageId="2" pageNumber="149">
|
||
<emphasis id="B905FC79A70B6C25FF42FEF03EF5FEB9" box="[199,403,325,348]" italics="true" pageId="2" pageNumber="149">Anterior gonopods</emphasis>
|
||
(
|
||
<figureCitation id="134A3CEEA70B6C25FE21FEF03D6BFEB9" box="[420,525,325,348]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,269,1875,1898]" captionTargetBox="[152,1411,703,1874]" captionTargetId="figure@1.[151,1436,678,1874]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Heterolatzelia karlstrasseri sp. n., holotype male gonopods. 1. anterior gonopods, oral view; 2. anterior gonopods, caudal view; 3. anterior gonopods, lateral view; 4. posterior gonopods, oral view. Designations: a: angiocoxite; b: comb-like process; c: colpocoxite; d: coxal cavity; e: wide subquadrangular part; f: teeth of comb-like process; l: connecting lamella between colpocoxites; t: rudimentary telopodites; cb: caudal bulges of angiocoxite. Scale line: 0.4 mm." httpUri="https://zenodo.org/record/244416/files/figure.png" pageId="2" pageNumber="149">Figs. 1–3</figureCitation>
|
||
): With a single, wide subquadrangular part (
|
||
<emphasis id="B905FC79A70B6C25FC79FEF03B6EFEB9" bold="true" box="[1020,1032,325,348]" pageId="2" pageNumber="149">e</emphasis>
|
||
) on the oral side. Anterior part with three elements. Angiocoxites (
|
||
<emphasis id="B905FC79A70B6C25FE59FEDD3E8FFE9A" bold="true" box="[476,489,360,383]" pageId="2" pageNumber="149">a</emphasis>
|
||
) are the most dominant and the longest elements; apically curved mesad; basal part wide in lateral view; with caudal bulges (
|
||
<emphasis id="B905FC79A70B6C25FD9FFE3F3D53FE44" bold="true" box="[538,565,394,417]" pageId="2" pageNumber="149">cb</emphasis>
|
||
). Large areas of angiocoxites are covered with papillae. Colpocoxites (
|
||
<emphasis id="B905FC79A70B6C25FAB0FE3F3A27FE44" bold="true" box="[1333,1345,394,417]" pageId="2" pageNumber="149">c</emphasis>
|
||
) shorter than angiocoxites, orally connected to each other by lamella (
|
||
<emphasis id="B905FC79A70B6C25FCACFE183C56FE21" bold="true" box="[809,816,429,452]" pageId="2" pageNumber="149">l</emphasis>
|
||
). Basal halves wide, subquadrangular in caudal view, and clearly separated from each other. Caudo-mesal sides of apical parts of colpocoxites with lobuses covered by papillae. From the widest parts of colpocoxites to their apical parts there are thin sutures. The third element is a comb-like process (
|
||
<emphasis id="B905FC79A70B6C25FF25FDA03FC9FDC9" bold="true" box="[160,175,533,556]" pageId="2" pageNumber="149">b</emphasis>
|
||
), starting from the oral side of anterior gonopods and slightly curving to the posterior part. These elements are sigmoid from the oral side, and 2/3 of their total length is covered with thin sutures. Apical parts are in the form of combs with 8+9 teeth (
|
||
<emphasis id="B905FC79A70B6C25FE95FDEF3E7FFD94" bold="true" box="[272,281,602,625]" pageId="2" pageNumber="149">f</emphasis>
|
||
) on the caudal side. Lateral sides of apical parts are devoid of teeth. Posterior part of anterior gonopods consists of large spoon-shaped coxal cavity (
|
||
<emphasis id="B905FC79A70B6C25FDF1FDC83DE5FD71" bold="true" box="[628,643,637,660]" pageId="2" pageNumber="149">d</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCE206BA70B6C25FF42FD153E58FD19" blockId="2.[151,1436,152,764]" pageId="2" pageNumber="149">
|
||
<emphasis id="B905FC79A70B6C25FF42FD153EFFFD52" box="[199,409,672,695]" italics="true" pageId="2" pageNumber="149">Posterior gonopods</emphasis>
|
||
(
|
||
<figureCitation id="134A3CEEA70B6C25FE2DFD153E8CFD52" box="[424,490,672,695]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,269,1875,1898]" captionTargetBox="[152,1411,703,1874]" captionTargetId="figure@1.[151,1436,678,1874]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Heterolatzelia karlstrasseri sp. n., holotype male gonopods. 1. anterior gonopods, oral view; 2. anterior gonopods, caudal view; 3. anterior gonopods, lateral view; 4. posterior gonopods, oral view. Designations: a: angiocoxite; b: comb-like process; c: colpocoxite; d: coxal cavity; e: wide subquadrangular part; f: teeth of comb-like process; l: connecting lamella between colpocoxites; t: rudimentary telopodites; cb: caudal bulges of angiocoxite. Scale line: 0.4 mm." httpUri="https://zenodo.org/record/244416/files/figure.png" pageId="2" pageNumber="149">Fig. 4</figureCitation>
|
||
): Massive, ovoid, consist of unique ladle-shaped coxites. Apical parts covered with short setae; orally with 2+2 setae (1+1 shorter and 1+1 longer). Posterior gonopods with telopodital remnants (
|
||
<emphasis id="B905FC79A70B6C25FA97FD773A7DFD3C" bold="true" box="[1298,1307,706,729]" pageId="2" pageNumber="149">t</emphasis>
|
||
), each with nine long setae.
|
||
</paragraph>
|
||
<paragraph id="8BCE206BA70B6C25FF12F9F93C3FF986" blockId="2.[151,857,1612,1635]" box="[151,857,1612,1635]" pageId="2" pageNumber="149">
|
||
<emphasis id="B905FC79A70B6C25FF12F9F93FBEF986" bold="true" box="[151,216,1612,1635]" pageId="2" pageNumber="149">MAP.</emphasis>
|
||
Distribution of the family
|
||
<taxonomicName id="4C715BE8A70B6C25FE73F9F93C33F986" authority="Verhoeff, 1897" authorityName="Verhoeff" authorityYear="1897" box="[502,853,1612,1635]" class="Diplopoda" family="Heterolatzeliidae" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="family">Heterolatzeliidae Verhoeff, 1897</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCE206BA70B6C25FF42F9273B0FF8F4" blockId="2.[151,1437,1682,2017]" pageId="2" pageNumber="149">
|
||
<emphasis id="B905FC79A70B6C25FF42F9273EE2F94C" bold="true" box="[199,388,1682,1705]" pageId="2" pageNumber="149">Taxonomic note.</emphasis>
|
||
Within the genus
|
||
<taxonomicName id="4C715BE8A70B6C25FDCBF9273D83F94C" box="[590,741,1682,1705]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905FC79A70B6C25FDCBF9273D83F94C" box="[590,741,1682,1705]" italics="true" pageId="2" pageNumber="149">Heterolatzelia</emphasis>
|
||
</taxonomicName>
|
||
, two species are closely related to each other:
|
||
<taxonomicName id="4C715BE8A70B6C25FB63F9273AFAF94C" box="[1254,1436,1682,1705]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="durmitorensis">
|
||
<emphasis id="B905FC79A70B6C25FB63F9273AFAF94C" box="[1254,1436,1682,1705]" italics="true" pageId="2" pageNumber="149">H. durmitorensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C715BE8A70B6C25FF43F9013E09F92E" box="[198,367,1716,1739]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="karlstrasseri" status="sp. nov.">
|
||
<emphasis id="B905FC79A70B6C25FF43F9013E09F92E" box="[198,367,1716,1739]" italics="true" pageId="2" pageNumber="149">H. karlstrasseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2364102A70B6C25FEFDF9013ED0F92E" box="[376,438,1716,1739]" pageId="2" pageNumber="149" rank="species">
|
||
<emphasis id="B905FC79A70B6C25FEFDF9013ED0F92E" bold="true" box="[376,438,1716,1739]" pageId="2" pageNumber="149">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
These species clearly differ from
|
||
<taxonomicName id="4C715BE8A70B6C25FCA8F9013CF0F92E" box="[813,918,1716,1739]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="nivalis">
|
||
<emphasis id="B905FC79A70B6C25FCA8F9013CF0F92E" box="[813,918,1716,1739]" italics="true" pageId="2" pageNumber="149">H. nivalis</emphasis>
|
||
</taxonomicName>
|
||
by the presence of long angiocoxites and short colpocoxites (these elements are almost the same length in
|
||
<taxonomicName id="4C715BE8A70B6C25FC8BF9623C10F90B" box="[782,886,1751,1774]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="nivalis">
|
||
<emphasis id="B905FC79A70B6C25FC8BF9623C10F90B" box="[782,886,1751,1774]" italics="true" pageId="2" pageNumber="149">H. nivalis</emphasis>
|
||
</taxonomicName>
|
||
). Apart from similarities in habitus and gonopodial structures,
|
||
<taxonomicName id="4C715BE8A70B6C25FE89F94F3ED9F8F4" box="[268,447,1786,1809]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="durmitorensis">
|
||
<emphasis id="B905FC79A70B6C25FE89F94F3ED9F8F4" box="[268,447,1786,1809]" italics="true" pageId="2" pageNumber="149">H. durmitorensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C715BE8A70B6C25FE76F94F3DFDF8F4" box="[499,667,1786,1809]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="karlstrasseri" status="sp. nov.">
|
||
<emphasis id="B905FC79A70B6C25FE76F94F3DFDF8F4" box="[499,667,1786,1809]" italics="true" pageId="2" pageNumber="149">H. karlstrasseri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B905FC79A70B6C25FD24F94F3DBBF8F4" bold="true" box="[673,733,1786,1809]" pageId="2" pageNumber="149">
|
||
<taxonomicNameLabel id="A2364102A70B6C25FD24F94F3DBBF8F4" box="[673,733,1786,1809]" pageId="2" pageNumber="149" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are also geographically close (Map).
|
||
</paragraph>
|
||
<paragraph id="8BCE206BA70B6C24FF42F8A93BFAFF34" blockId="2.[151,1437,1682,2017]" lastBlockId="3.[151,1435,152,209]" lastPageId="3" lastPageNumber="150" pageId="2" pageNumber="149">
|
||
In our description of the new species, we referred to the drawing of
|
||
<taxonomicName id="4C715BE8A70B6C25FC36F8A93B0EF8D6" box="[947,1128,1820,1843]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="durmitorensis">
|
||
<emphasis id="B905FC79A70B6C25FC36F8A93B0EF8D6" box="[947,1128,1820,1843]" italics="true" pageId="2" pageNumber="149">H. durmitorensis</emphasis>
|
||
</taxonomicName>
|
||
given in the paper of
|
||
<bibRefCitation id="EFE05D9AA70B6C25FADBF8A93F86F8B3" author="Mrsic" pageId="2" pageNumber="149" refString="Mrsic, N. (1992) Biokoviella mauriesi gen. nov., sp. nov. (Biokoviellidae fam. nov.), Macrochaetosomatinae subf. nov. (Anthogonidae) and superfamily Cleidogonoidea of the western Balkans (Craspedosomidea, Diplopoda). Razprave IV. Razreda SAZU, 33, 51 - 91." type="journal article" year="1992">Mršić (1992)</bibRefCitation>
|
||
, where he indicated that he examined numerous individuals of this species.
|
||
<taxonomicName id="4C715BE8A70B6C25FB8DF88A3A4BF8B3" box="[1032,1325,1855,1878]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="karlstrasseri" status="sp. nov.">
|
||
<emphasis id="B905FC79A70B6C25FB8DF88A3A4BF8B3" box="[1032,1325,1855,1878]" italics="true" pageId="2" pageNumber="149">Heterolatzelia karlstrasseri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2364102A70B6C25FAB6F88A3A09F8B3" box="[1331,1391,1855,1878]" pageId="2" pageNumber="149" rank="species">
|
||
<emphasis id="B905FC79A70B6C25FAB6F88A3A09F8B3" bold="true" box="[1331,1391,1855,1878]" pageId="2" pageNumber="149">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C715BE8A70B6C25FF12F8D73E28F89C" box="[151,334,1890,1913]" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="durmitorensis">
|
||
<emphasis id="B905FC79A70B6C25FF12F8D73E28F89C" box="[151,334,1890,1913]" italics="true" pageId="2" pageNumber="149">H. durmitorensis</emphasis>
|
||
</taxonomicName>
|
||
differ clearly from each other in some structures of the gonopods, such as the presence of a wide subquadrangular part on the oral side of the anterior gonopods in the new species
|
||
<emphasis id="B905FC79A70B6C25FBC9F8313B0CF87E" box="[1100,1130,1924,1947]" italics="true" pageId="2" pageNumber="149">vs.</emphasis>
|
||
absence of this part in
|
||
<taxonomicName id="4C715BE8A70B6C25FA07F8313E4CF85B" class="Diplopoda" family="Heterolatzeliidae" genus="Heterolatzelia" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="durmitorensis">
|
||
<emphasis id="B905FC79A70B6C25FA07F8313E4CF85B" italics="true" pageId="2" pageNumber="149">H. durmitorensis</emphasis>
|
||
</taxonomicName>
|
||
; apically curved mesad angiocoxites
|
||
<emphasis id="B905FC79A70B6C25FD7BF8123C7DF85B" box="[766,795,1959,1982]" italics="true" pageId="2" pageNumber="149">vs.</emphasis>
|
||
almost straight aniocoxites; colpocoxites with wide, subquadrangular basal halves
|
||
<emphasis id="B905FC79A70B6C25FE5CF87F3E90F804" box="[473,502,1994,2017]" italics="true" pageId="2" pageNumber="149">vs.</emphasis>
|
||
almost triangular colpocoxites; colpocoxites connected to each other by lamella on the oral side
|
||
<emphasis id="B905FC79A70A6C24FF7AFF2D3E7AFF4A" box="[255,284,152,175]" italics="true" pageId="3" pageNumber="150">vs.</emphasis>
|
||
absence of such connecting lamella; absence of small medial fimbriate processes on the inner edges of colpocoxites
|
||
<emphasis id="B905FC79A70A6C24FEA1FF0F3E27FF34" box="[292,321,186,209]" italics="true" pageId="3" pageNumber="150">vs.</emphasis>
|
||
presence of these processes; and ovoid vs. subquadrangular posterior gonopods.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |