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<document id="9ECCDA1F99D4ED282CD2E0ACA5EC9C7E" ID-DOI="10.11646/zootaxa.4751.1.13" ID-GBIF-Dataset="bc38c18d-80ad-463b-8c2c-a0ce1f0ade1f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3711750" ID-ZooBank="4D98A9A3-B510-49C3-A50C-CFBD7325EBD6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584349035548" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="03BFF97FFFE78272B4E0269DFC9CF86C" docLanguage="en" docName="zootaxa.4751.1.13.pdf" docOrigin="Zootaxa 4751 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Siamites langkatensis Jałoszyński 2020, sp. n." docType="treatment" docVersion="5" lastPageNumber="188" masterDocId="FF868107FFE68273B4772408FFFDFF94" masterDocTitle="Four new species of the Oriental genus Siamites Franz (Coleoptera, Staphylinidae Scydmaeninae)" masterLastPageNumber="195" masterPageNumber="187" pageNumber="188" updateTime="1698728428309" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="6EF42075579BCF16C27E21AD077CBDB6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F43FC97D96E271F788F40FD9909351FC">
<mods:title id="B0BC7FB15936E401347DF0196049E84D">Four new species of the Oriental genus Siamites Franz (Coleoptera, Staphylinidae Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="E6E254A05CD0B4B6A3942A10288929D8" type="personal">
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<mods:namePart id="27CF7E87C9ECB817E1397BB88174857A">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="6AED8F3C68E525F9E0267F7212395C0A">Zootaxa</mods:title>
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<mods:part id="F13FA7FF7922F2328B60272F2E19019A">
<mods:date id="6C61D381A79E75630E8AC9116630983A">2020</mods:date>
<mods:detail id="C3011F928516E667CEF1FEFA5A5DFC4D" type="pubDate">
<mods:number id="36C0C2F15A09F42213FFDBE66F316A26">2020-03-16</mods:number>
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<mods:number id="BA223B01E0B7427CB39583CFE8418AA6">4751</mods:number>
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<mods:number id="08DF36048220441DD8D2C46560B2F706">1</mods:number>
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<mods:classification id="7AB101ADF4D101EFE388E8B22E69477C">journal article</mods:classification>
<mods:identifier id="8A0113B2A9B319EC9FDD9CC56A298EBF" type="DOI">10.11646/zootaxa.4751.1.13</mods:identifier>
<mods:identifier id="10FC2132C6D905593AEFF978A1E03D79" type="GBIF-Dataset">bc38c18d-80ad-463b-8c2c-a0ce1f0ade1f</mods:identifier>
<mods:identifier id="E0FF24FDAC254AD135A1A5F802B2A6A2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="595DA6A1E79AF491AE6FA7C487F2038F" type="Zenodo-Dep">3711750</mods:identifier>
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<treatment id="03BFF97FFFE78272B4E0269DFC9CF86C" ID-DOI="http://doi.org/10.5281/zenodo.3718010" ID-GBIF-Taxon="162575683" ID-Zenodo-Dep="3718010" LSID="urn:lsid:plazi:treatment:03BFF97FFFE78272B4E0269DFC9CF86C" httpUri="http://treatment.plazi.org/id/03BFF97FFFE78272B4E0269DFC9CF86C" lastPageNumber="188" pageId="1" pageNumber="188">
<subSubSection id="C30C1BE2FFE78272B4E0269DFE1BFD3B" box="[151,486,661,688]" pageId="1" pageNumber="188" type="nomenclature">
<paragraph id="8BA94869FFE78272B4E0269DFE1BFD3B" blockId="1.[151,486,595,724]" box="[151,486,661,688]" pageId="1" pageNumber="188">
<heading id="D0E1FF05FFE78272B4E0269DFE1BFD3B" bold="true" box="[151,486,661,688]" fontSize="11" level="1" pageId="1" pageNumber="188" reason="1">
<emphasis id="B962947BFFE78272B4E0269DFE1BFD3B" bold="true" box="[151,486,661,688]" pageId="1" pageNumber="188">
<taxonomicName id="4C1633EAFFE78272B4E0269DFE60FD24" ID-CoL="8Y9KX" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,413,661,688]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="langkatensis" status="sp. nov.">
<emphasis id="B962947BFFE78272B4E0269DFE60FD24" bold="true" box="[151,413,661,688]" italics="true" pageId="1" pageNumber="188">Siamites langkatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2512900FFE78272B5D3269DFE1BFD3B" box="[420,486,661,687]" pageId="1" pageNumber="188" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C1BE2FFE78272B4E026B1FEF6FD40" box="[151,267,697,724]" pageId="1" pageNumber="188" type="description">
<paragraph id="8BA94869FFE78272B4E026B1FEF6FD40" blockId="1.[151,486,595,724]" box="[151,267,697,724]" pageId="1" pageNumber="188">
(
<figureCitation id="132D54ECFFE78272B4E826B1FEFFFD40" box="[159,258,697,724]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1571,1596]" captionTargetBox="[214,1361,179,1543]" captionTargetId="figure@2.[214,1374,179,1543]" captionTargetPageId="2" captionText="FIGURES 1–6. Siamites langkatensis sp. n., dorsal habitus (1); head in anterodorsal view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views." figureDoi="http://doi.org/10.5281/zenodo.3711752" httpUri="https://zenodo.org/record/3711752/files/figure.png" pageId="1" pageNumber="188">Figs 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C1BE2FFE78272B4E026F4FE9AFCC9" pageId="1" pageNumber="188" type="materials_examined">
<paragraph id="8BA94869FFE78272B4E026F4FE9AFCC9" blockId="1.[151,1437,762,2040]" pageId="1" pageNumber="188">
<emphasis id="B962947BFFE78272B4E026F4FEB8FC81" bold="true" box="[151,325,762,790]" pageId="1" pageNumber="188">Type material.</emphasis>
<materialsCitation id="3B7E4234FFE78272B53B26F4FE9AFCC9" ID-GBIF-Occurrence="2576149572" collectionCode="MHNG" country="Indonesia" determinerName="P. Jaloszynski" location="Sumatra" pageId="1" pageNumber="188" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="B962947BFFE78272B53B26F4FD0CFC81" bold="true" box="[332,753,762,790]" pageId="1" pageNumber="188">
<typeStatus id="54ADF6CBFFE78272B53B26F4FE40FC82" box="[332,445,764,790]" pageId="1" pageNumber="188" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F30108F9FFE78272B5BE26F2FD9BFC81" box="[457,614,762,789]" name="Indonesia" pageId="1" pageNumber="188">INDONESIA</collectingCountry>
(
<collectingRegion id="49D2868BFFE78272B60226F2FD19FC81" box="[629,740,762,789]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="188">Sumatra</collectingRegion>
):
</emphasis>
<specimenCount id="9D1083E0FFE78272B68F26F4FCF2FC82" box="[760,783,764,790]" pageId="1" pageNumber="188" type="male">♂</specimenCount>
, two labels: “
<collectingRegion id="49D2868BFFE78272B7D226F3FBD0FC81" box="[933,1069,763,789]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="188">SUMATRA</collectingRegion>
/ res. Langkat / bord Bohorok-5. / J.Robert-VII.84” [white, printed]; “
<taxonomicName id="4C1633EAFFE78272B6432717FD51FCAD" authorityName="Franz" authorityYear="1989" box="[564,684,799,825]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B962947BFFE78272B6432717FD51FCAD" box="[564,684,799,825]" italics="true" pageId="1" pageNumber="188">SIAMITES</emphasis>
</taxonomicName>
/
<taxonomicName id="4C1633EAFFE78272B6B62717FCB0FCAD" authorityName="Jałoszyński" authorityYear="2020" box="[705,845,799,825]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="langkatensis">
<emphasis id="B962947BFFE78272B6B62717FCB0FCAD" box="[705,845,799,825]" italics="true" pageId="1" pageNumber="188">langkatensis</emphasis>
</taxonomicName>
m. / det.
<determinerName id="7E75A4ADFFE78272B7C02728FBABFCAD" box="[951,1110,799,826]" pageId="1" pageNumber="188">P. Jałoszyński</determinerName>
, 2020 /
<typeStatus id="54ADF6CBFFE78272B0D92717FAA5FCAD" box="[1198,1368,799,825]" pageId="1" pageNumber="188" type="holotype">HOLOTYPUS</typeStatus>
” [red, printed] (
<collectionCode id="ED07D0ACFFE78272B488274BFEA6FCC9" box="[255,347,835,861]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="188" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30C1BE2FFE78272B4B0276EFBA8FC30" pageId="1" pageNumber="188" type="diagnosis">
<paragraph id="8BA94869FFE78272B4B0276EFBA8FC30" blockId="1.[151,1437,762,2040]" pageId="1" pageNumber="188">
<emphasis id="B962947BFFE78272B4B0276EFEBFFC15" bold="true" box="[199,322,870,897]" pageId="1" pageNumber="188">Diagnosis.</emphasis>
Male macrophthalmous; head with six separate tufts of bristles evenly distributed along posterior margin of vertex; aedeagus broadest near middle, with deeply bifurcate apical region.
</paragraph>
</subSubSection>
<subSubSection id="C30C1BE2FFE78272B4B027A5FE63F96A" pageId="1" pageNumber="188" type="description">
<paragraph id="8BA94869FFE78272B4B027A5FDBAFC78" blockId="1.[151,1437,762,2040]" pageId="1" pageNumber="188">
<emphasis id="B962947BFFE78272B4B027A5FEA4FC5C" bold="true" box="[199,345,941,968]" pageId="1" pageNumber="188">Description.</emphasis>
Body of male (
<figureCitation id="132D54ECFFE78272B67327A6FDBEFC5C" box="[516,579,942,968]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1571,1596]" captionTargetBox="[214,1361,179,1543]" captionTargetId="figure@2.[214,1374,179,1543]" captionTargetPageId="2" captionText="FIGURES 1–6. Siamites langkatensis sp. n., dorsal habitus (1); head in anterodorsal view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views." figureDoi="http://doi.org/10.5281/zenodo.3711752" httpUri="https://zenodo.org/record/3711752/files/figure.png" pageId="1" pageNumber="188">Fig. 1</figureCitation>
) moderately convex and moderately slender, uniformly light brown, covered with yellowish vestiture. BL
<quantity id="4CEEE58CFFE78272B5AB27DAFDBEFC78" box="[476,579,978,1004]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="188" unit="mm" value="1.1">1.10 mm</quantity>
.
</paragraph>
<paragraph id="8BA94869FFE78272B4B027FDFBFAFAB9" blockId="1.[151,1437,762,2040]" pageId="1" pageNumber="188">
Head (
<figureCitation id="132D54ECFFE78272B56427FDFEA5FB84" box="[275,344,1013,1040]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1571,1596]" captionTargetBox="[214,1361,179,1543]" captionTargetId="figure@2.[214,1374,179,1543]" captionTargetPageId="2" captionText="FIGURES 1–6. Siamites langkatensis sp. n., dorsal habitus (1); head in anterodorsal view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views." figureDoi="http://doi.org/10.5281/zenodo.3711752" httpUri="https://zenodo.org/record/3711752/files/figure.png" pageId="1" pageNumber="188">Fig. 2</figureCitation>
) broadest at conspicuously large, strongly convex and coarsely faceted eyes, HL
<quantity id="4CEEE58CFFE78272B08E27FDFA9CFB84" box="[1273,1377,1013,1040]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="188" unit="mm" value="0.15">0.15 mm</quantity>
, HW
<quantity id="4CEEE58CFFE78272B4E02011FF03FBA0" box="[151,254,1049,1076]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="188" unit="mm" value="0.23">0.23 mm</quantity>
; tempora in dorsal view nearly as long as eyes; vertex flattened at middle, strongly transverse; frons confluent with vertex, with shallow median impression between weakly elevated supraantennal tubercles. Punctures on vertex obscured by bristles, those on frons fine, inconspicuous. Vestiture of head dorsum (
<figureCitation id="132D54ECFFE78272B0F82069FB2EFBEF" box="[1167,1235,1121,1147]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1571,1596]" captionTargetBox="[214,1361,179,1543]" captionTargetId="figure@2.[214,1374,179,1543]" captionTargetPageId="2" captionText="FIGURES 1–6. Siamites langkatensis sp. n., dorsal habitus (1); head in anterodorsal view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views." figureDoi="http://doi.org/10.5281/zenodo.3711752" httpUri="https://zenodo.org/record/3711752/files/figure.png" pageId="1" pageNumber="188">Fig. 2</figureCitation>
) composed of six separate tufts of dense bristles forming a transverse row on vertex; long and moderately thick bristles directed posterodorsad and situated slightly in front of posterior tufts; and sparse, suberect setae on frons. Antennae short, with moderately broadened, indistinctly delimited tetramerous club, AnL
<quantity id="4CEEE58CFFE78272B7E220C3FC01FB72" box="[917,1020,1227,1254]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="188" unit="mm" value="0.45">0.45 mm</quantity>
; scape and pedicel strongly elongate; antennomeres III–VI each indistinctly elongate, VII about as long as broad, VIII–X each distinctly transverse, XI distinctly shorter than IX–X combined, narrower than X, indistinctly elongate.
</paragraph>
<paragraph id="8BA94869FFE78272B4B02130FDE8FA28" blockId="1.[151,1437,762,2040]" pageId="1" pageNumber="188">
Pronotum subconical with broad anterior margin, broadest near posterior third; PL
<quantity id="4CEEE58CFFE78272B010213FFB30FAC6" box="[1127,1229,1335,1362]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="188" unit="mm" value="0.3">0.30 mm</quantity>
, PW
<quantity id="4CEEE58CFFE78272B17E213FFA8DFAC6" box="[1289,1392,1335,1362]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="188" unit="mm" value="0.28">0.28 mm</quantity>
. All margins weakly and evenly rounded; lateral carinae distinct and almost as long as half length of pronotum; base with distinct transverse groove deepened at each end. Punctures on pronotum fine and sparse, unremarkable; setae sparse, suberect, moderately long.
</paragraph>
<paragraph id="8BA94869FFE78272B4B021CEFB98F9DF" blockId="1.[151,1437,762,2040]" pageId="1" pageNumber="188">
Elytra slightly flattened, together oval, broadest distinctly in front of middle; EL
<quantity id="4CEEE58CFFE78272B01321CDFB31FA74" box="[1124,1228,1477,1504]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="188" unit="mm" value="0.65">0.65 mm</quantity>
, EW
<quantity id="4CEEE58CFFE78272B17A21CEFA88FA74" box="[1293,1397,1478,1504]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="188" unit="mm" value="0.48">0.48 mm</quantity>
, EI 1.37; humeral calli weakly marked, each with indistinct, rounded humeral carina not exceeding 1/4 of elytral length; basal impressions shallow and short; apices of elytra rounded together. Punctures on elytra more distinct and denser than those on pronotum but superficial and unremarkable; setae sparse, short, suberect.
</paragraph>
<paragraph id="8BA94869FFE78272B4B0225DFE05F9FB" blockId="1.[151,1437,762,2040]" box="[199,504,1621,1647]" pageId="1" pageNumber="188">Wings long and functional.</paragraph>
<paragraph id="8BA94869FFE78272B4B02271FB02F94F" blockId="1.[151,1437,762,2040]" pageId="1" pageNumber="188">
Aedeagus (
<figureCitation id="132D54ECFFE78272B5332271FE5AF907" box="[324,423,1656,1683]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1571,1596]" captionTargetBox="[214,1361,179,1543]" captionTargetId="figure@2.[214,1374,179,1543]" captionTargetPageId="2" captionText="FIGURES 1–6. Siamites langkatensis sp. n., dorsal habitus (1); head in anterodorsal view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views." figureDoi="http://doi.org/10.5281/zenodo.3711752" httpUri="https://zenodo.org/record/3711752/files/figure.png" pageId="1" pageNumber="188">Figs 3–6</figureCitation>
) moderately stout; AeL
<quantity id="4CEEE58CFFE78272B6C32270FCE7F907" box="[692,794,1656,1683]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="188" unit="mm" value="0.15">0.15 mm</quantity>
; median lobe fusiform, broadest near middle and strongly narrowing toward base and apex, apical region bifurcate, with two pairs of long lateral projections; endophallic structures lightly sclerotized; parameres short and moderately broad, each with two tiny apical setae.
</paragraph>
<paragraph id="8BA94869FFE78272B4B022ECFE63F96A" blockId="1.[151,1437,762,2040]" box="[199,414,1764,1790]" pageId="1" pageNumber="188">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30C1BE2FFE78272B4B0230FFD5CF8B6" box="[199,673,1799,1826]" pageId="1" pageNumber="188" type="distribution">
<paragraph id="8BA94869FFE78272B4B0230FFD5CF8B6" blockId="1.[151,1437,762,2040]" box="[199,673,1799,1826]" pageId="1" pageNumber="188">
<emphasis id="B962947BFFE78272B4B0230FFE9CF8B6" bold="true" box="[199,353,1799,1826]" pageId="1" pageNumber="188">Distribution.</emphasis>
Northern
<collectingRegion id="49D2868BFFE78272B5A32300FDCCF8B6" box="[468,561,1800,1826]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="188">Sumatra</collectingRegion>
(
<figureCitation id="132D54ECFFE78272B637230FFD68F8B6" box="[576,661,1799,1826]" captionStart="FIGURE 25" captionStartId="7.[151,250,2004,2029]" captionTargetBox="[151,1436,1173,1979]" captionTargetId="figure@7.[151,1436,1173,1979]" captionTargetPageId="7" captionText="FIGURE 25. Distribution of nominal Siamites species." figureDoi="http://doi.org/10.5281/zenodo.3711762" httpUri="https://zenodo.org/record/3711762/files/figure.png" pageId="1" pageNumber="188">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30C1BE2FFE78272B4B02324FCDFF8D1" box="[199,802,1835,1862]" pageId="1" pageNumber="188" type="etymology">
<paragraph id="8BA94869FFE78272B4B02324FCDFF8D1" blockId="1.[151,1437,762,2040]" box="[199,802,1835,1862]" pageId="1" pageNumber="188">
<emphasis id="B962947BFFE78272B4B02324FEB3F8D2" bold="true" box="[199,334,1836,1862]" pageId="1" pageNumber="188">Etymology.</emphasis>
After the Indonesian regency of Langkat.
</paragraph>
</subSubSection>
<subSubSection id="C30C1BE2FFE78272B4B02347FC9CF86C" pageId="1" pageNumber="188" type="discussion">
<paragraph id="8BA94869FFE78272B4B02347FC9CF86C" blockId="1.[151,1437,762,2040]" pageId="1" pageNumber="188">
<emphasis id="B962947BFFE78272B4B02347FEC6F8FD" bold="true" box="[199,315,1871,1897]" pageId="1" pageNumber="188">Remarks.</emphasis>
Adults of
<taxonomicName id="4C1633EAFFE78272B5CC2347FDE7F8FD" authorityName="Franz" authorityYear="1989" box="[443,538,1871,1897]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B962947BFFE78272B5CC2347FDE7F8FD" box="[443,538,1871,1897]" italics="true" pageId="1" pageNumber="188">Siamites</emphasis>
</taxonomicName>
are externally uniform; differences can be seen in proportions of body parts, distribution of tufts of bristles on the vertex, and a relative diameter of eyes. However, these differences can be indistinct, and the most reliable way to identify a species is to examine the aedeagus. The aedeagi in
<taxonomicName id="4C1633EAFFE78272B08C239FFAA7F825" authorityName="Franz" authorityYear="1989" box="[1275,1370,1943,1969]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="genus">
<emphasis id="B962947BFFE78272B08C239FFAA7F825" box="[1275,1370,1943,1969]" italics="true" pageId="1" pageNumber="188">Siamites</emphasis>
</taxonomicName>
show far greater diversity than any external adult structures.
<taxonomicName id="4C1633EAFFE78272B77F23B3FC06F840" authorityName="Jałoszyński" authorityYear="2020" box="[776,1019,1978,2005]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="188" phylum="Arthropoda" rank="species" species="langkatensis">
<emphasis id="B962947BFFE78272B77F23B3FC06F840" box="[776,1019,1978,2005]" italics="true" pageId="1" pageNumber="188">Siamites langkatensis</emphasis>
</taxonomicName>
clearly differs from all congeners in the uniquely shaped, bifurcate apical region of the median lobe.
</paragraph>
</subSubSection>
</treatment>
</document>
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