317 lines
35 KiB
XML
317 lines
35 KiB
XML
<document id="0E0626E16AC1EE5583D921C17440A0D0" ID-DOI="10.5281/zenodo.5167791" ID-ISSN="1942-1354" ID-Zenodo-Dep="5167791" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628270932123" checkinUser="felipe" docAuthor="Mawdsley, Jonathan R." docDate="2009" docId="03D087FEFFA1FFD9FF2496A5FB9DF902" docLanguage="en" docName="InsectaMundi.0094.1-10.pdf" docOrigin="Insecta Mundi 2009 (94)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Cicindela obsoleta Say" docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="FFE9FF86FFA2FFDCFFE49020FFEEFF8F" masterDocTitle="Geographic variation in U. S. populations of the tiger beetle Cicindela obsoleta Say (Coleoptera: Cicindelidae)" masterLastPageNumber="10" masterPageNumber="1" pageNumber="2" updateTime="1699038641360" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="66A556BF6D0F8099C5F546B130735B84">
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<mods:title id="6B5A2CBB63143752185C6E325C2EB322">Geographic variation in U. S. populations of the tiger beetle Cicindela obsoleta Say (Coleoptera: Cicindelidae)</mods:title>
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<mods:name id="8007C04758839E90312BCF8E73F923E9" type="personal">
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<mods:roleTerm id="76C8E911494021C95490FC4967E6D30E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A5DE729369AC64C7C7D13385DEB917D3">Mawdsley, Jonathan R.</mods:namePart>
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<mods:typeOfResource id="61F74D2435BEB1A4EE73BE720CFD1389">text</mods:typeOfResource>
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<mods:titleInfo id="7A4FD0607670579E67309F0DDD752223">
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<mods:title id="4E4E17CAE67133E9D3E0635BC5E36845">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="CDAF9A09D6D00469E7790CD1B95B477E">
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<mods:date id="1A2EFAD7FD231717DA4C7470310D9310">2009</mods:date>
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<mods:detail id="F3BE2B772CB0BEC1CF0DF3CBEA8A53CB" type="pubDate">
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<mods:number id="D37F6DBE06993577F72F108270691653">2009-09-25</mods:number>
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</mods:detail>
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<mods:detail id="78C3C2EAB2E7750390EBBF5D4D2CA576" type="volume">
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<mods:number id="7419627676A4B088C18C1D69A43B3AC5">2009</mods:number>
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</mods:detail>
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<mods:detail id="F3B85B85A44FAA9239F627B1E9DA83E4" type="issue">
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<mods:number id="73D1918FF5487B32A693AC89D8697743">94</mods:number>
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</mods:detail>
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<mods:extent id="FC89DB64D41679F127076B7A8E4CD72C" unit="page">
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<mods:start id="46B429BF8096E7AD6D581B33F5586E69">1</mods:start>
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<mods:end id="778C40E3406BA4B3AF7FD730319B85D7">10</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="22F7BF19FF60B7AA497448E4DAD7CEB8">journal article</mods:classification>
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<mods:identifier id="EB27DFFCDC72AA1FC941BA9D311701A1" type="DOI">10.5281/zenodo.5167791</mods:identifier>
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<mods:identifier id="6399AAE021BEC7BE219F24705222BE39" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="D4B700D30E961D9FEB2A2DDCB2BBCC45" type="Zenodo-Dep">5167791</mods:identifier>
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</mods:mods>
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<treatment id="03D087FEFFA1FFD9FF2496A5FB9DF902" LSID="urn:lsid:plazi:treatment:03D087FEFFA1FFD9FF2496A5FB9DF902" httpUri="http://treatment.plazi.org/id/03D087FEFFA1FFD9FF2496A5FB9DF902" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
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<subSubSection id="C3636563FFA1FFDFFF2496A5FE1CF914" box="[192,498,1667,1692]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="8BC636E8FFA1FFDFFF2496A5FE1CF914" blockId="3.[192,498,1667,1692]" box="[192,498,1667,1692]" pageId="3" pageNumber="2">
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<heading id="D08E8184FFA1FFDFFF2496A5FE1CF914" bold="true" box="[192,498,1667,1692]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
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<taxonomicName id="4C794D6BFFA1FFDFFF2496A5FE1CF914" ID-CoL="V8C8" authority="Say" authorityName="Say" box="[192,498,1667,1692]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B90DEAFAFFA1FFDFFF2496A5FE1CF914" bold="true" box="[192,498,1667,1692]" pageId="3" pageNumber="2">
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<emphasis id="B90DEAFAFFA1FFDFFF2496A5FE55F913" bold="true" box="[192,443,1669,1692]" italics="true" pageId="3" pageNumber="2">Cicindela obsoleta</emphasis>
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Say
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3636563FFA1FFDFFF1496E3FB42F954" box="[240,1196,1731,1756]" pageId="3" pageNumber="2" type="reference_group">
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<paragraph id="8BC636E8FFA1FFDFFF1496E3FB42F954" blockId="3.[240,1196,1731,1756]" box="[240,1196,1731,1756]" pageId="3" pageNumber="2">
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<treatmentCitationGroup id="AB6911C6FFA1FFDFFF1496E3FB42F954" box="[240,1196,1731,1756]" pageId="3" pageNumber="2">
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<treatmentCitation id="0AD810F9FFA1FFDFFF1496E3FD6AF954" author="Say, T." box="[240,644,1731,1755]" page=")" pageId="3" pageNumber="2" year="1823">
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<taxonomicName id="4C794D6BFFA1FFDFFF1496E3FD6AF954" ID-CoL="V8C8" authority="Say (1823: 143)" authorityName="Say" authorityPageNumber="143" authorityYear="1823" box="[240,644,1731,1755]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B90DEAFAFFA1FFDFFF1496E3FE22F954" box="[240,460,1731,1755]" italics="true" pageId="3" pageNumber="2">Cicindela obsoleta</emphasis>
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<bibRefCitation id="EFE84B19FFA1FFDFFE3596E3FD6AF954" author="Say, T." box="[465,644,1731,1755]" pageId="3" pageNumber="2" pagination="139 - 216" refId="ref7261" refString="Say, T. 1823. Description of coleopterous insects collected in the late expedition to the Rocky Mountains, performed by order of Mr. Calhoun, Secretary of War, under the command of Major Long. Journal of the Academy of Natural Sciences of Philadelphia 3 (1): 139 - 216." type="journal article" year="1823">Say (1823:143)</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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<typeStatus id="54C2884AFFA1FFDFFD6D96E4FD2BF953" box="[649,709,1732,1756]" pageId="3" pageNumber="2">Type</typeStatus>
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destroyed, but identity well established.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3636563FFA1FFD9FF249723FB9DF902" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="diagnosis">
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<paragraph id="8BC636E8FFA1FFDFFF249723FA4EF8F4" blockId="3.[192,1441,1795,1915]" pageId="3" pageNumber="2">
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<emphasis id="B90DEAFAFFA1FFDFFF249723FEA7F894" bold="true" box="[192,329,1795,1819]" pageId="3" pageNumber="2">Diagnosis</emphasis>
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. A large to very large tiger beetle (length
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<quantity id="4C819B0DFFA1FFDFFCA09723FC2EF894" box="[836,960,1795,1819]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="3" pageNumber="2" unit="mm" value="16.5" valueMax="20.0" valueMin="13.0">13-20 mm</quantity>
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) associated with desert grasslands and woodlands in
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<collectingRegion id="49BDF80AFFA1FFDFFE869703FE2AF8B4" box="[354,452,1827,1851]" country="United States of America" name="Arizona" pageId="3" pageNumber="2">Arizona</collectingRegion>
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,
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<collectingRegion id="49BDF80AFFA1FFDFFE299703FDD5F8B4" box="[461,571,1827,1851]" country="United States of America" name="Colorado" pageId="3" pageNumber="2">Colorado</collectingRegion>
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,
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<collectingRegion id="49BDF80AFFA1FFDFFDA09704FD4EF8B4" box="[580,672,1828,1851]" country="United States of America" name="Kansas" pageId="3" pageNumber="2">Kansas</collectingRegion>
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,
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<collectingRegion id="49BDF80AFFA1FFDFFD4E9704FCD0F8B4" box="[682,830,1827,1851]" country="United States of America" name="New Mexico" pageId="3" pageNumber="2">New Mexico</collectingRegion>
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,
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<collectingRegion id="49BDF80AFFA1FFDFFCA39703FC29F8B4" box="[839,967,1827,1851]" country="United States of America" name="Oklahoma" pageId="3" pageNumber="2">Oklahoma</collectingRegion>
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, and
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<collectingRegion id="49BDF80AFFA1FFDFFBE79704FBA3F8B4" box="[1027,1101,1828,1851]" country="United States of America" name="Texas" pageId="3" pageNumber="2">Texas</collectingRegion>
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, with isolated populations in
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<collectingRegion id="49BDF80AFFA1FFDFFF249763FEDBF8D4" box="[192,309,1859,1883]" country="United States of America" name="Arkansas" pageId="3" pageNumber="2">Arkansas</collectingRegion>
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and
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<collectingRegion id="49BDF80AFFA1FFDFFE8A9763FE30F8D4" box="[366,478,1859,1883]" country="United States of America" name="Missouri" pageId="3" pageNumber="2">Missouri</collectingRegion>
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. Dorsal coloration may be matte black, brown, blue, olive-green, or bright green, but never strongly iridescent. Elytral markings variable, with yellow or white maculae and/or fasciae
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</paragraph>
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<caption id="DF066660FFA6FFD8FF24940FFDB9FB7E" ID-DOI="http://doi.org/10.5281/zenodo.5167793" ID-Zenodo-Dep="5167793" httpUri="https://zenodo.org/record/5167793/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,276,1071,1093]" targetBox="[192,1439,192,1041]" targetPageId="4">
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<paragraph id="8BC636E8FFA6FFD8FF24940FFDB9FB7E" blockId="4.[192,1440,1071,1266]" pageId="4" pageNumber="3">
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<emphasis id="B90DEAFAFFA6FFD8FF24940FFEAAFBCB" bold="true" box="[192,324,1071,1093]" pageId="4" pageNumber="3">Figure 1-9.</emphasis>
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Adult habitus photographs of pinned specimens of
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<taxonomicName id="4C794D6BFFA6FFD8FC97940FFBD2FBCB" box="[883,1084,1071,1092]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B90DEAFAFFA6FFD8FC97940FFBD2FBCB" box="[883,1084,1071,1092]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta</emphasis>
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</taxonomicName>
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in NMNH.
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<emphasis id="B90DEAFAFFA6FFD8FB5F940FFB3CFBCA" bold="true" box="[1211,1234,1071,1093]" pageId="4" pageNumber="3">1)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FB33940FFEA2FBEE" authority="obsoleta Say" authorityName="Say" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="obsoleta">
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<emphasis id="B90DEAFAFFA6FFD8FB33940FFEF4FBEE" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta obsoleta</emphasis>
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Say
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</taxonomicName>
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, male, Silver City, NM.
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<emphasis id="B90DEAFAFFA6FFD8FDBC946BFD81FBEE" bold="true" box="[600,623,1099,1121]" pageId="4" pageNumber="3">2)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FD91946CFC4FFBEE" authority="obsoleta" box="[629,929,1100,1121]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="obsoleta">
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<emphasis id="B90DEAFAFFA6FFD8FD91946CFC4FFBEE" box="[629,929,1100,1121]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta obsoleta</emphasis>
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</taxonomicName>
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, female, Carrizo Creek Picnic Area, Baca Co., CO.
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<emphasis id="B90DEAFAFFA6FFD8FF109448FEE5FBF1" bold="true" box="[244,267,1128,1150]" pageId="4" pageNumber="3">3)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FEF09449FDAAFBF1" authority="obsoleta" box="[276,580,1129,1150]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="obsoleta">
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<emphasis id="B90DEAFAFFA6FFD8FEF09449FDAAFBF1" box="[276,580,1129,1150]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta obsoleta</emphasis>
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</taxonomicName>
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, male, Silver City, NM.
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<emphasis id="B90DEAFAFFA6FFD8FCBB9448FC98FBF1" bold="true" box="[863,886,1128,1150]" pageId="4" pageNumber="3">4)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FC9B9449FB5EFBF1" authority="obsoleta" box="[895,1200,1129,1150]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="obsoleta">
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<emphasis id="B90DEAFAFFA6FFD8FC9B9449FB5EFBF1" box="[895,1200,1129,1150]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta obsoleta</emphasis>
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</taxonomicName>
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X
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<taxonomicName id="4C794D6BFFA6FFD8FB369449FEADFB15" authorityName="Bates" authorityYear="1890" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="santaclarae">
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<emphasis id="B90DEAFAFFA6FFD8FB369449FEADFB15" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta santaclarae</emphasis>
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</taxonomicName>
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, female, Silver City, NM.
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<emphasis id="B90DEAFAFFA6FFD8FD9C94A5FD61FB14" bold="true" box="[632,655,1157,1179]" pageId="4" pageNumber="3">5)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FD7D94A5FBAFFB14" authority="Bates" authorityName="Bates" authorityYear="1890" box="[665,1089,1157,1179]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="santaclarae">
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<emphasis id="B90DEAFAFFA6FFD8FD7D94A5FC1BFB15" box="[665,1013,1157,1178]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta santaclarae</emphasis>
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Bates
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</taxonomicName>
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, syntype female, Santa Clara, Chihuahua, México.
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<emphasis id="B90DEAFAFFA6FFD8FE419482FE52FB37" bold="true" box="[421,444,1186,1208]" pageId="4" pageNumber="3">6)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FE269482FCFAFB38" authorityName="Bates" authorityYear="1890" box="[450,788,1186,1207]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="santaclarae">
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<emphasis id="B90DEAFAFFA6FFD8FE269482FCFAFB38" box="[450,788,1186,1207]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta santaclarae</emphasis>
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</taxonomicName>
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, male, Ft. Wingate, NM.
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<emphasis id="B90DEAFAFFA6FFD8FBCF9482FBAFFB37" bold="true" box="[1067,1089,1186,1208]" pageId="4" pageNumber="3">7)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FBA39482FA77FB38" authorityName="Bates" authorityYear="1890" box="[1095,1433,1186,1207]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="santaclarae">
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<emphasis id="B90DEAFAFFA6FFD8FBA39482FA77FB38" box="[1095,1433,1186,1207]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta santaclarae</emphasis>
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</taxonomicName>
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, female, Ft. Wingate, NM.
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<emphasis id="B90DEAFAFFA6FFD8FE15949FFDE6FB5A" bold="true" box="[497,520,1215,1237]" pageId="4" pageNumber="3">8)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FDF6949FFC80FB5B" authorityName="Bates" authorityYear="1890" box="[530,878,1215,1236]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="santaclarae">
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<emphasis id="B90DEAFAFFA6FFD8FDF6949FFC80FB5B" box="[530,878,1215,1236]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta santaclarae</emphasis>
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</taxonomicName>
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, female, Ft. Wingate, NM.
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<emphasis id="B90DEAFAFFA6FFD8FB54949FFB28FB5A" bold="true" box="[1200,1222,1215,1237]" pageId="4" pageNumber="3">9)</emphasis>
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<taxonomicName id="4C794D6BFFA6FFD8FB35949FFEADFB7E" authorityName="Bates" authorityYear="1890" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="santaclarae">
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<emphasis id="B90DEAFAFFA6FFD8FB35949FFEADFB7E" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta santaclarae</emphasis>
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</taxonomicName>
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, male, Ft. Wingate, NM.
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</paragraph>
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</caption>
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<paragraph id="8BC636E8FFA6FFD8FF249533FF18FAC4" blockId="4.[192,1440,1299,1355]" pageId="4" pageNumber="3">often present. Pronotum subquadrate, with a thin band of reclinate setae along the lateral edges of the disc.</paragraph>
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<paragraph id="8BC636E8FFA6FFD8FF249553FC9EF9A4" blockId="4.[192,1440,1395,1900]" pageId="4" pageNumber="3">
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<emphasis id="B90DEAFAFFA6FFD8FF249553FE14FA04" bold="true" box="[192,506,1395,1419]" pageId="4" pageNumber="3">Subspecific taxonomy.</emphasis>
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Four subspecies are recognized here, confirming the traditional taxonomic arrangement in
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<taxonomicName id="4C794D6BFFA6FFD8FE6E95B3FD82FA24" box="[394,620,1427,1451]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obsoleta">
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<emphasis id="B90DEAFAFFA6FFD8FE6E95B3FD82FA24" box="[394,620,1427,1451]" italics="true" pageId="4" pageNumber="3">Cicindela obsoleta</emphasis>
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</taxonomicName>
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(
|
||
<bibRefCitation id="EFE84B19FFA6FFD8FD9895B3FCF1FA24" author="Freitag, R." box="[636,799,1427,1451]" pageId="4" pageNumber="3" refId="ref6816" refString="Freitag, R. 1999. Catalogue of the tiger beetles of Canada and the United States. NRC Research Press; Ottawa. 195 p." type="book" year="1999">Freitag 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE84B19FFA6FFD8FCCF95B4FBF3FA24" author="Pearson, D. L. & C. B. Knisley & C. J. Kazilek" box="[811,1053,1427,1451]" pageId="4" pageNumber="3" refId="ref7148" refString="Pearson, D. L., C. B. Knisley, and C. J. Kazilek. 2006. A field guide to tiger beetles of the United States and Canada: Identification, natural history, and distribution of the Cicindelidae. Oxford University Press; New York. vii + 227 p." type="book" year="2006">Pearson et al. 2006</bibRefCitation>
|
||
). Of the 1,424 specimens that I examined, 1,393 could be unequivocally assigned to a single subspecies on the basis of their elytral markings. The remaining
|
||
<specimenCount id="9D7FFD61FFA6FFD8FE5895F3FD8EFA64" box="[444,608,1491,1515]" pageId="4" pageNumber="3" type="generic">31 specimens</specimenCount>
|
||
are more or less intermediate between the two forms that have been conventionally recognized as
|
||
<emphasis id="B90DEAFAFFA6FFD8FDC395D4FD28F984" box="[551,710,1523,1547]" italics="true" pageId="4" pageNumber="3">
|
||
C. o.
|
||
<taxonomicName id="4C794D6BFFA6FFD8FD8695D3FD28F984" box="[610,710,1523,1547]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obsoleta">obsoleta</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B90DEAFAFFA6FFD8FCE595D4FC20F984" box="[769,974,1523,1547]" italics="true" pageId="4" pageNumber="3">C. o. santaclarae</emphasis>
|
||
Bates. As discussed below, I interpret these forms as intergrades between these two subspecies.
|
||
</paragraph>
|
||
<paragraph id="8BC636E8FFA6FFD9FF149613FAFAF982" blockId="4.[192,1440,1395,1900]" lastBlockId="5.[192,1441,1461,1677]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
|
||
<bibRefCitation id="EFE84B19FFA6FFD8FF149613FE7CF9C4" author="Cazier, M. A." box="[240,402,1587,1611]" pageId="4" pageNumber="3" pagination="231 - 309" refId="ref6746" refString="Cazier, M. A. 1954. A review of the Mexican tiger beetles of the genus Cicindela (Coleoptera: Cicindelidae). Bulletin of the American Museum of Natural History 103 (3): 231 - 309." type="journal article" year="1954">Cazier (1954)</bibRefCitation>
|
||
reviewed the Mexican subspecies of
|
||
<taxonomicName id="4C794D6BFFA6FFD8FCAE9614FC22F9C4" box="[842,972,1587,1611]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B90DEAFAFFA6FFD8FCAE9614FC22F9C4" box="[842,972,1587,1611]" italics="true" pageId="4" pageNumber="3">C. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
and recognized two additional subspecific taxa,
|
||
<emphasis id="B90DEAFAFFA6FFD8FEDA9674FDF0F9E4" box="[318,542,1619,1643]" italics="true" pageId="4" pageNumber="3">C. o. latemaculata</emphasis>
|
||
Becker from
|
||
<collectingRegion id="49BDF80AFFA6FFD8FD279674FCDEF9E3" box="[707,816,1620,1644]" country="Mexico" name="Durango" pageId="4" pageNumber="3">Durango</collectingRegion>
|
||
and
|
||
<emphasis id="B90DEAFAFFA6FFD8FC889674FBFDF9E4" box="[876,1043,1619,1643]" italics="true" pageId="4" pageNumber="3">C. o. juvenilis</emphasis>
|
||
Horn from
|
||
<collectingRegion id="49BDF80AFFA6FFD8FB469673FB14F9E4" box="[1186,1274,1619,1643]" country="Mexico" name="Jalisco" pageId="4" pageNumber="3">Jalisco</collectingRegion>
|
||
,
|
||
<collectingRegion id="49BDF80AFFA6FFD8FAE19673FA88F9E4" box="[1285,1382,1619,1643]" country="Mexico" name="Sinaloa" pageId="4" pageNumber="3">Sinaloa</collectingRegion>
|
||
, and
|
||
<collectingRegion id="49BDF80AFFA6FFD8FF249653FEF7F904" box="[192,281,1651,1675]" country="Mexico" name="Sonora" pageId="4" pageNumber="3">Sonora</collectingRegion>
|
||
. However, Cazier’s treatment of
|
||
<taxonomicName id="4C794D6BFFA6FFD8FD7F9654FCF2F904" box="[667,796,1651,1675]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B90DEAFAFFA6FFD8FD7F9654FCF2F904" box="[667,796,1651,1675]" italics="true" pageId="4" pageNumber="3">C. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
is limited, as it was based on material collected at just 11 localities in
|
||
<collectingCountry id="F36E7678FFA6FFD8FE9E96B3FE3FF924" box="[378,465,1683,1707]" name="Mexico" pageId="4" pageNumber="3">México</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFE84B19FFA6FFD8FE0596B3FD94F924" author="Cazier, M. A." box="[481,634,1683,1707]" pageId="4" pageNumber="3" pagination="231 - 309" refId="ref6746" refString="Cazier, M. A. 1954. A review of the Mexican tiger beetles of the genus Cicindela (Coleoptera: Cicindelidae). Bulletin of the American Museum of Natural History 103 (3): 231 - 309." type="journal article" year="1954">Cazier 1954</bibRefCitation>
|
||
). A more extensive study of the Mexican forms is needed;
|
||
<bibRefCitation id="EFE84B19FFA6FFD8FAA596B4FEE3F944" author="Murray, R. R." pageId="4" pageNumber="3" pagination="49 - 56" refId="ref7108" refString="Murray, R. R. 1979. The Cicindela fauna of Mexico: range extensions, additions, and ecological notes (Coleoptera: Cicindelidae). The Coleopterists Bulletin 33 (1): 49 - 56." type="journal article" year="1979">Murray (1979)</bibRefCitation>
|
||
reports the discovery of Mexican specimens which do not fit into any of the currently-recognized subspecific taxa. Even the status of the two subspecific taxa
|
||
<emphasis id="B90DEAFAFFA6FFD8FC7196F4FB9FF964" box="[917,1137,1747,1771]" italics="true" pageId="4" pageNumber="3">C. o. latemaculata</emphasis>
|
||
and
|
||
<emphasis id="B90DEAFAFFA6FFD8FB4E96F4FAA0F964" box="[1194,1358,1747,1771]" italics="true" pageId="4" pageNumber="3">C. o. juvenilis</emphasis>
|
||
can be questioned.
|
||
<bibRefCitation id="EFE84B19FFA6FFD8FEB796D3FE18F884" author="Cazier, M. A." box="[339,502,1779,1803]" pageId="4" pageNumber="3" pagination="231 - 309" refId="ref6746" refString="Cazier, M. A. 1954. A review of the Mexican tiger beetles of the genus Cicindela (Coleoptera: Cicindelidae). Bulletin of the American Museum of Natural History 103 (3): 231 - 309." type="journal article" year="1954">Cazier (1954)</bibRefCitation>
|
||
relied on the presence of wide elytral markings to separate
|
||
<emphasis id="B90DEAFAFFA6FFD8FB2096D4FA71F884" box="[1220,1439,1779,1803]" italics="true" pageId="4" pageNumber="3">C. o. latemaculata</emphasis>
|
||
from the other subspecies of
|
||
<taxonomicName id="4C794D6BFFA6FFD8FDF09734FD78F8A4" box="[532,662,1811,1835]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B90DEAFAFFA6FFD8FDF09734FD78F8A4" box="[532,662,1811,1835]" italics="true" pageId="4" pageNumber="3">C. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
. However, similar markings are present in individuals from
|
||
<collectingCountry id="F36E7678FFA6FFD8FA8F9733FA4EF8A4" box="[1387,1440,1811,1835]" name="United States of America" pageId="4" pageNumber="3">U.S.</collectingCountry>
|
||
populations of
|
||
<emphasis id="B90DEAFAFFA6FFD8FE979714FDD0F8C4" box="[371,574,1843,1867]" italics="true" pageId="4" pageNumber="3">C. o. santaclarae</emphasis>
|
||
(e.g.
|
||
<figureCitation id="13422A6DFFA6FFD8FD999713FCEDF8C4" box="[637,771,1843,1867]" captionStart="Figure 1-9" captionStartId="4.[192,276,1071,1093]" captionTargetBox="[192,1439,192,1041]" captionTargetId="figure-294@4.[192,1440,192,1041]" captionTargetPageId="4" captionText="Figure 1-9. Adult habitus photographs of pinned specimens of Cicindela obsoleta in NMNH. 1) Cicindela obsoleta obsoleta Say, male, Silver City, NM. 2) Cicindela obsoleta obsoleta, female, Carrizo Creek Picnic Area, Baca Co., CO. 3) Cicindela obsoleta obsoleta, male, Silver City, NM. 4) Cicindela obsoleta obsoleta X Cicindela obsoleta santaclarae, female, Silver City, NM. 5) Cicindela obsoleta santaclarae Bates, syntype female, Santa Clara, Chihuahua, México. 6) Cicindela obsoleta santaclarae, male, Ft. Wingate, NM. 7) Cicindela obsoleta santaclarae, female, Ft. Wingate, NM. 8) Cicindela obsoleta santaclarae, female, Ft. Wingate, NM. 9) Cicindela obsoleta santaclarae, male, Ft. Wingate, NM." figureDoi="http://doi.org/10.5281/zenodo.5167793" httpUri="https://zenodo.org/record/5167793/files/figure.png" pageId="4" pageNumber="3">Figure 7, 9</figureCitation>
|
||
). Thus,
|
||
<emphasis id="B90DEAFAFFA6FFD8FC879714FBAEF8C4" box="[867,1088,1843,1867]" italics="true" pageId="4" pageNumber="3">C. o. latemaculata</emphasis>
|
||
may ultimately prove to be a synonym of
|
||
<emphasis id="B90DEAFAFFA6FFD8FEB59774FDF2F8E4" box="[337,540,1875,1899]" italics="true" pageId="4" pageNumber="3">C. o. santaclarae</emphasis>
|
||
. Likewise,
|
||
<emphasis id="B90DEAFAFFA6FFD8FD419774FCA4F8E4" box="[677,842,1875,1899]" italics="true" pageId="4" pageNumber="3">C. o. juvenilis</emphasis>
|
||
differs from the other subspecies of
|
||
<taxonomicName id="4C794D6BFFA6FFD8FB1A9774FA6CF8E4" box="[1278,1410,1875,1899]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B90DEAFAFFA6FFD8FB1A9774FA6CF8E4" box="[1278,1410,1875,1899]" italics="true" pageId="4" pageNumber="3">C. obsoleta</emphasis>
|
||
</taxonomicName>
|
||
in lacking dense setae on the proepisterna and metepisterna (
|
||
<bibRefCitation id="EFE84B19FFA7FFD9FC669595FBF8FA42" author="Cazier, M. A." box="[898,1046,1461,1485]" pageId="5" pageNumber="4" pagination="231 - 309" refId="ref6746" refString="Cazier, M. A. 1954. A review of the Mexican tiger beetles of the genus Cicindela (Coleoptera: Cicindelidae). Bulletin of the American Museum of Natural History 103 (3): 231 - 309." type="journal article" year="1954">Cazier 1954</bibRefCitation>
|
||
), a significant morphological distinction which suggests that
|
||
<emphasis id="B90DEAFAFFA7FFD9FDC195F6FD23FA62" box="[549,717,1493,1517]" italics="true" pageId="5" pageNumber="4">C. o. juvenilis</emphasis>
|
||
may actually be a separate species. Additional material of these forms from additional collecting localities is needed in order to resolve these questions.
|
||
</paragraph>
|
||
<caption id="DF066660FFA7FFD9FF2494C4FC83FA05" ID-DOI="http://doi.org/10.5281/zenodo.5167797" ID-Zenodo-Dep="5167797" httpUri="https://zenodo.org/record/5167797/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,276,1252,1274]" targetBox="[193,1443,192,1226]" targetPageId="5">
|
||
<paragraph id="8BC636E8FFA7FFD9FF2494C4FC83FA05" blockId="5.[192,1440,1252,1418]" pageId="5" pageNumber="4">
|
||
<emphasis id="B90DEAFAFFA7FFD9FF2494C4FE8DFB76" bold="true" box="[192,355,1252,1274]" pageId="5" pageNumber="4">Figure 10-17.</emphasis>
|
||
Adult habitus photographs of pinned specimens of
|
||
<taxonomicName id="4C794D6BFFA7FFD9FC5594C4FB90FB76" box="[945,1150,1252,1273]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B90DEAFAFFA7FFD9FC5594C4FB90FB76" box="[945,1150,1252,1273]" italics="true" pageId="5" pageNumber="4">Cicindela obsoleta</emphasis>
|
||
</taxonomicName>
|
||
in NMNH.
|
||
<emphasis id="B90DEAFAFFA7FFD9FAE394C4FAC2FB75" bold="true" box="[1287,1324,1252,1274]" pageId="5" pageNumber="4">10)</emphasis>
|
||
<taxonomicName id="4C794D6BFFA7FFD9FAD194C4FE09FA99" authority="LeConte" authorityName="LeConte" authorityYear="1853" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="vulturina">
|
||
<emphasis id="B90DEAFAFFA7FFD9FAD194C4FE6AFA99" italics="true" pageId="5" pageNumber="4">Cicindela obsoleta vulturina</emphasis>
|
||
LeConte
|
||
</taxonomicName>
|
||
, female, Cisco, TX.
|
||
<emphasis id="B90DEAFAFFA7FFD9FD509521FD37FA98" bold="true" box="[692,729,1281,1303]" pageId="5" pageNumber="4">11)</emphasis>
|
||
<emphasis id="B90DEAFAFFA7FFD9FD399521FBFAFA99" box="[733,1044,1281,1302]" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C794D6BFFA7FFD9FD399521FBFEFA99" authorityName="LeConte" authorityYear="1853" box="[733,1040,1281,1302]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="vulturina">Cicindela obsoleta vulturina</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
female, Henrietta, TX.
|
||
<emphasis id="B90DEAFAFFA7FFD9FAEB9521FADAFA98" bold="true" box="[1295,1332,1281,1303]" pageId="5" pageNumber="4">12)</emphasis>
|
||
<emphasis id="B90DEAFAFFA7FFD9FADC9521FE63FABC" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C794D6BFFA7FFD9FADC9521FE67FABC" authorityName="LeConte" authorityYear="1853" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="vulturina">Cicindela obsoleta vulturina</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
male, Forestburg, TX.
|
||
<emphasis id="B90DEAFAFFA7FFD9FD63953EFD42FABB" bold="true" box="[647,684,1310,1332]" pageId="5" pageNumber="4">13)</emphasis>
|
||
<emphasis id="B90DEAFAFFA7FFD9FD55953EFC02FABC" box="[689,1004,1310,1331]" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C794D6BFFA7FFD9FD55953EFC06FABC" authorityName="LeConte" authorityYear="1853" box="[689,1000,1310,1331]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="vulturina">Cicindela obsoleta vulturina</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
male College Station, TX.
|
||
<emphasis id="B90DEAFAFFA7FFD9FAE9953EFADFFABB" bold="true" box="[1293,1329,1310,1332]" pageId="5" pageNumber="4">14)</emphasis>
|
||
<emphasis id="B90DEAFAFFA7FFD9FAD2953EFE7FFADF" italics="true" pageId="5" pageNumber="4">
|
||
<taxonomicName id="4C794D6BFFA7FFD9FAD2953EFE63FADF" authorityName="LeConte" authorityYear="1853" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="vulturina">Cicindela obsoleta vulturina</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
male, 2.5 miles SW Calico Rock, AR.
|
||
<emphasis id="B90DEAFAFFA7FFD9FCCB951BFCBAFADE" bold="true" box="[815,852,1339,1361]" pageId="5" pageNumber="4">15)</emphasis>
|
||
<taxonomicName id="4C794D6BFFA7FFD9FCBF951BFB00FADF" authority="Vogt" authorityName="Vogt" authorityYear="1949" box="[859,1262,1339,1360]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="neojuvenilis">
|
||
<emphasis id="B90DEAFAFFA7FFD9FCBF951BFB5CFADF" box="[859,1202,1339,1360]" italics="true" pageId="5" pageNumber="4">Cicindela obsoleta neojuvenilis</emphasis>
|
||
Vogt
|
||
</taxonomicName>
|
||
, paratype male, S. W. Hidalgo Co., TX.
|
||
<emphasis id="B90DEAFAFFA7FFD9FE5B9577FE0AFAE2" bold="true" box="[447,484,1367,1389]" pageId="5" pageNumber="4">16)</emphasis>
|
||
<taxonomicName id="4C794D6BFFA7FFD9FE0E9578FCAFFAE2" authorityName="Vogt" authorityYear="1949" box="[490,833,1368,1389]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="neojuvenilis">
|
||
<emphasis id="B90DEAFAFFA7FFD9FE0E9578FCAFFAE2" box="[490,833,1368,1389]" italics="true" pageId="5" pageNumber="4">Cicindela obsoleta neojuvenilis</emphasis>
|
||
</taxonomicName>
|
||
, paratype female, S.W. Hidalgo Co., TX.
|
||
<emphasis id="B90DEAFAFFA7FFD9FAEE9577FAC1FAE2" bold="true" box="[1290,1327,1367,1389]" pageId="5" pageNumber="4">17)</emphasis>
|
||
<taxonomicName id="4C794D6BFFA7FFD9FAD19578FE48FA06" authorityName="Vogt" authorityYear="1949" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="subSpecies" species="obsoleta" subSpecies="neojuvenilis">
|
||
<emphasis id="B90DEAFAFFA7FFD9FAD19578FE48FA06" italics="true" pageId="5" pageNumber="4">Cicindela obsoleta neojuvenilis</emphasis>
|
||
</taxonomicName>
|
||
, paratype female, S. W. Hidalgo Co., TX.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC636E8FFA7FFD9FF149635FB9DF902" blockId="5.[192,1441,1461,1677]" pageId="5" pageNumber="4">
|
||
The following key is offered to assist in the separation of the
|
||
<collectingCountry id="F36E7678FFA7FFD9FC599635FC1CF9A2" box="[957,1010,1557,1581]" name="United States of America" pageId="5" pageNumber="4">U.S.</collectingCountry>
|
||
subspecies of
|
||
<taxonomicName id="4C794D6BFFA7FFD9FB7D9636FA4EF9A2" authority="Say. Users" authorityName="Say. Users" box="[1177,1440,1557,1581]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="obsoleta">
|
||
<emphasis id="B90DEAFAFFA7FFD9FB7D9636FAF4F9A2" box="[1177,1306,1557,1581]" italics="true" pageId="5" pageNumber="4">C. obsoleta</emphasis>
|
||
Say. Users
|
||
</taxonomicName>
|
||
should be aware that putative intergrades between
|
||
<emphasis id="B90DEAFAFFA7FFD9FCDB9616FC0EF9C2" box="[831,992,1589,1613]" italics="true" pageId="5" pageNumber="4">
|
||
C. o.
|
||
<taxonomicName id="4C794D6BFFA7FFD9FC989615FC0EF9C2" box="[892,992,1589,1613]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="obsoleta">obsoleta</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B90DEAFAFFA7FFD9FBF99616FB05F9C2" box="[1053,1259,1589,1613]" italics="true" pageId="5" pageNumber="4">C. o. santaclarae</emphasis>
|
||
have been collected in
|
||
<collectingRegion id="49BDF80AFFA7FFD9FEC99676FE2CF9E2" box="[301,450,1621,1645]" country="United States of America" name="New Mexico" pageId="5" pageNumber="4">New Mexico</collectingRegion>
|
||
and
|
||
<collectingRegion id="49BDF80AFFA7FFD9FE189676FDABF9E2" box="[508,581,1622,1645]" country="United States of America" name="Texas" pageId="5" pageNumber="4">Texas</collectingRegion>
|
||
and may also occur in
|
||
<collectingRegion id="49BDF80AFFA7FFD9FCB39675FC56F9E2" box="[855,952,1621,1645]" country="United States of America" name="Arizona" pageId="5" pageNumber="4">Arizona</collectingRegion>
|
||
(see further discussion in the section on intergrades below). These intergrades will run to
|
||
<emphasis id="B90DEAFAFFA7FFD9FCFE9656FC0BF902" box="[794,997,1653,1677]" italics="true" pageId="5" pageNumber="4">C. o. santaclarae</emphasis>
|
||
in this key.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |