258 lines
31 KiB
XML
258 lines
31 KiB
XML
<document id="CE4E2AE8F307EBD57CFD909BEB83215F" ID-DOI="10.13140/rg.2.2.11871.87201" ID-ISSN="11755334" ID-Zenodo-Dep="5315376" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629623705209" checkinUser="felipe" docAuthor="Puente, Marta, Glaw, Frank, Vieites, David R. & Vences, Miguel" docDate="2009" docId="039D87D50F22FFE0FF0D2AE2EC59FC09" docLanguage="en" docName="zt02103p076.pdf" docOrigin="Zootaxa 2103" docSource="http://dx.doi.org/10.13140/rg.2.2.11871.87201" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Lygodactylus roavolana Puente & Glaw & Vieites & Vences 2009, sp. nov." docType="treatment" docVersion="5" lastPageNumber="34" masterDocId="FFA4FFAD0F3DFFC1FF9A2E1BE962FF99" masterDocTitle="Competitividad del Sector Forestal de México Tendencias y Perspectivas" masterLastPageNumber="76" masterPageNumber="1" pageNumber="32" updateTime="1699085337495" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E7D2DB246B30A7CA7A989A336575310B">
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<mods:title id="85B9B7F677DB553E22599A8CAC8EA8E4">Competitividad del Sector Forestal de México Tendencias y Perspectivas</mods:title>
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</mods:titleInfo>
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<mods:name id="6C73730EF2F588DB0A5E4A2127134F17" type="personal">
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<mods:roleTerm id="B29617DA4000C3DCFEC52DF9BA15FBCC">Author</mods:roleTerm>
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<mods:namePart id="53CC071B738848B342521AFB88F6C7AE">Puente, Marta</mods:namePart>
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<mods:namePart id="621F331D7AADAB83ED90E4163439843D">Glaw, Frank</mods:namePart>
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<mods:roleTerm id="961AC18927F0F6597E34BA7039714863">Author</mods:roleTerm>
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<mods:namePart id="748CE43A384829789F4966E710512AAC">Vieites, David R.</mods:namePart>
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<mods:name id="C9047F56F78AFF2A218EFF4C0688EC75" type="personal">
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<mods:namePart id="E30912F199888B8839FA29A80EA50ED8">Vences, Miguel</mods:namePart>
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<mods:title id="91B1A04AEDE1BEE43610622477BEB28A">Zootaxa</mods:title>
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<mods:part id="880857F5D81DEA6C36F7DA3E3DA314DC">
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<mods:date id="38A8CDFBEE3D1DA02BD7123CCE738CA5">2009</mods:date>
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<mods:number id="B5CB37171C27CD076D8E691B1804ECB0">2009-05-11</mods:number>
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<mods:detail id="D50A68B05C7E61CB728CFD65259EB5CA" type="volume">
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<mods:number id="8388660E6C698E938DFFE67658A1EB0E">2103</mods:number>
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<mods:url id="BC174F57D2221F8DBF6C7F90D5B5A756">http://dx.doi.org/10.13140/rg.2.2.11871.87201</mods:url>
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<mods:classification id="029663C66855948FEFD39778926A1BDB">journal article</mods:classification>
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<mods:identifier id="31C1A493BB96871DE010A7024B9F8DE3" type="DOI">10.13140/rg.2.2.11871.87201</mods:identifier>
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<treatment id="039D87D50F22FFE0FF0D2AE2EC59FC09" ID-DOI="http://doi.org/10.5281/zenodo.5323820" ID-Zenodo-Dep="5323820" LSID="urn:lsid:plazi:treatment:039D87D50F22FFE0FF0D2AE2EC59FC09" httpUri="http://treatment.plazi.org/id/039D87D50F22FFE0FF0D2AE2EC59FC09" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
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<subSubSection id="C32E65480F22FFDEFF0D2AE2EB79FA8B" box="[151,539,1272,1299]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8B8B36C30F22FFDEFF0D2AE2EB79FA8B" blockId="31.[151,539,1272,1338]" box="[151,539,1272,1299]" pageId="31" pageNumber="32">
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<heading id="D0C381AF0F22FFDEFF0D2AE2EB79FA8B" bold="true" box="[151,539,1272,1299]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
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<emphasis id="B940EAD10F22FFDEFF0D2AE2EB79FA8B" bold="true" box="[151,539,1272,1299]" pageId="31" pageNumber="32">
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<taxonomicName id="4C344D400F22FFDEFF0D2AE2E8D4FA8A" authority="Puente & Glaw & Vieites & Vences, 2009" authorityName="Puente & Glaw & Vieites & Vences" authorityYear="2009" box="[151,438,1273,1299]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="roavolana" status="sp. nov.">
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<emphasis id="B940EAD10F22FFDEFF0D2AE2E8D4FA8A" bold="true" box="[151,438,1273,1299]" italics="true" pageId="31" pageNumber="32">Lygodactylus roavolana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27357AA0F22FFDEFE242AE3EB79FA8B" box="[446,539,1272,1298]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32E65480F22FFDEFF0D2B3BE99FFAA3" box="[151,253,1312,1338]" pageId="31" pageNumber="32" type="description">
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<paragraph id="8B8B36C30F22FFDEFF0D2B3BE99FFAA3" blockId="31.[151,539,1272,1338]" box="[151,253,1312,1338]" pageId="31" pageNumber="32">
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(
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<figureCitation id="130F2A460F22FFDEFF052B3BE996FAA3" box="[159,244,1312,1338]" captionStart="FIGURE 15" captionStartId="32.[151,255,1840,1864]" captionTargetBox="[197,1417,641,1809]" captionTargetId="figure-284@32.[169,1417,622,1812]" captionTargetPageId="32" captionText="FIGURE 15. Scalation and hemipenial characters of Lygodactylus roavolana, all based on male holotype ZSM 5115/ 2005. (a) head in lateral view (scale bar = 1 mm); (b) black spot above forelimb insertion, photographed in preserved specimen; (c) dorsal view of head (scale bar = 2 mm); (d) throat in ventral view (scale bar = 2 mm), (e) preanal pores in ventral view (scale bar = 1 mm); (f) lamellae of fourth toe in ventral view (scale bar = 2 mm); (g) and (h), sulcal and asulcal views of the hemipenis (scale bar = 1 mm). All drawings by M. Puente; photograph by R. D. Randrianiaina." figureDoi="http://doi.org/10.5281/zenodo.5361429" httpUri="https://zenodo.org/record/5361429/files/figure.png" pageId="31" pageNumber="32">Fig. 15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E65480F22FFDEFF0D2B76EB4BFA4C" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="8B8B36C30F22FFDEFF0D2B76EB4BFA4C" blockId="31.[151,1436,1389,1493]" pageId="31" pageNumber="32">
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<materialsCitation id="3B5C3C9E0F22FFDEFF0D2B76EB4BFA4C" collectingDate="2005-02-03" collectionCode="ZSM, FGZC" collectorName="P. Bora & F. Glaw & M. Vences. No" country="Madagascar" latitude="-24.983334" location="Madagascar" longLatPrecision="1248" longitude="46.916668" pageId="31" pageNumber="32" specimenCode="ZSM 5115, FGZC 2324" specimenCount="2" typeStatus="holotype">
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<emphasis id="B940EAD10F22FFDEFF0D2B76E84FFA1E" bold="true" box="[151,301,1389,1415]" pageId="31" pageNumber="32">
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<typeStatus id="548F88610F22FFDEFF0D2B76E868FA1E" box="[151,266,1389,1415]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
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.—
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</emphasis>
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<specimenCode id="DB929EB80F22FFDEFEB72B76E8D0FA1E" box="[301,434,1389,1415]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="31" pageNumber="32" type="Museum">ZSM 5115</specimenCode>
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/2005 (field number
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<specimenCode id="DB929EB80F22FFDEFD232B76EA30FA1E" box="[697,850,1389,1415]" collectionCode="FGZC" pageId="31" pageNumber="32">FGZC 2324</specimenCode>
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), adult male, collected on
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<date id="FF8A10030F22FFDEFB332B76EC1AFA1E" box="[1193,1400,1389,1415]" pageId="31" pageNumber="32" value="2005-02-03">
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<collectingDate id="EFCEE9EB0F22FFDEFB332B76EC1AFA1E" box="[1193,1400,1389,1415]" pageId="31" pageNumber="32" value="2005-02-03">3 February 2005</collectingDate>
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</date>
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at Manantantely (approximately
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<geoCoordinate id="EE0050040F22FFDEFE672B8FEB3FFA37" box="[509,605,1428,1454]" degrees="46" direction="east" minutes="55" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="46.916668">46°55'E</geoCoordinate>
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,
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<geoCoordinate id="EE0050040F22FFDEFDF12B8FEBA8FA37" box="[619,714,1428,1454]" degrees="24" direction="south" minutes="59" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="-24.983334">24°59'S</geoCoordinate>
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, close to sea level), southeastern
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<collectingCountry id="F32376530F22FFDEFBC62B8FED89FA37" box="[1116,1259,1428,1454]" name="Madagascar" pageId="31" pageNumber="32">Madagascar</collectingCountry>
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, by
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<collectorName id="26C153150F22FFDEFA852B8FEC1BFA37" box="[1311,1401,1428,1454]" pageId="31" pageNumber="32">P. Bora</collectorName>
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,
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<collectorName id="26C153150F22FFDEFA1D2B8FE9B5FA4C" pageId="31" pageNumber="32">F. Glaw</collectorName>
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&
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<collectorName id="26C153150F22FFDEFF662BA0E8C8FA4C" box="[252,426,1467,1493]" pageId="31" pageNumber="32">M. Vences. No</collectorName>
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<typeStatus id="548F88610F22FFDEFE2B2BA0EB47FA4C" box="[433,549,1467,1493]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
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.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32E65480F22FFE1FF0D281CEB81FF41" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="diagnosis">
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<paragraph id="8B8B36C30F22FFDEFF0D281CE981F893" blockId="31.[151,1436,1543,2034]" pageId="31" pageNumber="32">
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<emphasis id="B940EAD10F22FFDEFF0D281CE86DF9B8" bold="true" box="[151,271,1543,1569]" pageId="31" pageNumber="32">Diagnosis</emphasis>
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. The new species can be assigned to the
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<taxonomicName id="4C344D400F22FFDEFD692813EA3BF9B8" baseAuthorityName="Peters" baseAuthorityYear="1883" box="[755,857,1544,1569]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pictus">
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<emphasis id="B940EAD10F22FFDEFD692813EA3BF9B8" box="[755,857,1544,1569]" italics="true" pageId="31" pageNumber="32">L. pictus</emphasis>
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</taxonomicName>
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group based on the characters listed in the group definition above, especially by the presence of a semi-divided mental scale and the presence of two postmental scales. Within this group, it differs from
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<taxonomicName id="4C344D400F22FFDEFC9B284EEA05F9F7" baseAuthorityName="Peters" baseAuthorityYear="1883" box="[769,871,1621,1646]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pictus">
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<emphasis id="B940EAD10F22FFDEFC9B284EEA05F9F7" box="[769,871,1621,1646]" italics="true" pageId="31" pageNumber="32">L. pictus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C344D400F22FFDEFC3B284EED50F9F7" authorityName="Mertens" authorityYear="1965" box="[929,1074,1621,1646]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="tuberosus">
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<emphasis id="B940EAD10F22FFDEFC3B284EED50F9F7" box="[929,1074,1621,1646]" italics="true" pageId="31" pageNumber="32">L. tuberosus</emphasis>
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</taxonomicName>
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by the presence of a large and very distinct black spot (encircling a smaller white spot) above and in front of the insertion of forelimbs (absent in
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<emphasis id="B940EAD10F22FFDEFE8C28B8E8E7F925" box="[278,389,1699,1724]" italics="true" pageId="31" pageNumber="32">
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<taxonomicName id="4C344D400F22FFDEFE8C28B8E8E3F925" baseAuthorityName="Peters" baseAuthorityYear="1883" box="[278,385,1699,1724]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
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,
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</emphasis>
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only small spots present in
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<taxonomicName id="4C344D400F22FFDEFD4728B8EA12F925" authorityName="Mertens" authorityYear="1965" box="[733,880,1699,1724]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="tuberosus">
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<emphasis id="B940EAD10F22FFDEFD4728B8EA12F925" box="[733,880,1699,1724]" italics="true" pageId="31" pageNumber="32">L. tuberosus</emphasis>
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</taxonomicName>
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). It further differs from
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<taxonomicName id="4C344D400F22FFDEFB0028B8EC60F925" baseAuthorityName="Peters" baseAuthorityYear="1883" box="[1178,1282,1699,1724]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pictus">
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<emphasis id="B940EAD10F22FFDEFB0028B8EC60F925" box="[1178,1282,1699,1724]" italics="true" pageId="31" pageNumber="32">L. pictus</emphasis>
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</taxonomicName>
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by having 7 infralabial scales (vs. 4–6), and from
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<taxonomicName id="4C344D400F22FFDEFDD528D2EBD6F97B" baseAuthorityName="Peters" baseAuthorityYear="1883" box="[591,692,1737,1762]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="pictus">
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<emphasis id="B940EAD10F22FFDEFDD528D2EBD6F97B" box="[591,692,1737,1762]" italics="true" pageId="31" pageNumber="32">L. pictus</emphasis>
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||
</taxonomicName>
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and
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<taxonomicName id="4C344D400F22FFDEFD7428D2EA1DF97B" authorityName="Mertens" authorityYear="1965" box="[750,895,1737,1762]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="tuberosus">
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<emphasis id="B940EAD10F22FFDEFD7428D2EA1DF97B" box="[750,895,1737,1762]" italics="true" pageId="31" pageNumber="32">L. tuberosus</emphasis>
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</taxonomicName>
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by the apparent absence of a claw on the first finger.
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</paragraph>
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<paragraph id="8B8B36C30F22FFDEFF5C290DECEDF83C" blockId="31.[151,1436,1543,2034]" pageId="31" pageNumber="32">
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Species in the
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<taxonomicName id="4C344D400F22FFDEFEEE290CEB0BF8A9" box="[372,617,1815,1840]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="madagascariensis">
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<emphasis id="B940EAD10F22FFDEFEEE290CEB0BF8A9" box="[372,617,1815,1840]" italics="true" pageId="31" pageNumber="32">L. madagascariensis</emphasis>
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</taxonomicName>
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group share with the new species a number of only two postmentals and the absence of a claw on the first finger, but in these species, the mental scale is undivided. A claw on the first finger is also absent in
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<taxonomicName id="4C344D400F22FFDEFE67297FEBEEF8E4" box="[509,652,1892,1917]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="mirabilis">
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<emphasis id="B940EAD10F22FFDEFE67297FEBEEF8E4" box="[509,652,1892,1917]" italics="true" pageId="31" pageNumber="32">L. mirabilis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C344D400F22FFDEFD3A297FEA2CF8E4" authorityName="Pasteur" authorityYear="1995" box="[672,846,1892,1917]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="intermedius">
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<emphasis id="B940EAD10F22FFDEFD3A297FEA2CF8E4" box="[672,846,1892,1917]" italics="true" pageId="31" pageNumber="32">L. intermedius</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C344D400F22FFDEFC09297FED4BF8E4" box="[915,1065,1892,1917]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="montanus">
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<emphasis id="B940EAD10F22FFDEFC09297FED4BF8E4" box="[915,1065,1892,1917]" italics="true" pageId="31" pageNumber="32">L. montanus</emphasis>
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</taxonomicName>
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, but these species are readily distinguished by their keeled or partially keeled dorsal scales (versus granular dorsal scales in
|
||
<taxonomicName id="4C344D400F22FFDEFB722990EC1CF83D" authorityName="Puente & Glaw & Vieites & Vences" authorityYear="2009" box="[1256,1406,1931,1956]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="roavolana">
|
||
<emphasis id="B940EAD10F22FFDEFB722990EC1CF83D" box="[1256,1406,1931,1956]" italics="true" pageId="31" pageNumber="32">L. roavolana</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8B36C30F22FFE1FF0D29AAEB81FF41" blockId="31.[151,1436,1543,2034]" lastBlockId="32.[151,1436,152,564]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
|
||
We cannot fully exclude that the absence of a claw on the first finger in the new species may be an artefact, because we could verify it on only one forelimb in a single specimen. However, the new species is also diagnosed by its unique colour pattern: the conspicuous pair of large ocellae above the forelimbs is not known from any other species of Malagasy
|
||
<taxonomicName id="4C344D400F1DFFE1FDDB2EA4EBBEFF41" box="[577,732,191,216]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940EAD10F1DFFE1FDDB2EA4EBBEFF41" box="[577,732,191,216]" italics="true" pageId="32" pageNumber="33">Lygodactylus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E65480F1DFFE0FF5C2EFEEDCBFD1B" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8B8B36C30F1DFFE1FF5C2EFEE860FDAD" blockId="32.[151,1436,152,564]" pageId="32" pageNumber="33">
|
||
<emphasis id="B940EAD10F1DFFE1FF5C2EFEE889FF66" bold="true" box="[198,491,229,255]" pageId="32" pageNumber="33">
|
||
Description of
|
||
<typeStatus id="548F88610F1DFFE1FE1A2EFEE889FF66" box="[384,491,229,255]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Adult male, hemipenes everted, in good state of preservation, although the tail is broken and one forelimb is partly removed as source of tissue for molecular analysis. SVL
|
||
<quantity id="4CCC9B260F1DFFE1FB662F17EC08FEBF" box="[1276,1386,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="32" pageNumber="33" unit="mm" value="23.6">23.6 mm</quantity>
|
||
, tail regenerated (TAL
|
||
<quantity id="4CCC9B260F1DFFE1FEE92F29E8BDFED5" box="[371,479,306,332]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="32" pageNumber="33" unit="mm" value="20.4">20.4 mm</quantity>
|
||
) for other measurements see
|
||
<tableCitation id="C6B603780F1DFFE1FCDB2F29EAFFFED5" box="[833,925,306,332]" captionStart="TABLE 2" captionStartId="25.[151,241,151,175]" captionText="TABLE 2. Character comparison of representative morphological characters in the species belonging to the Lygodactylus pictus group, separately for males and females. Values are given as mean (in bold) ± standard deviation, minimum-maximum, modal, and number of measured individuals (n)." pageId="32" pageNumber="33">Table 2</tableCitation>
|
||
; granular dorsal scales; first finger present but very small, without bearing a claw; 4 pairs of subdigital lamellae on the fourth toe; mental scale semidivided into three parts by sutures; only slight contact between posterior projection of mental scale and first infralabial; two symmetrical postmental scales; five postpostmental scales; seven infralabial scales; seven supralabial scales; two internasal scales; six preanal pores; tail without whorls; four weakly expressed dorsolateral tubercles, each composed of 1–2 scales; 185 dorsal scales along the body; 88 dorsal scales around the body.
|
||
</paragraph>
|
||
<caption id="DF4B664B0F1DFFE1FF0D292BEC0AF84B" ID-DOI="http://doi.org/10.5281/zenodo.5361429" ID-Zenodo-Dep="5361429" httpUri="https://zenodo.org/record/5361429/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,255,1840,1864]" targetBox="[197,1417,641,1809]" targetPageId="32">
|
||
<paragraph id="8B8B36C30F1DFFE1FF0D292BEC0AF84B" blockId="32.[151,1436,1840,2002]" pageId="32" pageNumber="33">
|
||
<emphasis id="B940EAD10F1DFFE1FF0D292BE843F8D1" bold="true" box="[151,289,1840,1864]" pageId="32" pageNumber="33">FIGURE 15</emphasis>
|
||
. Scalation and hemipenial characters of
|
||
<taxonomicName id="4C344D400F1DFFE1FD45292BEA80F8DE" authorityName="Puente & Glaw & Vieites & Vences" authorityYear="2009" box="[735,994,1840,1863]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="roavolana">
|
||
<emphasis id="B940EAD10F1DFFE1FD45292BEA80F8DE" box="[735,994,1840,1863]" italics="true" pageId="32" pageNumber="33">Lygodactylus roavolana</emphasis>
|
||
</taxonomicName>
|
||
, all based on male holotype ZSM 5115/ 2005. (a) head in lateral view (scale bar = 1 mm); (b) black spot above forelimb insertion, photographed in preserved specimen; (c) dorsal view of head (scale bar = 2 mm); (d) throat in ventral view (scale bar = 2 mm), (e) preanal pores in ventral view (scale bar = 1 mm); (f) lamellae of fourth toe in ventral view (scale bar = 2 mm); (g) and (h), sulcal and asulcal views of the hemipenis (scale bar = 1 mm). All drawings by M. Puente; photograph by R. D. Randrianiaina.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8B36C30F1CFFE0FF5C2E83E8D7FE59" blockId="33.[151,1437,152,912]" pageId="33" pageNumber="34">
|
||
Dorsally light brown on the dorsum and greyish on the flanks, with a pair of indistinct yellowish dorsolateral stripes, one on each side, that start just behind the eye and run onto the tail; at the height of the hindlimbs, these stripes become more distinct and the dorsal area between them more dark brown. Some whitish spots on the upper lip. Above and anterior to the forelimb, there is a large semicircular black marking with a central white spot and bordered ventrally by a distinct short white line. Tail dorsally with some irregular beige-brown marbling. Ventrally, in life, the throat and tail were yellow and the chest and venter white. The throat has an indistinct pattern of irregular and poorly marked dark chevrons and longitudinal stripes (
|
||
<figureCitation id="130F2A460F1CFFE0FF692FBDE819FE59" box="[243,379,422,448]" captionStart="FIGURE 12" captionStartId="24.[151,255,1434,1458]" captionTargetBox="[154,1433,189,1405]" captionTargetId="figure-72@24.[154,1434,189,1405]" captionTargetPageId="24" captionText="FIGURE 12. Photos of living specimens assigned to species in the Lygodactylus pictus group. (a) L. tuberosus, Berenty; (b) L. tuberosus, Ifaty; (c) and (d), L. pictus, near Sendrisoa, Andringitra; (e) and (f), dorsolateral and ventral view of L. roavolana, holotype specimen, from Manantantely. All photographs by F. Glaw & M. Vences." figureDoi="http://doi.org/10.5281/zenodo.5315402" httpUri="https://zenodo.org/record/5315402/files/figure.png" pageId="33" pageNumber="34">Figs. 12e–f</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A460F1CFFE0FE1D2FBDE8C5FE59" box="[391,423,422,448]" captionStart="FIGURE 15" captionStartId="32.[151,255,1840,1864]" captionTargetBox="[197,1417,641,1809]" captionTargetId="figure-284@32.[169,1417,622,1812]" captionTargetPageId="32" captionText="FIGURE 15. Scalation and hemipenial characters of Lygodactylus roavolana, all based on male holotype ZSM 5115/ 2005. (a) head in lateral view (scale bar = 1 mm); (b) black spot above forelimb insertion, photographed in preserved specimen; (c) dorsal view of head (scale bar = 2 mm); (d) throat in ventral view (scale bar = 2 mm), (e) preanal pores in ventral view (scale bar = 1 mm); (f) lamellae of fourth toe in ventral view (scale bar = 2 mm); (g) and (h), sulcal and asulcal views of the hemipenis (scale bar = 1 mm). All drawings by M. Puente; photograph by R. D. Randrianiaina." figureDoi="http://doi.org/10.5281/zenodo.5361429" httpUri="https://zenodo.org/record/5361429/files/figure.png" pageId="33" pageNumber="34">15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8B36C30F1CFFE0FF5C2FD6EDCBFD1B" blockId="33.[151,1437,152,912]" pageId="33" pageNumber="34">
|
||
<emphasis id="B940EAD10F1CFFE0FF5C2FD6E8B0FE7E" bold="true" box="[198,466,461,487]" pageId="33" pageNumber="34">Hemipenial structure</emphasis>
|
||
. Hemipenis subcylindrical, total length ca.
|
||
<quantity id="4CCC9B260F1CFFE0FC4E2FD6ED50FE7E" box="[980,1074,461,487]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="33" pageNumber="34" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, with a relatively large pedicel of ca.
|
||
<quantity id="4CCC9B260F1CFFE0FF792FEFE823FD97" box="[227,321,500,526]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="33" pageNumber="34" unit="mm" value="1.8">1.8 mm</quantity>
|
||
. The apex is divided in two short lobes each of which measures ca.
|
||
<quantity id="4CCC9B260F1CFFE0FBE92FEFED89FD97" box="[1139,1259,500,526]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="33" pageNumber="34" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
|
||
in length. The truncus and lobes are covered with fields and serrated ridges of pointed papillae. Sulcus spermaticus formed by two channels (running to the apex of each arm); there are no distinct sulcal lips and no papillae around the sulcus spermaticus. Very similar to the hemipenes of
|
||
<taxonomicName id="4C344D400F1CFFE0FC9D2C73EAF4FD18" authorityName="Mertens" authorityYear="1965" box="[775,918,616,641]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="tuberosus">
|
||
<emphasis id="B940EAD10F1CFFE0FC9D2C73EAF4FD18" box="[775,918,616,641]" italics="true" pageId="33" pageNumber="34">L. tuberosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C344D400F1CFFE0FC4A2C73ED57FD18" baseAuthorityName="Peters" baseAuthorityYear="1883" box="[976,1077,616,641]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B940EAD10F1CFFE0FC4A2C73ED57FD18" box="[976,1077,616,641]" italics="true" pageId="33" pageNumber="34">L. pictus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130F2A460F1CFFE0FBDF2C73EDFEFD1B" box="[1093,1180,616,642]" captionStart="FIGURE 15" captionStartId="32.[151,255,1840,1864]" captionTargetBox="[197,1417,641,1809]" captionTargetId="figure-284@32.[169,1417,622,1812]" captionTargetPageId="32" captionText="FIGURE 15. Scalation and hemipenial characters of Lygodactylus roavolana, all based on male holotype ZSM 5115/ 2005. (a) head in lateral view (scale bar = 1 mm); (b) black spot above forelimb insertion, photographed in preserved specimen; (c) dorsal view of head (scale bar = 2 mm); (d) throat in ventral view (scale bar = 2 mm), (e) preanal pores in ventral view (scale bar = 1 mm); (f) lamellae of fourth toe in ventral view (scale bar = 2 mm); (g) and (h), sulcal and asulcal views of the hemipenis (scale bar = 1 mm). All drawings by M. Puente; photograph by R. D. Randrianiaina." figureDoi="http://doi.org/10.5281/zenodo.5361429" httpUri="https://zenodo.org/record/5361429/files/figure.png" pageId="33" pageNumber="34">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E65480F1CFFE0FF5C2C95EC59FC09" pageId="33" pageNumber="34" type="etymology">
|
||
<paragraph id="8B8B36C30F1CFFE0FF5C2C95E855FD6F" blockId="33.[151,1437,152,912]" pageId="33" pageNumber="34">
|
||
<emphasis id="B940EAD10F1CFFE0FF5C2C95E831FD31" bold="true" box="[198,339,654,680]" pageId="33" pageNumber="34">Etymology.</emphasis>
|
||
The name “roavolana” refers to the Malagasy words
|
||
<emphasis id="B940EAD10F1CFFE0FC5C2C94EA8CFD31" box="[966,1006,655,680]" italics="true" pageId="33" pageNumber="34">roa</emphasis>
|
||
meaning “two” and
|
||
<emphasis id="B940EAD10F1CFFE0FB7A2C94EC52FD31" box="[1248,1328,655,680]" italics="true" pageId="33" pageNumber="34">volana</emphasis>
|
||
meaning “moon”, and refers to the two extremely conspicuous ocellae of the new species. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="8B8B36C30F1CFFE0FF5C2D19EC59FC09" blockId="33.[151,1437,152,912]" pageId="33" pageNumber="34">
|
||
<emphasis id="B940EAD10F1CFFE0FF5C2D19EB62FC85" bold="true" box="[198,512,770,796]" pageId="33" pageNumber="34">Habitat and distribution</emphasis>
|
||
. The
|
||
<typeStatus id="548F88610F1CFFE0FDD12D19EBD3FC85" box="[587,689,770,796]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
was collected in the late afternoon, active on a tree trunk in disturbed low-altitude rainforest, ca.
|
||
<quantity id="4CCC9B260F1CFFE0FDCE2D32EBE7FCDA" box="[596,645,809,835]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="m" value="1.0">1 m</quantity>
|
||
above the ground. This species has apparently not been detected during previous surveys in the nearby Andohahela reserve (
|
||
<bibRefCitation id="EFA54B320F1CFFE0FCC02D4BED97FCF3" author="Andreone, F. & Randriamahazo, H." box="[858,1269,848,874]" pageId="33" pageNumber="34" pagination="95 - 127" refId="ref48559" refString="Andreone, F. & Randriamahazo, H. (1997) Ecological and taxonomic observations on the amphibians and reptiles of the Andohahela low altitude rainforest, S. Madagascar. Revue francaise d'Aquariologie Herpetologie, 24, 95 - 127." type="journal article" year="1997">Andreone & Randriamahazo 1997</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA54B320F1CFFE0FA992D4BE999FC09" author="Nussbaum, R. A. & Raxworthy, C. J. & Raselimanana, A. P. & Ramanamanjato, J. B." pageId="33" pageNumber="34" pagination="155 - 173" refId="ref49736" refString="Nussbaum, R. A., Raxworthy, C. J., Raselimanana, A. P. & Ramanamanjato, J. B. (1999) Amphibians and reptiles of the Reserve Naturelle Integrale d'Andohahela, Madagascar. Fieldiana Zoology, 94, 155 - 173." type="journal article" year="1999">
|
||
Nussbaum
|
||
<emphasis id="B940EAD10F1CFFE0FA1D2D4BE9D7FC09" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
). However, the species has more recently been recorded from this reserve (
|
||
<bibRefCitation id="EFA54B320F1CFFE0FBF22D6DEC4DFC09" author="Theisinger, O." box="[1128,1327,886,912]" pageId="33" pageNumber="34" refId="ref50614" refString="Theisinger, O. (2008) Reptiles of the dry forest. What to find in the Andohahela National Park, Madagascar. Leaflet published by University of Hamburg." type="book" year="2008">Theisinger 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |