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<document id="3C675D404840BA27028BF2D36CE5699C" ID-CLB-Dataset="46146" ID-DOI="10.5281/zenodo.278958" ID-GBIF-Dataset="56fb15f0-b55b-4ce9-a9ad-d8a2f5a95029" ID-ISSN="1175-5326" ID-Zenodo-Dep="278958" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460401525037" checkinUser="plazi" docAuthor="Williams, Kevin A., Brothers, Denis J. &amp; Pitts, James P." docDate="2011" docId="03BF6217D823215206B64F41FD47FE73" docLanguage="en" docName="zt03064p068.pdf" docOrigin="Zootaxa 3064" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tobantilla krima Williams, Brothers &amp; Pitts, sp. nov." docType="treatment" docVersion="11" lastPageNumber="58" masterDocId="FF861A6FD833214306214C5EFF9CFFBC" masterDocTitle="New species of Tobantilla Casal, 1965 and a new genus and species, Gogoltilla chichikovi gen. et sp. nov., from Argentina (Hymenoptera: Mutillidae)" masterLastPageNumber="68" masterPageNumber="41" pageNumber="57" updateTime="1698299261469" updateUser="ExternalLinkService">
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<mods:title id="770932BAD5E4D0D6A237EC3B77088E6E">New species of Tobantilla Casal, 1965 and a new genus and species, Gogoltilla chichikovi gen. et sp. nov., from Argentina (Hymenoptera: Mutillidae)</mods:title>
</mods:titleInfo>
<mods:name id="FB4903936E8E0CC69CC74FE51D0BD78F" type="personal">
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<mods:namePart id="9695F4A43E224267F0403C4401923F7D">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="F7F3426E2CB0C23A1505265A27F93951">Brothers, Denis J.</mods:namePart>
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<mods:name id="0D1A5651E49920A99B3A45B02FD5C1B2" type="personal">
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<mods:namePart id="D8CC592D02AE7CF2146E1CBFA927E308">Pitts, James P.</mods:namePart>
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<mods:typeOfResource id="FA264A7A0001938742C8711FE10AC4C4">text</mods:typeOfResource>
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<mods:title id="11798E9B1DE1A7D47BC5F19DEA3991E3">Zootaxa</mods:title>
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<mods:part id="A4DC79C3E5998016E2521154C74686BC">
<mods:date id="9AD286CEF2F8302C62700F90C997940B">2011</mods:date>
<mods:detail id="5B87F3B2C2840BDC1A892DCBA8484215" type="volume">
<mods:number id="04A4B1A822DBD0AA70F26EB9794DB542">3064</mods:number>
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<treatment id="03BF6217D823215206B64F41FD47FE73" ID-DOI="http://doi.org/10.5281/zenodo.6191535" ID-GBIF-Taxon="119419968" ID-Zenodo-Dep="6191535" LSID="urn:lsid:zoobank.org:act:7B0C62B5-899F-43D8-9D6C-6C1FE84B6FF3" httpUri="http://treatment.plazi.org/id/03BF6217D823215206B64F41FD47FE73" lastPageId="17" lastPageNumber="58" pageId="16" pageNumber="57">
<subSubSection id="C30C808AD823215306B64F41FE84FCE6" pageId="16" pageNumber="57" type="nomenclature">
<paragraph id="8BA9D301D823215306B64F41FCBCFC85" blockId="16.[151,800,799,858]" box="[151,800,799,825]" pageId="16" pageNumber="57">
<heading id="D0E1646DD823215306B64F41FCBCFC85" bold="true" box="[151,800,799,825]" fontSize="11" level="1" pageId="16" pageNumber="57" reason="1">
<emphasis id="B9620F13D823215306B64F41FCBCFC85" bold="true" box="[151,800,799,825]" pageId="16" pageNumber="57">
<taxonomicName id="4C16A882D823215306B64F41FD2AFC85" authority="Williams, Brothers &amp; Pitts" authorityName="Williams, Brothers &amp; Pitts" box="[151,694,799,825]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="krima" status="sp. nov.">
<emphasis id="B9620F13D823215306B64F41FEFDFC85" bold="true" box="[151,353,799,825]" italics="true" pageId="16" pageNumber="57">Tobantilla krima</emphasis>
Williams, Brothers &amp; Pitts
</taxonomicName>
,
<taxonomicNameLabel id="A251B268D823215304E24F41FCBCFC85" box="[707,800,799,825]" pageId="16" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA9D301D823215306B64F1CFE84FCE6" blockId="16.[151,800,799,858]" box="[151,280,834,858]" pageId="16" pageNumber="57">
(
<figureCitation id="132DCF84D823215306BE4F1CFF7BFCE6" box="[159,231,834,858]" captionStart="FIGURES 1 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="16" pageNumber="57">Figs 5</figureCitation>
,
<figureCitation id="132DCF84D823215306D24F1CFE92FCE6" box="[243,270,834,858]" captionStart="FIGURES 7 14" captionStartId="6.[151,264,1742,1765]" captionTargetBox="[151,1436,193,1719]" captionTargetId="figure@6.[151,1436,193,1721]" captionTargetPageId="6" captionText="FIGURES 7 14. Tobantilla species, females. 7 - 12. Habitus, lateral view. 7. T. aleatrix sp. nov., holotype; 8. T. charrasca Casal; 9. T. frigidula (Cresson); 10. T. kolasma sp. nov., holotype; 11. T. krima sp. nov., holotype; 12. T. montonera Casal. 13, 14. Pygidium (T 6). 13. T. frigidula; 14. T. aleatrix." httpUri="https://zenodo.org/record/278960/files/figure.png" pageId="16" pageNumber="57">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD823215306B64FD4FDE9FC56" pageId="16" pageNumber="57" type="diagnosis">
<paragraph id="8BA9D301D823215306B64FD4FDE9FC56" blockId="16.[151,1436,906,2011]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306B64FD4FE8DFC1F" bold="true" box="[151,273,906,931]" pageId="16" pageNumber="57">Diagnosis.</emphasis>
FEMALE. The pronotum is clothed with black setae that contrast with the pale golden mesonotal setae (
<figureCitation id="132DCF84D823215306BE4FF0FF7FFC7A" box="[159,227,942,966]" captionStart="FIGURES 1 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="16" pageNumber="57">Fig. 5</figureCitation>
); the scutellar scale is separated from the dorsal propodeal carina; and T2 has a single pair of yellow integumental spots, on the posterior half (
<figureCitation id="132DCF84D823215304024F8CFDFBFC56" box="[547,615,978,1002]" captionStart="FIGURES 1 6" captionStartId="5.[151,264,1969,1992]" captionTargetBox="[151,1436,411,1944]" captionTargetId="figure@5.[151,1436,411,1945]" captionTargetPageId="5" captionText="FIGURES 1 6. Tobantilla species, females, habitus, dorsal view. 1. T. aleatrix sp. nov.; holotype; 2. T. charrasca Casal; 3. T. frigidula (Cresson); 4. T. kolasma sp. nov., holotype; 5. T. krima sp. nov., holotype; 6. T. montonera Casal." httpUri="https://zenodo.org/record/278959/files/figure.png" pageId="16" pageNumber="57">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD823215306E44FA8FE04F8D2" pageId="16" pageNumber="57" type="description">
<paragraph id="8BA9D301D823215306E44FA8FE6BF8F6" blockId="16.[151,1436,906,2011]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306E44FA8FECAFBB2" bold="true" box="[197,342,1014,1039]" pageId="16" pageNumber="57">Description.</emphasis>
FEMALE. Body length
<quantity id="4CEE7EE4D8232153044F4FA8FD54FBB2" box="[622,712,1014,1038]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="16" pageNumber="57" unit="mm" value="4.5">4.5 mm</quantity>
.
<emphasis id="B9620F13D823215304F54FA8FCCAFBB2" box="[724,854,1014,1038]" italics="true" pageId="16" pageNumber="57">Coloration.</emphasis>
Head and mesosoma pale brown, except metapleuron and lateral face of propodeum brown. Metasoma dark brown to black, except T46 brown and T2 having a pair of yellow circular spots in basal half. Legs, antennae and mandibles pale brown. Tibial spurs white. Head and mesosoma clothed with pale golden or silvery setae, except pronotum clothed with black setae and dorsum of head and mesosoma having a few scattered erect brown setae. T23 clothed with black setae, except lateral margins and integumental spots clothed with silver setae. T56 and S16 clothed with silvery white setae.
<emphasis id="B9620F13D8232153029548F5FB6CFB7F" box="[1204,1264,1195,1219]" italics="true" pageId="16" pageNumber="57">Head</emphasis>
. Rounded posteriorly, with lowly produced rectangular glabrous tubercles on posterolateral margin. Head width 1.0 × pronotal width. Eye almost circular. Front and vertex coarsely contiguously punctate; gena shallowly reticulate. Genal carina lowly produced, extending anteriorly to hypostomal carina. Mandible oblique, tapering, bidentate apically, unarmed ventrally. Antennal scrobe lacking dorsal carina. Antennal tubercle moderately punctate basally. Scape simply punctate. Flagellomere 1 1.2 × pedicel length; flagellomere 2 1.2 × pedicel length.
<emphasis id="B9620F13D823215302B5493EFA90FACB" box="[1172,1292,1376,1399]" italics="true" pageId="16" pageNumber="57">Mesosoma</emphasis>
. Mesosomal length 1.1 × width; pronotum 0.95 × as wide as mesothorax. Mesosomal dorsum reticulate. Humeral carina low, extending to epaulet. Lateral face of pronotum and mesopleuron micropunctate, dull, mesopleural ridge broadly and shallowly reticulate; clothed with sparse silver setae. Metapleuron dorsal to endophragmal pit and dorsal twothirds of lateral face of propodeum shagreened; ventral regions of metapleuron and lateral face of propodeum shagreened. In dorsal view, mesosoma gradually narrowed posterior to pronotum, lateral margin of mesothorax weakly sinuate anterior to propodeal spiracle. Scutellar scale highly raised, having transverse w-shaped carinae anteriorly. Dorsal and posterior propodeal faces separated by transverse carina. Dorsal propodeal face convergent posterior to spiracle, posterolateral angle rounded, posterior face virtually flat, vertical, reticulate and clothed with moderately dense silver setae on dorsal half.
<emphasis id="B9620F13D823215304BB4AFAFC93F907" box="[666,783,1700,1723]" italics="true" pageId="16" pageNumber="57">Metasoma</emphasis>
. T1 narrow and petiolate, somewhat cylindrical, 0.5 × as wide as T2. T2 1.15 × longer than wide, with maximum width situated medially. T2 densely punctate, punctation similar on integumental spots and disc of T2; S2 moderately punctate. T35 densely punctate, S25 moderately punctate. Pygidium well defined by lateral carinae, posterior margin rounded, having 8 widely separated striae, spaces between striae glabrous.
</paragraph>
<paragraph id="8BA9D301D823215306E44B09FE04F8D2" blockId="16.[151,1436,906,2011]" box="[197,408,1878,1903]" pageId="16" pageNumber="57">MALE. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30C808AD823215306E44B25FC73F80A" pageId="16" pageNumber="57" type="materials_examined">
<paragraph id="8BA9D301D823215306E44B25FC73F80A" blockId="16.[151,1436,906,2011]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306E44B25FEE8F82E" bold="true" box="[197,372,1914,1939]" pageId="16" pageNumber="57">
<typeStatus id="54AD6DA3D823215306E44B25FE9CF82F" box="[197,256,1915,1939]" pageId="16" pageNumber="57">Type</typeStatus>
material.
</emphasis>
<emphasis id="B9620F13D8232153075C4B25FE7FF82F" box="[381,483,1915,1939]" italics="true" pageId="16" pageNumber="57">
<typeStatus id="54AD6DA3D8232153075C4B25FE7FF82F" box="[381,483,1915,1939]" pageId="16" pageNumber="57" type="holotype">Holotype</typeStatus>
</emphasis>
, Ƥ,
<collectingCountry id="F3019391D823215304394B24FD2EF82E" box="[536,690,1914,1938]" name="Colombia" pageId="16" pageNumber="57">COLOMBIA</collectingCountry>
: Bolivar, Zambrano, Alrededores Hacienda Toleteme, 9°N 74°W,
<date id="FFA8F5C1D823215306B64BC0FEBEF80A" box="[151,290,1950,1974]" pageId="16" pageNumber="57" value="1994-12-13">13.XII.1994</date>
,
<emphasis id="B9620F13D8232153070C4BC0FDCBF80A" box="[301,599,1950,1975]" italics="true" pageId="16" pageNumber="57">Sobre carretera destapada</emphasis>
(on dirt road), M. Ospina (
<collectionCode id="ED074BC4D823215305A64BC0FC7FF80A" LSID="urn:lsid:biocol.org:col:34925" box="[903,995,1950,1974]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="57">AMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD823215206E44B9CFE1DFF13" lastPageId="17" lastPageNumber="58" pageId="16" pageNumber="57" type="distribution">
<paragraph id="8BA9D301D823215306E44B9CFC0FF866" blockId="16.[151,1436,906,2011]" box="[197,915,1986,2011]" pageId="16" pageNumber="57">
<emphasis id="B9620F13D823215306E44B9CFEC2F866" bold="true" box="[197,350,1986,2011]" pageId="16" pageNumber="57">Distribution.</emphasis>
This species is known from the
<typeStatus id="54AD6DA3D823215304E84B9DFD65F867" box="[713,761,1987,2011]" pageId="16" pageNumber="57">type</typeStatus>
locality only.
</paragraph>
<paragraph id="8BA9D301D822215206E44CC6FE1DFF13" blockId="17.[151,1436,151,464]" box="[197,385,151,176]" pageId="17" pageNumber="58">
<emphasis id="B9620F13D822215206E44CC6FE98FF0C" bold="true" box="[197,260,152,176]" pageId="17" pageNumber="58">Host.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD822215206E44CE2FE23FEA0" pageId="17" pageNumber="58" type="etymology">
<paragraph id="8BA9D301D822215206E44CE2FE23FEA0" blockId="17.[151,1436,151,464]" pageId="17" pageNumber="58">
<emphasis id="B9620F13D822215206E44CE2FED7FF68" bold="true" box="[197,331,188,212]" pageId="17" pageNumber="58">Etymology.</emphasis>
From the Greek
<taxonomicName id="4C16A882D8222152043A4CE2FDC1FF68" box="[539,605,188,212]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="krima">
<emphasis id="B9620F13D8222152043A4CE2FDC1FF68" box="[539,605,188,212]" italics="true" pageId="17" pageNumber="58">krima</emphasis>
</taxonomicName>
“crime”, a noun in apposition. This species, along with
<taxonomicName id="4C16A882D822215202DB4CE3FAE9FF68" box="[1274,1397,188,212]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="kolasma">
<emphasis id="B9620F13D822215202DB4CE3FAE9FF68" box="[1274,1397,188,212]" italics="true" pageId="17" pageNumber="58">T. kolasma</emphasis>
</taxonomicName>
, is named after the Russian novel Преступление и наказание “Crime and Punishment” written by Fyodor Mikhailovich Dostoevsky.
</paragraph>
</subSubSection>
<subSubSection id="C30C808AD822215206E44D76FD47FE73" pageId="17" pageNumber="58" type="discussion">
<paragraph id="8BA9D301D822215206E44D76FC22FE3B" blockId="17.[151,1436,151,464]" pageId="17" pageNumber="58">
<emphasis id="B9620F13D822215206E44D76FEA5FE83" bold="true" box="[197,313,296,320]" pageId="17" pageNumber="58">Remarks.</emphasis>
This species was collected over
<quantity id="4CEE7EE4D822215204914D79FC9BFE83" box="[688,775,295,319]" metricMagnitude="5" metricUnit="m" metricValue="8.0" pageId="17" pageNumber="58" unit="km" value="800.0">800 km</quantity>
away from the only known Colombian male,
<taxonomicName id="4C16A882D822215203374D77FA09FEFC" box="[1302,1429,296,320]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="andrikos">
<emphasis id="B9620F13D822215203374D77FA09FEFC" box="[1302,1429,296,320]" italics="true" pageId="17" pageNumber="58">T. andrikos</emphasis>
</taxonomicName>
, but was collected
<quantity id="4CEE7EE4D8222152077E4D12FE28FED8" box="[351,436,332,356]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="58" unit="km" value="400.0">400 km</quantity>
away from the Venezuelan male,
<taxonomicName id="4C16A882D8222152050C4D13FC03FED8" box="[813,927,332,356]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="xouthos">
<emphasis id="B9620F13D8222152050C4D13FC03FED8" box="[813,927,332,356]" italics="true" pageId="17" pageNumber="58">T. xouthos</emphasis>
</taxonomicName>
(
<figureCitation id="132DCF84D8222152058F4D12FC63FED8" box="[942,1023,332,356]" captionStart="FIGURE 39" captionStartId="12.[151,250,1315,1338]" captionTargetBox="[195,1400,384,1293]" captionTargetId="figure@12.[180,1407,370,1294]" captionTargetPageId="12" captionText="FIGURE 39. Distribution of Tobantilla species." httpUri="https://zenodo.org/record/278963/files/figure.png" pageId="17" pageNumber="58">Fig. 39</figureCitation>
). More field observations, collection data or molecular data are necessary to make a concrete sex association.
</paragraph>
<paragraph id="8BA9D301D822215206B64DCAFD47FE73" blockId="17.[151,1436,151,464]" pageId="17" pageNumber="58">
The scutellar armature of this species is similar to that of the Argentinean species,
<taxonomicName id="4C16A882D8222152021B4DCBFB5BFE10" box="[1082,1223,404,428]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="charrasca">
<emphasis id="B9620F13D8222152021B4DCBFB5BFE10" box="[1082,1223,404,428]" italics="true" pageId="17" pageNumber="58">T. charrasca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C16A882D822215202DE4DCBFA08FE10" box="[1279,1428,405,428]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="montonera">
<emphasis id="B9620F13D822215202DE4DCBFA08FE10" box="[1279,1428,405,428]" italics="true" pageId="17" pageNumber="58">T. montonera</emphasis>
</taxonomicName>
, and dissimilar to that of
<taxonomicName id="4C16A882D822215207894DE7FDB9FE6C" box="[424,549,440,464]" class="Insecta" family="Mutillidae" genus="Lomachaeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="frigidula">
<emphasis id="B9620F13D822215207894DE7FDB9FE6C" box="[424,549,440,464]" italics="true" pageId="17" pageNumber="58">T. frigidula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C16A882D8222152047A4DE7FD4FFE6C" box="[603,723,440,464]" class="Insecta" family="Mutillidae" genus="Tobantilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="58" phylum="Arthropoda" rank="species" species="kolasma">
<emphasis id="B9620F13D8222152047A4DE7FD4FFE6C" box="[603,723,440,464]" italics="true" pageId="17" pageNumber="58">T. kolasma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>