treatments-xml/data/03/C1/25/03C1251D7E75FFC9038DFF3BFAC12E0D.xml
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<document id="72FFA9C2CD851718B7D8A5F4479BF673" ID-DOI="10.11606/1807-0205/2020.60.26" ID-ISSN="1807-0205" ID-Zenodo-Dep="4614913" ID-ZooBank="0EF2A463-5ED2-420F-AF80-70660A774232" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616008014004" checkinUser="carolina" docAuthor="Bravo, Freddy, Vilarinho, Naiara &amp; Araújo, Maíra Xavier" docDate="2020" docId="03C1251D7E75FFC9038DFF3BFAC12E0D" docLanguage="en" docName="PapAvZool.60.e20206026.pdf" docOrigin="Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 60" docSource="http://dx.doi.org/10.11606/1807-0205/2020.60.26" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Tonnoira igrapiunensis Bravo &amp; Vilarinho &amp; Araújo 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2" masterDocId="FFF85D657E74FFC8036EFFE1FFF22F7E" masterDocTitle="New species and new records of Tonnoira (Diptera: Psychodidae) from Brazil" masterLastPageNumber="5" masterPageNumber="5" pageNumber="2" updateTime="1698889211807" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
<mods:mods id="88C68493B731129D6D58E9F45ED35BF4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E40A6FDD74ADFC33183A8FDBC6B19A6E">
<mods:title id="C78723E96A7888B800E900865DED6525">New species and new records of Tonnoira (Diptera: Psychodidae) from Brazil</mods:title>
</mods:titleInfo>
<mods:name id="76D27AAD766EF3C9E43EEB3BD57AE5FD" type="personal">
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<mods:namePart id="1583B54DB5962FC664B2BB2B36FA8F84">Bravo, Freddy</mods:namePart>
</mods:name>
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<mods:role id="36AF44CC4EDF0AD0E93A259CA215EBD2">
<mods:roleTerm id="F98D5E475C418D9724CDC7A2EC49D66F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8DABD80F30ECB2317B742C0E1FF40348">Vilarinho, Naiara</mods:namePart>
</mods:name>
<mods:name id="C694FC7500C3B3D520580F040558D20C" type="personal">
<mods:role id="B27BA352CBA7ADF5B0305742AB3E94D3">
<mods:roleTerm id="8CB799D689D66F5986EA43B0FA0AE011">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC2D77966AFEB725EBA04DC64B9B9C31">Araújo, Maíra Xavier</mods:namePart>
</mods:name>
<mods:typeOfResource id="8B3DB0A5E9A9A6587F088AAB094B7A95">text</mods:typeOfResource>
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<mods:title id="4F6A6C7F8D19CBCC3D560B16A1831B26">Papéis Avulsos de Zoologia</mods:title>
</mods:titleInfo>
<mods:part id="81B728D761970695C2960671367693B8">
<mods:date id="752C9A5390AAD1D2780F5378EB5CE8ED">2020</mods:date>
<mods:detail id="2F706D2ED403682291A5233946B1AC5B" type="series">
<mods:title id="DE79615F0DECCAE81F866DD6C05434E5">Pap. Avulsos Zool., S. Paulo</mods:title>
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<mods:number id="0D244168CE2D7C30156541B06F6C3F71">2020-06-08</mods:number>
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<mods:detail id="0B3576168B83FEE1B2327F55BBC90A9D" type="volume">
<mods:number id="7CB44085B92975608A1FE6D2A500E1B8">60</mods:number>
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<mods:url id="6C44E6EE1CB203D9DD25BB4F8E8EEDE4">http://dx.doi.org/10.11606/1807-0205/2020.60.26</mods:url>
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<mods:classification id="E8B9D7B37B48296103125DEDF91EDAE0">journal article</mods:classification>
<mods:identifier id="6A819889F49CC66C93821DEECD048EF9" type="DOI">10.11606/1807-0205/2020.60.26</mods:identifier>
<mods:identifier id="B1A8F70AE728392B1D423FBBF987CCA6" type="ISSN">1807-0205</mods:identifier>
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<mods:identifier id="4DF8952BF3797AEE0087A35E10BD7764" type="ZooBank">0EF2A463-5ED2-420F-AF80-70660A774232</mods:identifier>
</mods:mods>
<treatment id="03C1251D7E75FFC9038DFF3BFAC12E0D" LSID="urn:lsid:plazi:treatment:03C1251D7E75FFC9038DFF3BFAC12E0D" httpUri="http://treatment.plazi.org/id/03C1251D7E75FFC9038DFF3BFAC12E0D" lastPageNumber="2" pageId="1" pageNumber="2">
<subSubSection id="C372C7807E75FFC9038DFF3BFDB82F8D" box="[227,586,218,243]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BD7940B7E75FFC9038DFF3BFDB82F8D" blockId="1.[227,586,218,275]" box="[227,586,218,243]" pageId="1" pageNumber="2">
<heading id="D09F23677E75FFC9038DFF3BFDB82F8D" box="[227,586,218,243]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="5">
<emphasis id="B91C48197E75FFC9038DFF3BFDB82F8D" bold="true" box="[227,586,218,243]" pageId="1" pageNumber="2">
<taxonomicName id="4C68EF887E75FFC9038DFF3BFE192F8D" ID-CoL="9HZ4Z" authority="Bravo &amp; Vilarinho &amp; Araújo, 2020" authorityName="Bravo &amp; Vilarinho &amp; Araújo" authorityYear="2020" box="[227,491,218,243]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="igrapiunensis" status="sp. nov.">
<emphasis id="B91C48197E75FFC9038DFF3BFE192F8D" bold="true" box="[227,491,218,243]" italics="true" pageId="1" pageNumber="2">Tonnoira igrapiunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22FF5627E75FFC9029EFF3BFDB82F8D" box="[496,586,218,243]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C372C7807E75FFC90230FF1BFE3D2E6D" box="[350,463,250,275]" pageId="1" pageNumber="2" type="description">
<paragraph id="8BD7940B7E75FFC90230FF1BFE3D2E6D" blockId="1.[227,586,218,275]" box="[350,463,250,275]" pageId="1" pageNumber="2">
<figureCitation id="1353888E7E75FFC90230FF1BFE3D2E6D" box="[350,463,250,275]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">
<emphasis id="B91C48197E75FFC90230FF1BFE3D2E6D" bold="true" box="[350,463,250,275]" pageId="1" pageNumber="2">Figs. 212</emphasis>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C372C7807E75FFC90330FEDBFE4E2D0D" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BD7940B7E75FFC90330FEDBFE4E2D0D" blockId="1.[94,719,314,627]" pageId="1" pageNumber="2">
<emphasis id="B91C48197E75FFC90330FEDBFEEE2E2D" bold="true" box="[94,284,314,339]" pageId="1" pageNumber="2">Type material:</emphasis>
<materialsCitation id="3B009E567E75FFC9025BFEDBFE2C2ECD" collectingDate="2012-05-28" collectingDateMax="2012-07-12" collectingDateMin="2012-05-28" collectionCode="MZFS" collectorName="Aragao, M. &amp; Menezes, E. &amp; Mota, E. &amp; Andena, S" country="Brazil" county="Igrapiuna" location="Vila" municipality="Reserva Ecologica Michelin" pageId="1" pageNumber="2" specimenCode="MZFS #56225" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="B91C48197E75FFC9025BFEDBFE582E2D" bold="true" box="[309,426,314,339]" pageId="1" pageNumber="2">
<typeStatus id="54D32AA97E75FFC9025BFEDBFE542E2D" box="[309,422,314,339]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
:
</emphasis>
male,
<collectingCountry id="F37FD49B7E75FFC90172FEDBFD9C2E2D" box="[540,622,314,339]" name="Brazil" pageId="1" pageNumber="2">BRAZIL</collectingCountry>
,
<collectingRegion id="49AC5AE97E75FFC901E5FEDBFD392E2D" box="[651,715,314,339]" country="Brazil" name="Bahia" pageId="1" pageNumber="2">Bahia</collectingRegion>
,
<collectingCounty id="62B6EC877E75FFC90330FEBBFF392E0D" box="[94,203,346,371]" pageId="1" pageNumber="2">Igrapiúna</collectingCounty>
,
<collectingMunicipality id="6BB30E717E75FFC903B5FEBBFDE42E0D" box="[219,534,346,371]" pageId="1" pageNumber="2">Reserva Ecológica Michelin</collectingMunicipality>
,
<location id="8EB7C2D07E75FFC9014AFEBBFDBE2E0D" LSID="urn:lsid:plazi:treatment:03C1251D7E75FFC9038DFF3BFAC12E0D:8EB7C2D07E75FFC9014AFEBBFDBE2E0D" box="[548,588,346,371]" country="Brazil" county="Igrapiuna" municipality="Reserva Ecologica Michelin" name="Vila" pageId="1" pageNumber="2" stateProvince="Bahia">Vila</location>
5,
<date id="FFD6B2CB7E75FFC90119FEBBFF452EED" pageId="1" pageNumber="2" value="2012-05-28" valueMax="2012-07-12" valueMin="2012-05-28">
<collectingDate id="EF924B237E75FFC90119FEBBFF452EED" pageId="1" pageNumber="2" value="2012-05-28" valueMax="2012-07-12" valueMin="2012-05-28">28.v-12. vii.2012</collectingDate>
</date>
,
<location id="8EB7C2D07E75FFC903AAFE9BFEEA2EED" LSID="urn:lsid:plazi:treatment:03C1251D7E75FFC9038DFF3BFAC12E0D:8EB7C2D07E75FFC903AAFE9BFEEA2EED" box="[196,280,378,403]" country="Brazil" county="Igrapiuna" municipality="Reserva Ecologica Michelin" name="Malaise" pageId="1" pageNumber="2" stateProvince="Bahia">Malaise</location>
[trap].
<collectorName id="269DF1DD7E75FFC90201FE9BFE192EED" box="[367,491,378,403]" pageId="1" pageNumber="2">Aragão, M.</collectorName>
;
<collectorName id="269DF1DD7E75FFC90294FE9BFD732EED" box="[506,641,378,403]" pageId="1" pageNumber="2">Menezes, E.</collectorName>
;
<collectorName id="269DF1DD7E75FFC901E1FE9BFF812ECD" pageId="1" pageNumber="2">Mota, E.</collectorName>
;
<collectorName id="269DF1DD7E75FFC903EEFE7BFF042ECD" box="[128,246,410,435]" pageId="1" pageNumber="2">Andena, S</collectorName>
, leg. (
<specimenCode id="DBCE3C707E75FFC90256FE7BFE2A2ECD" box="[312,472,410,435]" collectionCode="MZFS" pageId="1" pageNumber="2">MZFS #56225</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B009E567E75FFC90284FE7BFE872E8D" collectingDate="2012-05-28" collectingDateMax="2012-07-12" collectingDateMin="2012-05-28" collectionCode="MZFS" collectorName="Aragao, M. &amp; Menezes, E. &amp; Mota, E. &amp; Andena, S" country="Brazil" county="Igrapiuna" location="Vila" municipality="Reserva Ecologica Michelin" pageId="1" pageNumber="2" specimenCode="MZFS #56227, 56229, 56230, 56231, 56232, 56233" specimenCount="6" specimenCount-male="6" stateProvince="Bahia" typeStatus="paratype">
<emphasis id="B91C48197E75FFC90284FE7BFD952ECD" bold="true" box="[490,615,410,435]" pageId="1" pageNumber="2">
<typeStatus id="54D32AA97E75FFC90284FE7BFD912ECD" box="[490,611,410,435]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D6E5F827E75FFC9011FFE7BFD3E2ECD" box="[625,716,410,435]" pageId="1" pageNumber="2" type="male">6 males</specimenCount>
, same data as holotype (
<specimenCode id="DBCE3C707E75FFC90200FE5BFDFE2EAD" box="[366,524,442,467]" collectionCode="MZFS" pageId="1" pageNumber="2">MZFS #56227</specimenCode>
, #
<specimenCode id="DBCE3C707E75FFC9014BFE5BFD9E2EAD" box="[549,620,442,467]" collectionCode="MZFS" pageId="1" pageNumber="2">56229</specimenCode>
, #
<specimenCode id="DBCE3C707E75FFC901EBFE5BFD3E2EAD" box="[645,716,442,467]" collectionCode="MZFS" pageId="1" pageNumber="2">56230</specimenCode>
, #
<specimenCode id="DBCE3C707E75FFC90303FE3BFF412E8D" box="[109,179,474,499]" collectionCode="MZFS" pageId="1" pageNumber="2">56231</specimenCode>
, #
<specimenCode id="DBCE3C707E75FFC903A4FE3BFEE32E8D" box="[202,273,474,499]" collectionCode="MZFS" pageId="1" pageNumber="2">56232</specimenCode>
, #
<specimenCode id="DBCE3C707E75FFC90249FE3BFE9D2E8D" box="[295,367,474,499]" collectionCode="MZFS" pageId="1" pageNumber="2">56233</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B009E567E75FFC90210FE3BFE1D2D4D" collectingDate="2012-10-27" collectingDateMax="2012-10-28" collectingDateMin="2012-10-27" collectionCode="MZFS" collectorName="Aragao, M. &amp; Menezes, E. &amp; Mota, E." country="Brazil" county="Igrapiuna" location="Vila" municipality="Reserva Ecologica Michelin" pageId="1" pageNumber="2" specimenCode="MZFS #56226, 56234" specimenCount="2" specimenCount-male="2" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="9D6E5F827E75FFC90210FE3BFE242E8D" box="[382,470,474,499]" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
, same locality as holotype,
<date id="FFD6B2CB7E75FFC903F3FE1BFEC62D6D" box="[157,308,506,531]" pageId="1" pageNumber="2" value="2012-10-27" valueMax="2012-10-28" valueMin="2012-10-27">
<collectingDate id="EF924B237E75FFC903F3FE1BFEC62D6D" box="[157,308,506,531]" pageId="1" pageNumber="2" value="2012-10-27" valueMax="2012-10-28" valueMin="2012-10-27">27-28.x. 2012</collectingDate>
</date>
, light [trap],
<collectorName id="269DF1DD7E75FFC902ADFE1BFDCE2D6D" box="[451,572,506,531]" pageId="1" pageNumber="2">Aragão, M.</collectorName>
,
<collectorName id="269DF1DD7E75FFC90129FE1BFD392D6D" box="[583,715,506,531]" pageId="1" pageNumber="2">Menezes, E.</collectorName>
,
<collectorName id="269DF1DD7E75FFC9030EFDFBFF4B2D4D" box="[96,185,538,563]" pageId="1" pageNumber="2">Mota, E.</collectorName>
leg. (
<specimenCode id="DBCE3C707E75FFC9039EFDFBFE792D4D" box="[240,395,538,563]" collectionCode="MZFS" pageId="1" pageNumber="2">MZFS #56226</specimenCode>
, #
<specimenCode id="DBCE3C707E75FFC902CFFDFBFE1B2D4D" box="[417,489,538,563]" collectionCode="MZFS" pageId="1" pageNumber="2">56234</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B009E567E75FFC90296FDFBFE4A2D0D" collectingDate="2012-10-28" collectingDateMax="2012-11-18" collectingDateMin="2012-10-28" collectionCode="MZFS" collectorName="Aragao, M. &amp; Menezes, E." country="Brazil" county="Igrapiuna" location="Vila" municipality="Reserva Ecologica Michelin" pageId="1" pageNumber="2" specimenCode="MZFS #56228" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="9D6E5F827E75FFC90296FDFBFDB62D4D" box="[504,580,538,563]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
, same locality as holotype,
<date id="FFD6B2CB7E75FFC9026AFDDBFE5E2D2D" box="[260,428,570,595]" pageId="1" pageNumber="2" value="2012-10-28" valueMax="2012-11-18" valueMin="2012-10-28">
<collectingDate id="EF924B237E75FFC9026AFDDBFE5E2D2D" box="[260,428,570,595]" pageId="1" pageNumber="2" value="2012-10-28" valueMax="2012-11-18" valueMin="2012-10-28">28.x-18.xi.2012</collectingDate>
</date>
, Malaise [trap],
<collectorName id="269DF1DD7E75FFC9013AFDDBFD382D2D" box="[596,714,570,595]" pageId="1" pageNumber="2">Aragão, M.</collectorName>
,
<collectorName id="269DF1DD7E75FFC90330FDBBFF2C2D0D" box="[94,222,602,627]" pageId="1" pageNumber="2">Menezes, E.</collectorName>
leg. (
<specimenCode id="DBCE3C707E75FFC90278FDBBFE402D0D" box="[278,434,602,627]" collectionCode="MZFS" pageId="1" pageNumber="2">MZFS #56228</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C372C7807E75FFC90330FD7BFEC629ED" pageId="1" pageNumber="2" type="description">
<paragraph id="8BD7940B7E75FFC90330FD7BFD5D292D" blockId="1.[93,721,666,1619]" pageId="1" pageNumber="2">
<emphasis id="B91C48197E75FFC90330FD7BFE8C2DCD" bold="true" box="[94,382,666,691]" pageId="1" pageNumber="2">Description: Male: Head:</emphasis>
slightly wider than long; short vertex; eye bridge with four facet rows; eyes separated by almost one facet diameter; interocular suture inverted Y-shaped (
<figureCitation id="1353888E7E75FFC903BCFD1BFEE32C6D" box="[210,273,762,787]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 2</figureCitation>
).
<emphasis id="B91C48197E75FFC9024DFD1BFE7D2C6D" bold="true" box="[291,399,762,787]" pageId="1" pageNumber="2">Antenna:</emphasis>
scape wider apically than at base, 2.0 times the length of the pedicel (
<figureCitation id="1353888E7E75FFC90140FCFBFD9E2C4D" box="[558,620,794,819]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
); pedicel spherical, with two groups of three small sensorial setae both in front of and behind the proximal margin (
<figureCitation id="1353888E7E75FFC901E6FCBBFD342C0D" box="[648,710,858,883]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 6</figureCitation>
); flagellum with 14 flagellomeres, all cylindrical (
<figureCitation id="1353888E7E75FFC90108FC9BFD372CED" box="[614,709,890,915]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Figs. 3-5</figureCitation>
); ascoids not observed,probably lost;14
<superScript id="7C1D39437E75FFC90290FC78FDFE2CD6" attach="left" box="[510,524,921,936]" fontSize="6" pageId="1" pageNumber="2">th</superScript>
flagellomere with long apiculus, 0.3 times the length of the flagellomere (
<figureCitation id="1353888E7E75FFC9030BFC3BFF572C8D" box="[101,165,986,1011]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
).
<emphasis id="B91C48197E75FFC903D6FC3BFEA42C8D" bold="true" box="[184,342,986,1011]" pageId="1" pageNumber="2">
Wing (
<figureCitation id="1353888E7E75FFC9026AFC3BFEBA2C8D" box="[260,328,986,1011]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 8</figureCitation>
):
</emphasis>
Sc short not reaching C; R₂₊₃ not connected to R₄; R₅ ending slightly beyond the wing apex; radial fork basal to medial fork.
<emphasis id="B91C48197E75FFC902D1FBFBFDB02B4D" bold="true" box="[447,578,1050,1075]" pageId="1" pageNumber="2">Terminalia:</emphasis>
hypandrium bent backwards, narrower laterally (
<figureCitation id="1353888E7E75FFC9029BFBDBFDC42B2D" box="[501,566,1082,1107]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 9</figureCitation>
); gonocoxites 1.2 times the length of gonostyli, with setae only dorsally (
<figureCitation id="1353888E7E75FFC9030BFB9BFF272BED" box="[101,213,1146,1171]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Figs. 9, 11</figureCitation>
); gonostyli bifurcate; lateral branch of gonostylus longer than mesal branch (
<figureCitation id="1353888E7E75FFC902ABFB7BFDC52BCD" box="[453,567,1178,1203]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Figs. 9, 10</figureCitation>
), divergent in lateral view (
<figureCitation id="1353888E7E75FFC9039CFB5BFEB02BAD" box="[242,322,1210,1235]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 10</figureCitation>
); aedeagus bipartite, asymmetrical, short shaft 0.8 times the length of the long shaft; short shaft with two sclerites fused basally, dorsal sclerite platelike, shorter than the narrow ventral sclerite, both covered by an apical membrane (
<figureCitation id="1353888E7E75FFC90203FADBFE592A2D" box="[365,427,1338,1363]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 9</figureCitation>
); parameres asymmetrical, long shaft with curved apex and short shaft plate-like with rounded apex (
<figureCitation id="1353888E7E75FFC90222FA9BFE362AED" box="[332,452,1402,1427]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Figs. 9, 11</figureCitation>
); gonocoxal apodemes fused medially and articulated with parameres (
<figureCitation id="1353888E7E75FFC90119FA7BFD302ACD" box="[631,706,1434,1459]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 11</figureCitation>
); epandrium wider than long, bare ventrally, with one basal aperture (
<figureCitation id="1353888E7E75FFC90386FA3BFEC72A8D" box="[232,309,1498,1523]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 12</figureCitation>
); hypoproct longer than epandrium, with apex tapered; epiproct shorter than hypoproct, with rounded apex; hypoproct and epiproct both with apical micropilosity (
<figureCitation id="1353888E7E75FFC90396F9DBFEB0292D" box="[248,322,1594,1619]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 12</figureCitation>
); cerci with two tenacula (
<figureCitation id="1353888E7E75FFC90134F9DBFD56292D" box="[602,676,1594,1619]" captionStart="Figures 2-12" captionStartId="2.[170,238,1978,2003]" captionTargetBox="[173,1490,210,1918]" captionTargetId="figure-1@2.[170,1493,204,1924]" captionTargetPageId="2" captionText="Figures 2-12. Tonnoira igrapiunensis sp. nov., adult male. (2) Head in frontal view (paratype 56227). (3) scape, pedicel and first flagellomeres (paratype 56227). (4)Flagellomeres 4,5and 6 (paratype 56227).(5)Apical flagellomeres andapiculus (paratype56226).(6) pedicelwiththe marginalsensorysetae(paratype 56227). (7) Palpus(paratype56227).(8)wing.(9)Hypandrium,gonocoxites,gonostylus,ejaculatoryapodeme,aedeagus and parameres (holotype).(10)gonostylus,lateral view (paratype 56227).(11) gonocoxites,gonocoxal apodemes fused,parameres and ejaculatory apodeme (holotype 56225).(12) Epandrium,cerci and hypoproct (paratype 56227).Abbreviations:am = apical membrane;lsae = long shaft of aedeagus;pr = paramere;ssae = small shaft of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4614919" httpUri="https://zenodo.org/record/4614919/files/figure.png" pageId="1" pageNumber="2">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BD7940B7E75FFC90330F99BFEC629ED" blockId="1.[94,308,1658,1683]" box="[94,308,1658,1683]" pageId="1" pageNumber="2">
<emphasis id="B91C48197E75FFC90330F99BFF4929ED" bold="true" box="[94,187,1658,1683]" pageId="1" pageNumber="2">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C372C7807E75FFC90330F95BFDBE272D" pageId="1" pageNumber="2" type="discussion">
<paragraph id="8BD7940B7E75FFC90330F95BFDBE272D" blockId="1.[92,719,1722,2131]" pageId="1" pageNumber="2">
<emphasis id="B91C48197E75FFC90330F95BFF3829AD" bold="true" box="[94,202,1722,1747]" pageId="1" pageNumber="2">Remarks:</emphasis>
The new species can be distinguished from oth- er species of
<taxonomicName id="4C68EF887E75FFC9039AF93BFEA3298D" authorityName="Enderlein" authorityYear="1937" box="[244,337,1754,1779]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B91C48197E75FFC9039AF93BFEA3298D" box="[244,337,1754,1779]" italics="true" pageId="1" pageNumber="2">Tonnoira</emphasis>
</taxonomicName>
by the eye bridge with four facet rows, separated by 0.5 facet diameter; R₅ ending slightly posterior to apex of wing; gonostyli bifurcated, dorsal arm shorter than ventral arm;long arm of gonostyli shorter than length of gonocoxites; cerci with two tenacula; aedeagus bipartite, asymmetrical, short shaft 0.8 times the length of the long shaft; parameres asymmetrical, longer shaft with apex curved.The gonostylus is similar to that of
<taxonomicName id="4C68EF887E75FFC90135F85BFF19288D" authority="Santos, 2014" authorityName="Santos" authorityYear="2014" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="andradei">
<emphasis id="B91C48197E75FFC90135F85BFD3F28AD" box="[603,717,1978,2003]" italics="true" pageId="1" pageNumber="2">T.andradei</emphasis>
<bibRefCitation id="EFF9E9FA7E75FFC90330F83BFF19288D" author="Santos, C. B. &amp; Curler, G. R." box="[94,235,2010,2035]" pageId="1" pageNumber="2" pagination="463 - 470" refId="ref4267" refString="Santos, C. B. &amp; Curler, G. R. 2014. Four new species of Tonnoira Enderlein (Diptera: Psychodidae, Psychodinae) from the Brazilian Atlantic forest. Zootaxa, 3760: 463 - 470." type="journal article" year="2014">Santos, 2014</bibRefCitation>
</taxonomicName>
, but differs, mainly, by the bifid aedeagus in the new species and with only one shaft in
<emphasis id="B91C48197E75FFC90144F81BFD50276D" box="[554,674,2042,2067]" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="4C68EF887E75FFC90144F81BFD6C276D" authorityName="Santos" authorityYear="2014" box="[554,670,2042,2067]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="andradei">T.andradei</taxonomicName>
.
</emphasis>
The ejaculatory apodeme and hypoproct in the new species is longer than the same structures in
<emphasis id="B91C48197E75FFC902BAF7DBFDBE272D" box="[468,588,2106,2131]" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="4C68EF887E75FFC902BAF7DBFDBA272D" authorityName="Santos" authorityYear="2014" box="[468,584,2106,2131]" class="Insecta" family="Psychodidae" genus="Tonnoira" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="andradei">T.andradei</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C372C7807E75FFC90074FF3BFBE92E4D" pageId="1" pageNumber="2" type="etymology">
<paragraph id="8BD7940B7E75FFC90074FF3BFBE92E4D" blockId="1.[794,1418,218,307]" pageId="1" pageNumber="2">
<emphasis id="B91C48197E75FFC90074FF3BFC532F8D" bold="true" box="[794,929,218,243]" pageId="1" pageNumber="2">Etymology:</emphasis>
The species name is a reference to the municipality of Igrapiúna where the specimens of the new species were collected.
</paragraph>
</subSubSection>
<subSubSection id="C372C7807E75FFC90074FEBBFAC12E0D" box="[794,1331,346,371]" pageId="1" pageNumber="2" type="distribution">
<paragraph id="8BD7940B7E75FFC90074FEBBFAC12E0D" blockId="1.[794,1331,346,371]" box="[794,1331,346,371]" pageId="1" pageNumber="2">
<emphasis id="B91C48197E75FFC90074FEBBFC432E0D" bold="true" box="[794,945,346,371]" pageId="1" pageNumber="2">Distribution:</emphasis>
Known only from the
<typeStatus id="54D32AA97E75FFC907C8FEBBFB2A2E0D" box="[1190,1240,346,371]" pageId="1" pageNumber="2">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>