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<document id="3429C433ADD177A59CA6FAC2CB1D42D8" ID-DOI="10.24199/j.mmv.2004.61.11" ID-ISSN="1447-2554" ID-Zenodo-Dep="12207880" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718988272896" checkinUser="felipe" docAuthor="Bock, Philip E. &amp; Cook, Patricia L." docDate="2004" docId="03C18788102EFFC067064D2AFCB2F9CD" docLanguage="en" docName="MemMusVictoria.61.2.135-182.pdf" docOrigin="Memoirs of Museum Victoria 61 (2)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-61-issue-2-2004/pages-135-182/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Trochosodon anomalus Bock &amp; Cook, 2004, sp. nov." docType="treatment" docVersion="2" lastPageNumber="170" masterDocId="FFF8FFF0100CFFE364374B1DFFDFFF91" masterDocTitle="A review of Australian Conescharellinidae (Bryozoa: Cheilostomata)" masterLastPageNumber="182" masterPageNumber="135" pageNumber="169" updateTime="1718989335086" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="67C6A8FB8B16969A952737FE8DA1CDDF">A review of Australian Conescharellinidae (Bryozoa: Cheilostomata)</mods:title>
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<mods:namePart id="27EC822334D7BD3B135435BD86F1468A">Bock, Philip E.</mods:namePart>
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<mods:namePart id="EF3C7A673231650F218EF0283ACAA395">Cook, Patricia L.</mods:namePart>
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<mods:title id="F6532E19A87D71523B4D39457ACEC492">Memoirs of Museum Victoria</mods:title>
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<mods:date id="2CFBFA252DAA9AE4F994DF8FCF2498CB">2004</mods:date>
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<mods:number id="226259B1B6840A611976F8D3ED404816">2004-12-31</mods:number>
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<treatment id="03C18788102EFFC067064D2AFCB2F9CD" LSID="urn:lsid:plazi:treatment:03C18788102EFFC067064D2AFCB2F9CD" httpUri="http://treatment.plazi.org/id/03C18788102EFFC067064D2AFCB2F9CD" lastPageId="35" lastPageNumber="170" pageId="34" pageNumber="169">
<subSubSection id="C3726515102EFFC167064D2AFBA1F9DC" box="[817,1150,1591,1613]" pageId="34" pageNumber="169" type="nomenclature">
<paragraph id="8BD7369E102EFFC167064D2AFBA1F9DC" blockId="34.[817,1150,1591,1613]" box="[817,1150,1591,1613]" pageId="34" pageNumber="169">
<heading id="D09F81F2102EFFC167064D2AFBA1F9DC" box="[817,1150,1591,1613]" fontSize="9" level="2" pageId="34" pageNumber="169" reason="5">
<taxonomicName id="4C684D1D102EFFC167064D2AFBF7F9DC" ID-CoL="597QD" authorityName="Bock &amp; Cook" authorityYear="2004" box="[817,1064,1591,1613]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="34" pageNumber="169" phylum="Bryozoa" rank="species" species="anomalus" status="sp. nov.">
<emphasis id="B91CEA8C102EFFC167064D2AFBF7F9DC" bold="true" box="[817,1064,1591,1613]" italics="true" pageId="34" pageNumber="169">Trochosodon anomalus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57F7102EFFC160184D2AFBA1F9DC" box="[1071,1150,1591,1613]" pageId="34" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3726515102EFFC067064D79FCB2F9CD" lastPageId="35" lastPageNumber="170" pageId="34" pageNumber="169" type="description">
<paragraph id="8BD7369E102EFFC167064D79FC15F9EB" blockId="34.[817,970,1636,1658]" box="[817,970,1636,1658]" pageId="34" pageNumber="169">
<figureCitation id="13532A1B102EFFC167064D79FC15F9EB" box="[817,970,1636,1658]" captionStart="Figure 24" captionStartId="36.[139,198,1275,1295]" captionTargetBox="[137,1460,221,1257]" captionTargetId="figure-283@36.[137,1460,221,1257]" captionTargetPageId="36" captionText="Figure 24. Trochosodon anomalus sp.nov. A–B, NMV F99018, holotype. A, lateral view of colony, direction of growth arrowed, scale = 500 _m. B, antapical view showing developing zooid, scale = 500 µm. C–D, BMNH 2003.11.27.2. C, lateral view of colony, growth direction arrowed, scale = 500 µm. D, antapical view, scale = 500 µm. E–F, NMV F99018, holotype. E, adapical region showing root pores and avicularia, scale = 200 µm. F, detail of orifices and avicularia, scale = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.12207935" httpUri="https://zenodo.org/record/12207935/files/figure.png" pageId="34" pageNumber="169">Figures 24A–F</figureCitation>
</paragraph>
<paragraph id="8BD7369E102EFFC167064D93FB49F933" blockId="34.[817,1457,1678,1805]" box="[817,1174,1678,1698]" pageId="34" pageNumber="169">
<materialsCitation id="3B003CC3102EFFC167064D93FB49F933" box="[817,1174,1678,1698]" collectionCode="NMV" pageId="34" pageNumber="169" specimenCode="F99018" specimenCount="1" typeStatus="holotype">
<emphasis id="B91CEA8C102EFFC167064D93FC5BF933" box="[817,900,1678,1698]" italics="true" pageId="34" pageNumber="169">
<typeStatus id="54D3883C102EFFC167064D93FC5BF933" box="[817,900,1678,1698]" pageId="34" pageNumber="169" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectionCode id="ED79AE5B102EFFC167A74D93FC1AF933" box="[912,965,1678,1698]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="34" pageNumber="169" type="Museum">NMV</collectionCode>
<specimenCode id="DBCE9EE5102EFFC167FC4D93FBCFF933" box="[971,1040,1678,1698]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="34" pageNumber="169" type="Museum">F99018</specimenCode>
stn SLOPE-7.
</materialsCitation>
</paragraph>
<paragraph id="8BD7369E102EFFC1677B4DB4FAF0F92C" blockId="34.[817,1457,1678,1805]" box="[844,1327,1705,1725]" pageId="34" pageNumber="169">
<materialsCitation id="3B003CC3102EFFC1677B4DB4FAF4F92C" box="[844,1323,1705,1725]" collectionCode="NMV" pageId="34" pageNumber="169" specimenCode="F99019" specimenCount="1" typeStatus="paratype">
<emphasis id="B91CEA8C102EFFC1677B4DB4FC79F92C" box="[844,934,1705,1725]" italics="true" pageId="34" pageNumber="169">
<typeStatus id="54D3883C102EFFC1677B4DB4FC79F92C" box="[844,934,1705,1725]" pageId="34" pageNumber="169" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectionCode id="ED79AE5B102EFFC167864DB4FC39F92C" box="[945,998,1705,1725]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="34" pageNumber="169" type="Museum">NMV</collectionCode>
<specimenCode id="DBCE9EE5102EFFC167DB4DB4FBEBF92C" box="[1004,1076,1705,1725]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="34" pageNumber="169" type="Museum">F99019</specimenCode>
, stn SLOPE-7 (2 colonies)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD7369E102EFFC1677B4DDEFC07F89C" blockId="34.[817,1457,1678,1805]" pageId="34" pageNumber="169">
<emphasis id="B91CEA8C102EFFC1677B4DDEFC2FF946" box="[844,1008,1731,1751]" italics="true" pageId="34" pageNumber="169">Other specimens.</emphasis>
BMNH 1976.1.6.2,
<emphasis id="B91CEA8C102EFFC1608E4DDEFAC0F946" box="[1209,1311,1731,1751]" italics="true" pageId="34" pageNumber="169">Challenger</emphasis>
stn 185, Cape York,
<collectingCountry id="F37F760E102EFFC1675E4DC3FC61F963" box="[873,958,1758,1778]" name="Australia" pageId="34" pageNumber="169">Australia</collectingCountry>
,
<quantity id="4C909B7B102EFFC167FE4DC3FBDBF963" box="[969,1028,1758,1778]" metricMagnitude="2" metricUnit="m" metricValue="2.79" pageId="34" pageNumber="169" unit="m" value="279.0">279 m</quantity>
(26 colonies); NMV F101974,
<emphasis id="B91CEA8C102EFFC1611D4DC3FA4FF963" box="[1322,1424,1758,1778]" italics="true" pageId="34" pageNumber="169">Challenger</emphasis>
stn 185 (10 colonies).
</paragraph>
<caption id="DF176616102EFFC164BC4DE8FD47F8E5" ID-DOI="http://doi.org/10.5281/zenodo.12207929" ID-Zenodo-Dep="12207929" httpUri="https://zenodo.org/record/12207929/files/figure.png" pageId="34" pageNumber="169" startId="34.[139,198,1781,1801]" targetBox="[171,762,221,1759]" targetPageId="34" targetType="figure">
<paragraph id="8BD7369E102EFFC164BC4DE8FD47F8E5" blockId="34.[139,777,1781,1908]" pageId="34" pageNumber="169">
Figure 21.
<taxonomicName id="4C684D1D102EFFC164C44DE8FE0CF898" authorityName="Bock &amp; Cook" authorityYear="2004" box="[243,467,1781,1801]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="34" pageNumber="169" phylum="Bryozoa" rank="species" species="diommatus" status="sp. nov.">
<emphasis id="B91CEA8C102EFFC164C44DE8FE0CF898" box="[243,467,1781,1801]" italics="true" pageId="34" pageNumber="169">Trochosodon diommatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57F7102EFFC165EB4DE8FDFAF898" box="[476,549,1781,1801]" pageId="34" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
A. NMV F99012, holotype. Detail of orifices and adapical pores, scale = 200 µm. B, NMV F99013, paratype, antapical view of marginal peristomes, showing paired avicularia, scale =200 µm. C. NMV F99014, paratype, detail of peristome, orifice, and lunate root pores, scale = 200 µm.
</paragraph>
</caption>
<paragraph id="8BD7369E102EFFC167064C38FA7AF8E2" blockId="34.[817,1457,1827,1907]" pageId="34" pageNumber="169">
<emphasis id="B91CEA8C102EFFC167064C38FC4FF8A8" box="[817,912,1829,1849]" italics="true" pageId="34" pageNumber="169">Etymology</emphasis>
. From
<emphasis id="B91CEA8C102EFFC167EE4C3EFBE5F8A8" box="[985,1082,1827,1849]" italics="true" pageId="34" pageNumber="169">anomalos</emphasis>
(Gr.) – irregular, inconsistent, deviating, with reference to the combination of character states found in several genera, that are uniquely possessed by this species.
</paragraph>
<caption id="DF176616102FFFC064B449D9FE4CFC9C" ID-DOI="http://doi.org/10.5281/zenodo.12207931" ID-Zenodo-Dep="12207931" httpUri="https://zenodo.org/record/12207931/files/figure.png" pageId="35" pageNumber="170" startId="35.[131,190,708,728]" targetBox="[129,1452,221,687]" targetPageId="35" targetType="figure">
<paragraph id="8BD7369E102FFFC064B449D9FE4CFC9C" blockId="35.[131,1449,708,781]" pageId="35" pageNumber="170">
Figure 22. A–D
<taxonomicName id="4C684D1D102FFFC0652C49D9FE1AFD49" authorityName="Bock &amp; Cook" authorityYear="2004" box="[283,453,708,728]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="species" species="aster" status="sp. nov.">
<emphasis id="B91CEA8C102FFFC0652C49D9FE1AFD49" box="[283,453,708,728]" italics="true" pageId="35" pageNumber="170">Trochosodon aster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F57F7102FFFC065FB49D9FDCAFD49" box="[460,533,708,728]" pageId="35" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
A, NMV F99015, paratype, young colony showing alternating triad structure; root pores and avicularia arrowed, scale = 200 µm. B, NMV 99016, paratype, older colony, scale = 500µm. C, NMV P99017, paratype, colony showing adapical calcification, scale = 500 µm.
</paragraph>
</caption>
<paragraph id="8BD7369E102FFFC0671E4820FBDAFBE9" blockId="35.[809,1450,829,1203]" pageId="35" pageNumber="170">
<emphasis id="B91CEA8C102FFFC0671E4820FBE6FCC2" box="[809,1081,829,851]" italics="true" pageId="35" pageNumber="170">Diagnosis and description.</emphasis>
Colonies very small, less than
<quantity id="4C909B7B102FFFC061584820FA77FCC2" box="[1391,1448,829,851]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="170" unit="mm" value="2.0">2 mm</quantity>
in height and diameter but appearing to be higher than wide. Calcification mamillate. Zooids arranged in alternating whorls, each of 3 zooids, appearing to be in radial series; peristomes elongated and prominent. Primary orifice with a shallow, rounded sinus and paired condyles, adapical pore symmetrically placed on the edge of the peristome. Avicularia paired, lateral, widely separated from the orifices, alternating in radial series; rostra rounded, bar without a ligula. Adapical region with avicularia and small rounded pores; antapical region with a few avicularia only.
</paragraph>
<paragraph id="8BD7369E102FFFC0677E4F9DFB25FB22" blockId="35.[809,1450,829,1203]" pageId="35" pageNumber="170">
Colony diameter
<quantity id="4C909B7B102FFFC067CA4F9DFBA3FB07" box="[1021,1148,1152,1174]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="35" pageNumber="170" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 mm</quantity>
, height
<quantity id="4C909B7B102FFFC060E74F9DFA90FB07" box="[1232,1359,1152,1174]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="35" pageNumber="170" unit="mm" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 mm</quantity>
, number of whorls 2–4, number of zooids per whorl 3.
</paragraph>
<paragraph id="8BD7369E102FFFC0671E4FD4FC6DFA5B" blockId="35.[809,1449,1225,1628]" pageId="35" pageNumber="170">
<emphasis id="B91CEA8C102FFFC0671E4FD4FC5DFB4E" box="[809,898,1225,1247]" italics="true" pageId="35" pageNumber="170">Remarks</emphasis>
. Specimens of
<taxonomicName id="4C684D1D102FFFC0602C4FD4FB49FB4E" authorityName="Bock &amp; Cook" authorityYear="2004" box="[1051,1174,1225,1247]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="species" species="anomalus">
<emphasis id="B91CEA8C102FFFC0602C4FD4FB49FB4E" box="[1051,1174,1225,1247]" italics="true" pageId="35" pageNumber="170">T. anomalus</emphasis>
</taxonomicName>
are of great interest as they include characteristics “typical” of both
<taxonomicName id="4C684D1D102FFFC060C64FFBFAABFB6D" authorityName="Canu and Bassler" authorityYear="1927" box="[1265,1396,1254,1276]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="genus">
<emphasis id="B91CEA8C102FFFC060C64FFBFAABFB6D" box="[1265,1396,1254,1276]" italics="true" pageId="35" pageNumber="170">Trochosodon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D1D102FFFC0671E4E19FC10FA8B" authorityName="d'Orbigny" authorityYear="1852" box="[809,975,1284,1306]" class="Gymnolaemata" family="Conescharellinidae" genus="Conescharellina" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="genus">
<emphasis id="B91CEA8C102FFFC0671E4E19FC10FA8B" box="[809,975,1284,1306]" italics="true" pageId="35" pageNumber="170">Conescharellina</emphasis>
</taxonomicName>
; in some features they even resemble species of
<taxonomicName id="4C684D1D102FFFC067714E3CFC7AFAA6" authorityName="Reuss" authorityYear="1867" box="[838,933,1313,1335]" class="Gymnolaemata" family="Batoporidae" genus="Batopora" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="genus">
<emphasis id="B91CEA8C102FFFC067714E3CFC7AFAA6" box="[838,933,1313,1335]" italics="true" pageId="35" pageNumber="170">Batopora</emphasis>
</taxonomicName>
, from which they are readily distinguished by the presence of an adapical pore. The arrangement of the radial series of avicularia suggests assignment to
<taxonomicName id="4C684D1D102FFFC060EB4E41FA5DFAE3" authorityName="d'Orbigny" authorityYear="1852" box="[1244,1410,1372,1394]" class="Gymnolaemata" family="Conescharellinidae" genus="Conescharellina" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="genus">
<emphasis id="B91CEA8C102FFFC060EB4E41FA5DFAE3" box="[1244,1410,1372,1394]" italics="true" pageId="35" pageNumber="170">Conescharellina</emphasis>
</taxonomicName>
but the lack of basal cancelli and the presence of prominent, tubular peristomes allow its inclusion here with other species of
<taxonomicName id="4C684D1D102FFFC0671E4EA9FC73FA5B" authorityName="Canu and Bassler" authorityYear="1927" box="[809,940,1460,1482]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="genus">
<emphasis id="B91CEA8C102FFFC0671E4EA9FC73FA5B" box="[809,940,1460,1482]" italics="true" pageId="35" pageNumber="170">Trochosodon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD7369E102FFFC0677E4ECCFCB2F9CD" blockId="35.[809,1449,1225,1628]" pageId="35" pageNumber="170">
The numerous colonies from Cape York indicate that the distribution of this tiny species extends from
<collectingRegion id="49ACF87C102FFFC061244EF3FA54F995" box="[1299,1419,1518,1540]" country="Australia" name="Queensland" pageId="35" pageNumber="170">Queensland</collectingRegion>
to
<collectingRegion id="49ACF87C102FFFC0671E4D11FC3DF9B3" box="[809,994,1548,1570]" country="Australia" name="New South Wales" pageId="35" pageNumber="170">New South Wales</collectingRegion>
, from lower shelf to slope depths. The similarities between
<taxonomicName id="4C684D1D102FFFC067F94D34FB96F9AE" authorityName="Bock &amp; Cook" authorityYear="2004" box="[974,1097,1577,1599]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="species" species="anomalus">
<emphasis id="B91CEA8C102FFFC067F94D34FB96F9AE" box="[974,1097,1577,1599]" italics="true" pageId="35" pageNumber="170">T. anomalus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C684D1D102FFFC0604E4D34FAC2F9AE" authorityName="Bock &amp; Cook" authorityYear="2004" box="[1145,1309,1577,1599]" class="Gymnolaemata" family="Conescharellinidae" genus="Trochosodon" kingdom="Animalia" order="Cheilostomatida" pageId="35" pageNumber="170" phylum="Bryozoa" rank="species" species="asymmetricus">
<emphasis id="B91CEA8C102FFFC0604E4D34FAC2F9AE" box="[1145,1309,1577,1599]" italics="true" pageId="35" pageNumber="170">T. asymmetricus</emphasis>
</taxonomicName>
are described above.
</paragraph>
</subSubSection>
</treatment>
</document>
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