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<document id="28FE2FF2231AA3657670A210307420F0" ID-DOI="10.17109/AZH.65.2.123.2019" ID-ISSN="2064-2474" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="julia" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638275451890" checkinUser="felipe" docAuthor="Szederjesi, Tímea, Pop, Victor V., Márton, Orsolya &amp; Csuzdi, Csaba" docDate="2019" docId="03C187DC1141FF9042FAFB78686FAD82" docLanguage="en" docName="ActaZoolAcadSciH.65.2.123-142.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 65 (2)" docSource="http://dx.doi.org/10.17109/azh.65.2.123.2019" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.5:ActaZoolAcadSciH.2015-2023.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-2023.journal_article" docStyleVersion="5" docTitle="Lumbricus terrestris Linnaeus 1758" docType="treatment" docVersion="1" lastPageNumber="136" masterDocId="FFF8FFA4114AFF9E4356FFD66A26AD7B" masterDocTitle="New Earthworm Species And Records From The Southern Carpathians (Megadrili: Lumbricidae)" masterLastPageNumber="142" masterPageNumber="123" pageNumber="134" updateTime="1719927152464" updateUser="julia" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="5F02FC7B119DC8CE53E2A923BBB2833C">New Earthworm Species And Records From The Southern Carpathians (Megadrili: Lumbricidae)</mods:title>
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<mods:namePart id="F132D0740BC6E0D43813545E73C78823">Szederjesi, Tímea</mods:namePart>
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<mods:namePart id="0012EE1B5F26A608BB00BBAF3C8125E7">Pop, Victor V.</mods:namePart>
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<taxonomicName id="4C684D491141FF9542FAFB78694EA9B2" ID-CoL="6R26F" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[428,872,1198,1225]" class="Clitellata" family="Lumbricidae" genus="Lumbricus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="134" phylum="Annelida" rank="species" species="terrestris">
<emphasis id="B91CEAD81141FF9542FAFB786886A9B3" box="[428,672,1198,1224]" italics="true" pageId="11" pageNumber="134">Lumbricus terrestris</emphasis>
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<taxonomicName id="4C684D491141FF9543F8FB30686AA987" ID-CoL="6R26F" authority="Linnaeus, 1758: 647" authorityName="Linnaeus" authorityPageNumber="647" authorityYear="1758" box="[174,588,1254,1276]" class="Clitellata" family="Lumbricidae" genus="Lumbricus" kingdom="Animalia" order="Crassiclitellata" pageId="11" pageNumber="134" phylum="Annelida" rank="species" species="terrestris">
<emphasis id="B91CEAD81141FF9543F8FB306B53A987" box="[174,373,1254,1276]" italics="true" pageId="11" pageNumber="134">Lumbricus terrestris</emphasis>
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.
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<bibRefCitation id="EFF94B3B1141FF954103FB316936A987" author="POP, V. V. &amp; POP, A. A. &amp; CSUZDI, CS" box="[597,784,1254,1276]" pageId="11" pageNumber="134" pagination="59 - 70" refId="ref11375" refString="POP, V. V., POP, A. A. &amp; CSUZDI, CS. (2012): An annotated checklist of the Romanian earth- worm fauna (Oligochaeta, Lumbricidae). - Zoology in the Middle East 4: 59 - 70." type="journal article" year="2012">
Pop
<emphasis id="B91CEAD81141FF9541D3FB316892A980" box="[645,692,1254,1276]" italics="true" pageId="11" pageNumber="134">et al.</emphasis>
2012: 64
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.
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<collectingMunicipality id="6BB3ACB01141FF9543BCFAF66B65A84D" box="[234,323,1312,1334]" pageId="11" pageNumber="134">Material</collectingMunicipality>
examined.
<specimenCode id="DBCE9EB11141FF954294FAF6687FA84E" box="[450,601,1312,1333]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="11" pageNumber="134">HNHM/17214</specimenCode>
<specimenCount id="9D6EFD431141FF954137FAF668B3A84E" box="[609,661,1312,1333]" pageId="11" pageNumber="134" type="generic">4 ex.</specimenCount>
,
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, stream valley NW of
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,
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,
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, chestnut forest patch, leg.
<collectorName id="269D531C1141FF95404EFAEB6981A829" box="[792,935,1341,1363]" pageId="11" pageNumber="134">Cs. Csuzdi, L</collectorName>
. Dányi,
<collectionCode id="ED79AE0F1141FF9540A8FAEB6E37A829" box="[1022,1041,1341,1362]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="134" type="Museum">V</collectionCode>
.
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.
<collectorName id="269D531C1141FF954760FAEB6E44A829" box="[1078,1122,1341,1362]" pageId="11" pageNumber="134">Pop</collectorName>
,
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.
<collectorName id="269D531C1141FF95439CFA8C6B10A80B" box="[202,310,1370,1392]" pageId="11" pageNumber="134">Szederjesi</collectorName>
,
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<collectingDate id="EF92E9E21141FF954214FA8F6B97A814" box="[322,433,1369,1391]" pageId="11" pageNumber="134" value="2016-11-13">13.11.2016</collectingDate>
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.
<materialsCitation id="3B003C971141FF9542EAFA8F69D2A8F7" collectingDate="2018-08-29" collectionCode="T" collectorName="Asami, Z" elevation="1070" location="Coliba Haiducilor" longitude="45.5885" municipality="Poiana" pageId="11" pageNumber="134" specimenCode="HNHM/17362" specimenCount="1" stateProvince="Brasov">
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<specimenCount id="9D6EFD431141FF95410CFA8C68A8A814" box="[602,654,1370,1391]" pageId="11" pageNumber="134" type="generic">1 ex.</specimenCount>
,
<collectingRegion id="49ACF8281141FF9541CCFA8C6910A814" box="[666,822,1370,1393]" country="Romania" name="Brasov" pageId="11" pageNumber="134">Brașov county</collectingRegion>
,
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<collectingRegion id="49ACF8281141FF9540C5FA8C69F8A80A" box="[915,990,1370,1393]" country="Romania" name="Brasov" pageId="11" pageNumber="134">Brașov</collectingRegion>
, near
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, forest,
<quantity id="4C909B2F1141FF954223FAA16BE6A8F7" box="[373,448,1399,1420]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="11" pageNumber="134" unit="m" value="1070.0">
<elevation id="0045D1F91141FF954223FAA16BE6A8F7" box="[373,448,1399,1420]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="11" pageNumber="134" unit="m" value="1070.0">1070 m</elevation>
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N
<geoCoordinate id="EE5C500D1141FF95428EFAA0681BA8F7" box="[472,573,1398,1420]" degrees="45.5885" direction="east" orientation="longitude" pageId="11" pageNumber="134" precision="5" value="45.5885">45.5885 E</geoCoordinate>
25.554, leg.
<collectionCode id="ED79AE0F1141FF9541E1FAA168EEA8F7" box="[695,712,1399,1420]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="134">T</collectionCode>
.
<collectorName id="269D531C1141FF954186FAA06912A8F7" box="[720,820,1398,1420]" pageId="11" pageNumber="134">Asami, Z</collectorName>
. Fehér,
<date id="FFD6100A1141FF9540D3FAA169D2A8F7" box="[901,1012,1399,1420]" pageId="11" pageNumber="134" value="2018-08-29">
<collectingDate id="EF92E9E21141FF9540D3FAA169D2A8F7" box="[901,1012,1399,1420]" pageId="11" pageNumber="134" value="2018-08-29">29.08.2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD736CA1146FF92425AFF68699BADA2" blockId="12.[268,957,190,217]" box="[268,957,190,217]" pageId="12" pageNumber="135">
<taxonomicName id="4C684D491146FF92425AFF686956ADA3" authority="Szederjesi, Pop et Csuzdi" authorityName="Szederjesi, Pop et Csuzdi" box="[268,880,190,217]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="135" phylum="Annelida" rank="species" species="banaticus" status="sp. nov.">
<emphasis id="B91CEAD81146FF92425AFF686805ADA3" bold="true" box="[268,547,190,216]" pageId="12" pageNumber="135">Octodrilus banaticus</emphasis>
Szederjesi, Pop et Csuzdi
</taxonomicName>
<taxonomicNameLabel id="A22F57A31146FF92402EFF69699BADA2" box="[888,957,191,217]" pageId="12" pageNumber="135" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BD736CA1146FF924161FF3468B4AD87" blockId="12.[567,658,226,252]" box="[567,658,226,252]" pageId="12" pageNumber="135">
<heading id="D09F81A61146FF924161FF3468B4AD87" box="[567,658,226,252]" centered="true" fontSize="11" level="3" pageId="12" pageNumber="135" reason="3">
(
<figureCitation id="13532A4F1146FF924116FF3468AEAD87" box="[576,648,226,252]" captionStart="Fig" captionStartId="12.[136,172,1487,1509]" captionTargetBox="[147,1064,1233,1461]" captionTargetId="figure-398@12.[136,1088,1225,1469]" captionTargetPageId="12" captionText="Fig. 3. Ventrolateral view of Octodrilus banaticus sp. n. Mp = male pore, Cl = clitellum, Tb = tubercles" pageId="12" pageNumber="135">Fig. 3</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BD736CA1146FF924393FECC69F8ACC4" blockId="12.[136,1089,281,448]" pageId="12" pageNumber="135">
<materialsCitation id="3B003C971146FF924393FECC6B6DAC12" collectingDate="2016-10-11" collectionCode="V, T" collectorName="Cs. Csuzdi, L &amp; L. Danyi &amp; Pop" elevation="495" latitude="45.341938" location="Tarcu Mts" longLatPrecision="1" longitude="22.407238" pageId="12" pageNumber="135" specimenCode="HNHM/17361" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D388681146FF924393FECC6B0AAC4B" box="[197,300,282,304]" pageId="12" pageNumber="135" type="holotype">Holotype</typeStatus>
.
<specimenCode id="DBCE9EB11146FF92426CFECF6BF6AC54" box="[314,464,281,303]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="12" pageNumber="135">HNHM/17361</specimenCode>
<location id="8EB760111146FF92428AFECC6876AC54" LSID="urn:lsid:plazi:treatment:03C187DC1141FF9042FAFB78686FAD82:8EB760111146FF92428AFECC6876AC54" box="[476,592,282,303]" latitude="45.341938" longLatPrecision="1" longitude="22.407238" name="Tarcu Mts" pageId="12" pageNumber="135">Ţarcu Mts</location>
,
<locationDeviation id="9A652FE81146FF924136FECC69C1AC4B" box="[608,999,282,304]" firstLocation="Borlova" pageId="12" pageNumber="135" secondLocation="Muntele Mic">between Borlova and Muntele Mic</locationDeviation>
,
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<elevation id="0045D1F91146FF9240A0FECF6E1AAC54" box="[1014,1084,281,303]" metricMagnitude="2" metricUnit="m" metricValue="4.95" pageId="12" pageNumber="135" unit="m" value="495.0">495 m</elevation>
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,
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<geoCoordinate id="EE5C500D1146FF92424EFEE16B86AC37" box="[280,416,311,332]" degrees="22" direction="east" minutes="24.4343" orientation="longitude" pageId="12" pageNumber="135" precision="1" value="22.407238">22°24.4343’E</geoCoordinate>
, mixed beech forest, leg.
<collectorName id="269D531C1146FF9241FEFEE16911AC37" box="[680,823,310,332]" pageId="12" pageNumber="135">Cs. Csuzdi, L</collectorName>
. Dányi,
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.
<collectionCode id="ED79AE0F1146FF9240FFFEE1699AAC37" box="[937,956,311,332]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="135" type="Museum">V</collectionCode>
.
<collectorName id="269D531C1146FF924093FEE169D7AC37" box="[965,1009,311,332]" pageId="12" pageNumber="135">Pop</collectorName>
,
<collectionCode id="ED79AE0F1146FF9240ADFEE16E2AAC37" box="[1019,1036,311,332]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="135">T</collectionCode>
. Sze- derjesi,
<date id="FFD6100A1146FF92438AFE856B6DAC12" box="[220,331,339,361]" pageId="12" pageNumber="135" value="2016-10-11">
<collectingDate id="EF92E9E21146FF92438AFE856B6DAC12" box="[220,331,339,361]" pageId="12" pageNumber="135" value="2016-10-11">10.11.2016</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B003C971146FF924200FE826BAFACFD" collectionCode="HNHM" pageId="12" pageNumber="135" specimenCode="HNHM/17231" specimenCount="7" typeStatus="paratype">
<typeStatus id="54D388681146FF924200FE826BE7AC12" box="[342,449,340,361]" pageId="12" pageNumber="135" type="paratype">Paratypes</typeStatus>
.
<specimenCode id="DBCE9EB11146FF92429AFE856844AC13" box="[460,610,339,360]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="12" pageNumber="135">HNHM/17231</specimenCode>
<specimenCount id="9D6EFD431146FF92413CFE8568BAAC12" box="[618,668,339,361]" pageId="12" pageNumber="135" type="generic">1 ex.</specimenCount>
praeadult +
<specimenCount id="9D6EFD431146FF924071FE85697FAC12" box="[807,857,339,361]" pageId="12" pageNumber="135" type="generic">6 ex.</specimenCount>
juvenile, locality and date same as that of the
</materialsCitation>
<materialsCitation id="3B003C971146FF9242C7FEA669FDACC4" collectingDate="2016-10-11" collectionCode="V, T" collectorName="Cs. Csuzdi, L &amp; L. Danyi &amp; Pop &amp; Szederjesi" elevation="443" latitude="45.434624" location="Tarcu Mts" longLatPrecision="1" longitude="22.457136" pageId="12" pageNumber="135" specimenCode="HNHM/17240" specimenCount="4" typeStatus="holotype">
<typeStatus id="54D388681146FF9242C7FEA66BDEACFD" box="[401,504,368,390]" pageId="12" pageNumber="135" type="holotype">Holotype</typeStatus>
.
<specimenCode id="DBCE9EB11146FF924155FEA668BFACFE" box="[515,665,368,389]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="12" pageNumber="135">HNHM/17240</specimenCode>
<specimenCount id="9D6EFD431146FF9241F7FEA668F5ACFE" box="[673,723,368,389]" pageId="12" pageNumber="135" type="generic">1 ex.</specimenCount>
subadult +
<specimenCount id="9D6EFD431146FF924005FEA669A3ACFE" box="[851,901,368,389]" pageId="12" pageNumber="135" type="generic">2 ex.</specimenCount>
praeadult +
<specimenCount id="9D6EFD431146FF924759FEA66E67ACFE" box="[1039,1089,368,389]" pageId="12" pageNumber="135" type="generic">1 ex.</specimenCount>
juvenile,
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,
<locationDeviation id="9A652FE81146FF924209FE5B68F4ACD8" box="[351,722,397,419]" firstLocation="Maru" pageId="12" pageNumber="135" secondLocation="Poiana Marului">between Măru and Poiana Mărului</locationDeviation>
,
<quantity id="4C909B2F1146FF92418AFE5B693AACD9" box="[732,796,397,418]" metricMagnitude="2" metricUnit="m" metricValue="4.43" pageId="12" pageNumber="135" unit="m" value="443.0">
<elevation id="0045D1F91146FF92418AFE5B693AACD9" box="[732,796,397,418]" metricMagnitude="2" metricUnit="m" metricValue="4.43" pageId="12" pageNumber="135" unit="m" value="443.0">443 m</elevation>
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,
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<geoCoordinate id="EE5C500D1146FF9240E0FE5B6E1AACD9" box="[950,1084,397,418]" degrees="22" direction="east" minutes="27.4282" orientation="longitude" pageId="12" pageNumber="135" precision="1" value="22.457136">22°27.4282’E</geoCoordinate>
, mixed beech forest, leg.
<collectorName id="269D531C1146FF9242DEFE7C6831ACC4" box="[392,535,426,448]" pageId="12" pageNumber="135">Cs. Csuzdi, L</collectorName>
. Dányi,
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.
<collectionCode id="ED79AE0F1146FF9241DCFE7C68BBACC4" box="[650,669,426,447]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="135" type="Museum">V</collectionCode>
.
<collectorName id="269D531C1146FF9241F0FE7C68F4ACC4" box="[678,722,426,447]" pageId="12" pageNumber="135">Pop</collectorName>
,
<collectionCode id="ED79AE0F1146FF92418AFE7C68CBACC4" box="[732,749,426,447]" name="Tavera, Department of Geology and Geophysics" pageId="12" pageNumber="135">T</collectionCode>
.
<collectorName id="269D531C1146FF9241A3FE7C6947ACBB" box="[757,865,426,448]" pageId="12" pageNumber="135">Szederjesi</collectorName>
,
<date id="FFD6100A1146FF92403DFE7F69FDACC4" box="[875,987,425,447]" pageId="12" pageNumber="135" value="2016-10-11">
<collectingDate id="EF92E9E21146FF92403DFE7F69FDACC4" box="[875,987,425,447]" pageId="12" pageNumber="135" value="2016-10-11">10.11.2016</collectingDate>
</date>
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.
</paragraph>
<paragraph id="8BD736CA1146FF924393FE356AF5AF6D" blockId="12.[136,1088,483,534]" pageId="12" pageNumber="135">
Etymology. The name refers to the geographic region of the
<typeStatus id="54D388681146FF92401DFE32695CAC82" box="[843,890,484,505]" pageId="12" pageNumber="135">type</typeStatus>
locality, the Banat region.
</paragraph>
<paragraph id="8BD736CA1146FF924393FDED6BE7AE65" blockId="12.[136,1089,571,798]" pageId="12" pageNumber="135">
Diagnosis. Length
<quantity id="4C909B2F1146FF9242FAFDED686FAF2E" box="[428,585,571,598]" metricMagnitude="-1" metricUnit="m" metricValue="1.58" metricValueMax="1.71" metricValueMin="1.45" pageId="12" pageNumber="135" unit="mm" value="158.0" valueMax="171.0" valueMin="145.0">145–171 mm</quantity>
, diameter
<quantity id="4C909B2F1146FF92419CFDED6909AF2E" box="[714,815,571,597]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="12" pageNumber="135" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
, setae distantly stand- ing. Pigmentation greyish. First dorsal pore in 11/12. Clitellum on ½ 29–½ 39, ½ 40, tubercles on 30–½ 39, ½ 40. Male pores on segment 15. Nephridial pores aligned above setal line
<emphasis id="B91CEAD81146FF9242EAFD496BF6AFC2" box="[444,464,671,697]" italics="true" pageId="12" pageNumber="135">b.</emphasis>
Three pairs of seminal vesicles in 9, 10, 12; sper- mathecae 7 (6–8) pairs in 5/6–11/12 (12/13) in setal line
<emphasis id="B91CEAD81146FF92406DFD17696FAFA0" box="[827,841,705,731]" italics="true" pageId="12" pageNumber="135">d</emphasis>
. Calciferous glands with well-developed diverticula in 10. Hearts in segments 7–11, nephridial bladders ocarina-shaped.
</paragraph>
<paragraph id="8BD736CA1146FF924393FC946B4EA97F" blockId="12.[136,1089,833,1201]" pageId="12" pageNumber="135">
External characters.
<typeStatus id="54D388681146FF9242F6FC946822AE23" box="[416,516,834,856]" pageId="12" pageNumber="135" type="holotype">Holotype</typeStatus>
<quantity id="4C909B2F1146FF92415AFC976844AE2C" box="[524,610,833,855]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" pageId="12" pageNumber="135" unit="mm" value="145.0">145 mm</quantity>
long and
<quantity id="4C909B2F1146FF924185FC946937AE2C" box="[723,785,834,855]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="135" unit="mm" value="7.0">7 mm</quantity>
wide. Number of segments 184.
<typeStatus id="54D388681146FF9243EFFC896B07AE0F" box="[185,289,863,884]" pageId="12" pageNumber="135" type="paratype">Paratypes</typeStatus>
<quantity id="4C909B2F1146FF92427FFC886B88AE0F" box="[297,430,862,884]" metricMagnitude="-1" metricUnit="m" metricValue="1.585" metricValueMax="1.71" metricValueMin="1.46" pageId="12" pageNumber="135" unit="mm" value="158.5" valueMax="171.0" valueMin="146.0">146–171 mm</quantity>
long and
<quantity id="4C909B2F1146FF92414AFC8868A5AE0F" box="[540,643,862,884]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="12" pageNumber="135" unit="mm" value="6.25" valueMax="6.5" valueMin="6.0">6–6.5 mm</quantity>
wide. Number of segments 180–223. Pig- mentation greyish. Prostomium epilobous ⅓ closed. First dorsal pore at the intersegmen- tal furrow 11/12. Setae distantly standing. Setal arrangement behind clitellum: aa:ab:bc: cd:dd = 3.38:2.13:1.22:1:5,07. Male pores on segment 15, small. Nephridial pores aligned above setal line
<emphasis id="B91CEAD81146FF924261FC046B6FAE9C" box="[311,329,978,999]" italics="true" pageId="12" pageNumber="135">b.</emphasis>
Clitellum on segments ½ 29–½ 39, ½ 40. Tubercula pubertatis on seg- ments 30–½ 39, ½ 40.
</paragraph>
<paragraph id="8BD736CA1146FF934393FBDD68B4AE83" blockId="12.[136,1089,833,1201]" lastBlockId="13.[173,1127,908,1589]" lastPageId="13" lastPageNumber="136" pageId="12" pageNumber="135">
Internal characters. Septa 6/7–8/9 thickened, 12/13–14/15 strongly. Testes and fun- nels paired in segments 10–11, covered by oesophageal testis sacs in segment 10. The dis- sepiments of 10/11 and 11/12 merged together, completely enclosing the male funnels of 11. Three pairs of seminal vesicles in 9, 10, 12. Usually seven (sometimes 6 or 8) pairs of spermathecae in 5/6–11/12 with external openings in setal line
<emphasis id="B91CEAD81146FF924001FBA8694CA9E8" box="[855,874,1150,1171]" italics="true" pageId="12" pageNumber="135">d.</emphasis>
In some specimens asymmetrically missing from 5/6 or extra pair present in 12/13. Calciferous glands with well-developed diverticula in segment 10. Paired hearts in segments 7–11. with a pair of small extraoesophageal vessels in 12. Nephridial bladders ocarina-shaped. Crop in seg- ments 15–16, and gizzard in segments 17–18. Typhlosolis lamelliform. The cross-section of the longitudinal muscle layer of pinnate
<typeStatus id="54D388681147FF93410AFC3568A8AE83" box="[604,654,995,1016]" pageId="13" pageNumber="136">type</typeStatus>
.
</paragraph>
<caption id="DF1766421146FF9243DEFA1968B2AB79" pageId="12" pageNumber="135" startId="12.[136,172,1487,1509]" targetBox="[147,1064,1233,1461]" targetPageId="12">
<paragraph id="8BD736CA1146FF9243DEFA1968B2AB79" blockId="12.[136,1089,1487,1538]" pageId="12" pageNumber="135">
<emphasis id="B91CEAD81146FF9243DEFA196AEFA89F" bold="true" box="[136,201,1487,1509]" pageId="12" pageNumber="135">Fig. 3.</emphasis>
Ventrolateral view of
<taxonomicName id="4C684D491146FF9242E3FA1968A6A89F" box="[437,640,1487,1508]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="12" pageNumber="135" phylum="Annelida" rank="species" species="banaticus" status="sp. nov.">
<emphasis id="B91CEAD81146FF9242E3FA1968A6A89F" box="[437,640,1487,1508]" italics="true" pageId="12" pageNumber="135">Octodrilus banaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57A31146FF9241D0FA066899A89E" box="[646,703,1488,1509]" pageId="12" pageNumber="135" rank="species">sp. n.</taxonomicNameLabel>
Mp = male pore, Cl = clitellum, Tb = tubercles
</paragraph>
</caption>
<caption id="DF1766421147FF9343A7FF176E02ADAC" box="[241,1060,193,215]" pageId="13" pageNumber="136" startId="13.[241,302,193,214]">
<paragraph id="8BD736CA1147FF9343A7FF176E02ADAC" blockId="13.[241,1060,193,215]" box="[241,1060,193,215]" pageId="13" pageNumber="136">
<emphasis id="B91CEAD81147FF9343A7FF176B60ADAD" bold="true" box="[241,326,193,214]" pageId="13" pageNumber="136">Table 4.</emphasis>
Distinguishing characters of the species similar to
<taxonomicName id="4C684D491147FF934036FF1469C3ADAD" box="[864,997,193,215]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="136" phylum="Annelida" rank="species" species="banaticus" status="sp. nov.">
<emphasis id="B91CEAD81147FF934036FF1469C3ADAD" box="[864,997,193,215]" italics="true" pageId="13" pageNumber="136">Oc. banaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57A31147FF9340BDFF146E02ADAC" box="[1003,1060,194,215]" pageId="13" pageNumber="136" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BD736CA1147FF9343EFFF3E6E1BAE2F" pageId="13" pageNumber="136">
<table id="F968C46A1147006143EFFF3E6E7FAE2D" box="[185,1113,232,854]" gridcols="7" gridrows="9" pageId="13" pageNumber="136">
<tr id="355834881147006143EFFF3E6E7FAC4C" box="[185,1113,232,311]" gridrow="0" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFF3E6BA8AC4C" box="[185,398,232,311]" gridcol="0" gridrow="0" pageId="13" pageNumber="136">Species</th>
<th id="76895DF41147006142FFFF3E6804AC4C" box="[425,546,232,311]" gridcol="1" gridrow="0" pageId="13" pageNumber="136">Clitellum</th>
<th id="76895DF411470061416FFF3E68B9AC4C" box="[569,671,232,311]" gridcol="2" gridrow="0" pageId="13" pageNumber="136">Tubercles</th>
<th id="76895DF41147006141E9FF3E68D3AC4C" box="[703,757,232,311]" gridcol="3" gridrow="0" pageId="13" pageNumber="136">Vesi- cles</th>
<th id="76895DF411470061404EFF3E6957AC4C" box="[792,881,232,311]" gridcol="4" gridrow="0" pageId="13" pageNumber="136">Sperma- thecae</th>
<th id="76895DF41147006140DFFF3E69C5AC4C" box="[905,995,232,311]" gridcol="5" gridrow="0" pageId="13" pageNumber="136">Testic sac</th>
<th id="76895DF41147006140ADFF3E6E7FAC4C" box="[1019,1113,232,311]" gridcol="6" gridrow="0" pageId="13" pageNumber="136">Nephridial pores</th>
</tr>
<tr id="355834881147006143EFFE916E7FAC07" box="[185,1113,327,380]" gridrow="1" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFE916BA8AC07" box="[185,398,327,380]" gridcol="0" gridrow="1" pageId="13" pageNumber="136">
<taxonomicName id="4C684D491147FF9343EFFE816B18AC17" box="[185,318,343,364]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="136" phylum="Annelida" rank="species" species="banaticus" status="sp. nov.">
<emphasis id="B91CEAD81147FF9343EFFE816B18AC17" box="[185,318,343,364]" italics="true" pageId="13" pageNumber="136">Oc. banaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57A31147FF934212FE8E6B5BAC16" box="[324,381,344,365]" pageId="13" pageNumber="136" rank="species">sp. n.</taxonomicNameLabel>
</th>
<td id="76895DF41147006142FFFE916804AC07" box="[425,546,327,380]" gridcol="1" gridrow="1" pageId="13" pageNumber="136">½ 29–½ 39, ½ 40</td>
<td id="76895DF411470061416FFE9168B9AC07" box="[569,671,327,380]" gridcol="2" gridrow="1" pageId="13" pageNumber="136">30–½ 39, ½ 40</td>
<td id="76895DF41147006141E9FE9168D3AC07" box="[703,757,327,380]" gridcol="3" gridrow="1" pageId="13" pageNumber="136">9, 10, 12</td>
<td id="76895DF411470061404EFE916957AC07" box="[792,881,327,380]" gridcol="4" gridrow="1" pageId="13" pageNumber="136">7 (5/6– 11/12)</td>
<td id="76895DF41147006140DFFE9169C5AC07" box="[905,995,327,380]" gridcol="5" gridrow="1" pageId="13" pageNumber="136">10 oeso</td>
<td id="76895DF41147006140ADFE916E7FAC07" box="[1019,1113,327,380]" gridcol="6" gridrow="1" pageId="13" pageNumber="136">
aligned in
<emphasis id="B91CEAD81147FF934764FEB36E1BAC01" box="[1074,1085,357,378]" italics="true" pageId="13" pageNumber="136">b</emphasis>
</td>
</tr>
<tr id="355834881147006143EFFE5A6E7FACC4" box="[185,1113,396,447]" gridrow="2" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFE5A6BA8ACC4" box="[185,398,396,447]" gridcol="0" gridrow="2" pageId="13" pageNumber="136">
<taxonomicName id="4C684D491147FF9343EFFE4D6BA8ACCB" authority="Pop, 1989" authorityName="Pop" authorityYear="1989" box="[185,398,411,432]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="136" phylum="Annelida" rank="species" species="aporus">
<emphasis id="B91CEAD81147FF9343EFFE4D6B04ACCB" box="[185,290,411,432]" italics="true" pageId="13" pageNumber="136">Oc. aporus</emphasis>
Pop, 1989
</taxonomicName>
</th>
<td id="76895DF41147006142FFFE5A6804ACC4" box="[425,546,396,447]" gridcol="1" gridrow="2" pageId="13" pageNumber="136">½ 29, 30–40</td>
<td id="76895DF411470061416FFE5A68B9ACC4" box="[569,671,396,447]" gridcol="2" gridrow="2" pageId="13" pageNumber="136">30–40</td>
<td id="76895DF41147006141E9FE5A68D3ACC4" box="[703,757,396,447]" gridcol="3" gridrow="2" pageId="13" pageNumber="136">9–12</td>
<td id="76895DF411470061404EFE5A6957ACC4" box="[792,881,396,447]" gridcol="4" gridrow="2" pageId="13" pageNumber="136">6 (5/6– 10/11)</td>
<td id="76895DF41147006140DFFE5A69C5ACC4" box="[905,995,396,447]" gridcol="5" gridrow="2" pageId="13" pageNumber="136">10, 11 perioeso</td>
<td id="76895DF41147006140ADFE5A6E7FACC4" box="[1019,1113,396,447]" gridcol="6" gridrow="2" pageId="13" pageNumber="136">irregular</td>
</tr>
<tr id="355834881147006143EFFE066E7FAF78" box="[185,1113,464,515]" gridrow="3" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFE066BA8AF78" box="[185,398,464,515]" gridcol="0" gridrow="3" pageId="13" pageNumber="136">
<emphasis id="B91CEAD81147FF9343EFFE066B72AC9E" box="[185,340,464,485]" italics="true" pageId="13" pageNumber="136">Oc. hemiandrus</emphasis>
(Cognetti, 1901)
</th>
<td id="76895DF41147006142FFFE066804AF78" box="[425,546,464,515]" gridcol="1" gridrow="3" pageId="13" pageNumber="136">28–36, 37</td>
<td id="76895DF411470061416FFE0668B9AF78" box="[569,671,464,515]" gridcol="2" gridrow="3" pageId="13" pageNumber="136">28–38, 39</td>
<td id="76895DF41147006141E9FE0668D3AF78" box="[703,757,464,515]" gridcol="3" gridrow="3" pageId="13" pageNumber="136">9, 10, 12</td>
<td id="76895DF411470061404EFE066957AF78" box="[792,881,464,515]" gridcol="4" gridrow="3" pageId="13" pageNumber="136">7 (6/7– 12/13)</td>
<td id="76895DF41147006140DFFE0669C5AF78" box="[905,995,464,515]" gridcol="5" gridrow="3" pageId="13" pageNumber="136">11 oeso</td>
<td id="76895DF41147006140ADFE066E7FAF78" box="[1019,1113,464,515]" gridcol="6" gridrow="3" pageId="13" pageNumber="136">irregular</td>
</tr>
<tr id="355834881147006143EFFDC26E7FAF3C" box="[185,1113,532,583]" gridrow="4" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFDC26BA8AF3C" box="[185,398,532,583]" gridcol="0" gridrow="4" pageId="13" pageNumber="136">
<emphasis id="B91CEAD81147FF9343EFFDC26B61AF52" box="[185,327,532,553]" italics="true" pageId="13" pageNumber="136">Oc. velebiticus</emphasis>
<bibRefCitation id="EFF94B3B1147FF9343EFFDE76B17AF3D" author="MRSIC, N." box="[185,305,561,583]" pageId="13" pageNumber="136" refId="ref11082" refString="MRSIC, N. (1991): Monograph on earthworms (Lumbricidae) of the Balkans I - II. - Slovenska Akademija Znanosti in Umetnosti, Ljubljana, 757 pp." type="book" year="1991">Mršić, 1991</bibRefCitation>
</th>
<td id="76895DF41147006142FFFDC26804AF3C" box="[425,546,532,583]" gridcol="1" gridrow="4" pageId="13" pageNumber="136">½ 29–39</td>
<td id="76895DF411470061416FFDC268B9AF3C" box="[569,671,532,583]" gridcol="2" gridrow="4" pageId="13" pageNumber="136">30–39</td>
<td id="76895DF41147006141E9FDC268D3AF3C" box="[703,757,532,583]" gridcol="3" gridrow="4" pageId="13" pageNumber="136">9–12</td>
<td id="76895DF411470061404EFDC26957AF3C" box="[792,881,532,583]" gridcol="4" gridrow="4" pageId="13" pageNumber="136">7 (6/7– 12/13)</td>
<td id="76895DF41147006140DFFDC269C5AF3C" box="[905,995,532,583]" gridcol="5" gridrow="4" pageId="13" pageNumber="136">10, 11 oeso</td>
<td id="76895DF41147006140ADFDC26E7FAF3C" box="[1019,1113,532,583]" gridcol="6" gridrow="4" pageId="13" pageNumber="136">irregular</td>
</tr>
<tr id="355834881147006143EFFD816E7FAFF1" box="[185,1113,599,650]" gridrow="5" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFD816BA8AFF1" box="[185,398,599,650]" gridcol="0" gridrow="5" pageId="13" pageNumber="136">
<emphasis id="B91CEAD81147FF9343EFFD8E6B7AAF17" box="[185,348,599,621]" italics="true" pageId="13" pageNumber="136">Oc. complanatus</emphasis>
(Dugès, 1828)
</th>
<td id="76895DF41147006142FFFD816804AFF1" box="[425,546,599,650]" gridcol="1" gridrow="5" pageId="13" pageNumber="136">28, 29–37</td>
<td id="76895DF411470061416FFD8168B9AFF1" box="[569,671,599,650]" gridcol="2" gridrow="5" pageId="13" pageNumber="136">30–39, 40</td>
<td id="76895DF41147006141E9FD8168D3AFF1" box="[703,757,599,650]" gridcol="3" gridrow="5" pageId="13" pageNumber="136">9–12</td>
<td id="76895DF411470061404EFD816957AFF1" box="[792,881,599,650]" gridcol="4" gridrow="5" pageId="13" pageNumber="136">7 (6/7– 12/13)</td>
<td id="76895DF41147006140DFFD8169C5AFF1" box="[905,995,599,650]" gridcol="5" gridrow="5" pageId="13" pageNumber="136">10, 11 oeso</td>
<td id="76895DF41147006140ADFD816E7FAFF1" box="[1019,1113,599,650]" gridcol="6" gridrow="5" pageId="13" pageNumber="136">irregular</td>
</tr>
<tr id="355834881147006143EFFD4D6E7FAFB5" box="[185,1113,667,718]" gridrow="6" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFD4D6BA8AFB5" box="[185,398,667,718]" gridcol="0" gridrow="6" pageId="13" pageNumber="136">
<emphasis id="B91CEAD81147FF9343EFFD4A6B09AFCB" box="[185,303,667,689]" italics="true" pageId="13" pageNumber="136">Oc. vallorus</emphasis>
(Bal- dasseroni, 1920)
</th>
<td id="76895DF41147006142FFFD4D6804AFB5" box="[425,546,667,718]" gridcol="1" gridrow="6" pageId="13" pageNumber="136">30–39</td>
<td id="76895DF411470061416FFD4D68B9AFB5" box="[569,671,667,718]" gridcol="2" gridrow="6" pageId="13" pageNumber="136">29, 30–39</td>
<td id="76895DF41147006141E9FD4D68D3AFB5" box="[703,757,667,718]" gridcol="3" gridrow="6" pageId="13" pageNumber="136">9–12</td>
<td id="76895DF411470061404EFD4D6957AFB5" box="[792,881,667,718]" gridcol="4" gridrow="6" pageId="13" pageNumber="136">6 (6/7– 11/12)</td>
<td id="76895DF41147006140DFFD4D69C5AFB5" box="[905,995,667,718]" gridcol="5" gridrow="6" pageId="13" pageNumber="136">10, 11 perioeso</td>
<td id="76895DF41147006140ADFD4D6E7FAFB5" box="[1019,1113,667,718]" gridcol="6" gridrow="6" pageId="13" pageNumber="136">irregular</td>
</tr>
<tr id="355834881147006143EFFD096E7FAE69" box="[185,1113,735,786]" gridrow="7" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFD096BA8AE69" box="[185,398,735,786]" gridcol="0" gridrow="7" pageId="13" pageNumber="136">
<emphasis id="B91CEAD81147FF9343EFFD096B02AF8E" box="[185,292,735,757]" italics="true" pageId="13" pageNumber="136">Oc. mimus</emphasis>
(Rosa, 1889)
</th>
<td id="76895DF41147006142FFFD096804AE69" box="[425,546,735,786]" gridcol="1" gridrow="7" pageId="13" pageNumber="136">28, 29–40, 41</td>
<td id="76895DF411470061416FFD0968B9AE69" box="[569,671,735,786]" gridcol="2" gridrow="7" pageId="13" pageNumber="136">28, 29–40, 41</td>
<td id="76895DF41147006141E9FD0968D3AE69" box="[703,757,735,786]" gridcol="3" gridrow="7" pageId="13" pageNumber="136">9–12</td>
<td id="76895DF411470061404EFD096957AE69" box="[792,881,735,786]" gridcol="4" gridrow="7" pageId="13" pageNumber="136">7 (6/7– 12/13)</td>
<td id="76895DF41147006140DFFD0969C5AE69" box="[905,995,735,786]" gridcol="5" gridrow="7" pageId="13" pageNumber="136">10, 11 perioeso</td>
<td id="76895DF41147006140ADFD096E7FAE69" box="[1019,1113,735,786]" gridcol="6" gridrow="7" pageId="13" pageNumber="136">irregular</td>
</tr>
<tr id="355834881147006143EFFCF56E7FAE2D" box="[185,1113,803,854]" gridrow="8" pageId="13" pageNumber="136">
<th id="76895DF41147006143EFFCF56BA8AE2D" box="[185,398,803,854]" gridcol="0" gridrow="8" pageId="13" pageNumber="136">
<emphasis id="B91CEAD81147FF9343EFFCF56AFEAE2E" italics="true" pageId="13" pageNumber="136">Oc. pseudocomplana- tus</emphasis>
(Omodeo, 1962)
</th>
<td id="76895DF41147006142FFFCF56804AE2D" box="[425,546,803,854]" gridcol="1" gridrow="8" pageId="13" pageNumber="136">1/n 29, 30–37, ½38</td>
<td id="76895DF411470061416FFCF568B9AE2D" box="[569,671,803,854]" gridcol="2" gridrow="8" pageId="13" pageNumber="136">1/n 29, 30–39, 40</td>
<td id="76895DF41147006141E9FCF568D3AE2D" box="[703,757,803,854]" gridcol="3" gridrow="8" pageId="13" pageNumber="136">9–12</td>
<td id="76895DF411470061404EFCF56957AE2D" box="[792,881,803,854]" gridcol="4" gridrow="8" pageId="13" pageNumber="136">7 (6/7– 12/13)</td>
<td id="76895DF41147006140DFFCF569C5AE2D" box="[905,995,803,854]" gridcol="5" gridrow="8" pageId="13" pageNumber="136">10, 11 perioeso</td>
<td id="76895DF41147006140ADFCF56E7FAE2D" box="[1019,1113,803,854]" gridcol="6" gridrow="8" pageId="13" pageNumber="136">aligned in b</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD736CA1147FF9043BCFBD7686FAD82" blockId="13.[173,1127,908,1589]" lastBlockId="14.[136,1089,189,250]" lastPageId="14" lastPageNumber="137" pageId="13" pageNumber="136">
Remarks. Morphologically the new species is most close to
<taxonomicName id="4C684D491147FF9340B1FBD76B01A947" authority="Pop, 1989" authorityName="Pop" authorityYear="1989" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="136" phylum="Annelida" rank="species" species="aporus">
<emphasis id="B91CEAD81147FF9340B1FBD76E40A961" box="[999,1126,1025,1051]" italics="true" pageId="13" pageNumber="136">Oc. aporus</emphasis>
Pop, 1989
</taxonomicName>
(cl: ½ 29, 30–40 and t: 30–40), but differs from it with the number of vesicles (3
<emphasis id="B91CEAD81147FF934202FB936B4BA925" box="[340,365,1093,1118]" italics="true" pageId="13" pageNumber="136">vs</emphasis>
. 4 pairs), spermathecae (7
<emphasis id="B91CEAD81147FF9341ECFB9368F5A925" box="[698,723,1093,1118]" italics="true" pageId="13" pageNumber="136">vs</emphasis>
. 6 pairs) and the
<typeStatus id="54D388681147FF9340FAFB9269C5A925" box="[940,995,1092,1118]" pageId="13" pageNumber="136">type</typeStatus>
of the tes- tic sacs (oesophageal
<emphasis id="B91CEAD81147FF934294FBB06BFDA904" box="[450,475,1126,1151]" italics="true" pageId="13" pageNumber="136">vs</emphasis>
. perioesophageal).
<taxonomicName id="4C684D491147FF934185FBB06954A904" box="[723,882,1125,1152]" class="Clitellata" family="Lumbricidae" genus="Octodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="13" pageNumber="136" phylum="Annelida" rank="species" species="banaticus" status="sp. nov.">
<emphasis id="B91CEAD81147FF934185FBB06954A904" box="[723,882,1125,1152]" italics="true" pageId="13" pageNumber="136">Oc. banaticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57A31147FF93402BFBB069E5A9FB" box="[893,963,1126,1152]" pageId="13" pageNumber="136" rank="species">sp. n.</taxonomicNameLabel>
is similar to
<emphasis id="B91CEAD81147FF9343F8FB5E6B40A9DA" box="[174,358,1159,1186]" italics="true" pageId="13" pageNumber="136">Oc. hemiandrus</emphasis>
(Cognetti, 1901) with the lack of vesicles in segment 11, the number of spermathecae and the oesophageal testis sacs, but differs with the clitellar organs (cl: 28–36, 37, t: 28–38, 39). It is close to
<emphasis id="B91CEAD81147FF93403EFB1D6E36A99F" box="[872,1040,1226,1253]" italics="true" pageId="13" pageNumber="136">Oc. velebiticus</emphasis>
<bibRefCitation id="EFF94B3B1147FF93474FFB1C6AC0A87D" author="MRSIC, N." pageId="13" pageNumber="136" refId="ref11082" refString="MRSIC, N. (1991): Monograph on earthworms (Lumbricidae) of the Balkans I - II. - Slovenska Akademija Znanosti in Umetnosti, Ljubljana, 757 pp." type="book" year="1991">Mršić, 1991</bibRefCitation>
with its clitellum and tubercles (cl: ½ 29–39, t: 30–39), but differs with the number of vesicles (3
<emphasis id="B91CEAD81147FF9342B0FAD86BD9A85C" box="[486,511,1294,1319]" italics="true" pageId="13" pageNumber="136">vs</emphasis>
. 4 pairs). The tubercles of
<emphasis id="B91CEAD81147FF934014FAD86E26A85C" box="[834,1024,1293,1320]" italics="true" pageId="13" pageNumber="136">Oc. complanatus</emphasis>
(Dugès, 1828) are quite similar (30–39, 40) but its clitellum is shorter (28, 29–37) and has 4 pairs of vesicles.
<emphasis id="B91CEAD81147FF934290FA876876A810" box="[454,592,1361,1387]" italics="true" pageId="13" pageNumber="136">Oc. vallorus</emphasis>
(Baldasseroni, 1920) (cl: 30–39, t: 29, 30–39) and
<emphasis id="B91CEAD81147FF9343B3FAA56B42A8F7" box="[229,356,1395,1421]" italics="true" pageId="13" pageNumber="136">Oc. mimus</emphasis>
(Rosa, 1889) (cl: 28, 29–40, 41 t: 28, 29–40, 41) are similar with their clitellar organs, but differ with the number of spermathecae (6 and 6–7 pairs), the number of vesicles (4 pairs) and the periesophageal testis sacs.
<emphasis id="B91CEAD81147FF934768FA606BADA88A" italics="true" pageId="13" pageNumber="136">Oc. pseudocomplanatus</emphasis>
(Omodeo, 1962) is very similar to the new species with its tubercles (1/n 29, 30–39, 40) and the aligned nephridiopores, but differs with its shorter clitellum (1/n 29, 30–37, ½38), reddish colour, the perioesophageal testis sacs and the four pairs of vesicles. The distinguishing characters of the similar species are shown in
<tableCitation id="C6EA03711144FF9042BDFF096863AD82" box="[491,581,223,249]" captionStart="Table 4" captionStartId="13.[241,302,193,214]" captionText="Table 4. Distinguishing characters of the species similar to Oc. banaticus sp. n." pageId="14" pageNumber="137">Table 4</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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