treatments-xml/data/53/68/CD/5368CD864BA15A4691858B647A50E114.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.366.6172" ID-PMC="PMC3890666" ID-Pensoft-Pub="1313-2970-366-1" ID-Pensoft-UUID="FFD8DC462108382BCB68FFC9FF97F235" ID-PubMed="24453569" ID-Zenodo-Dep="577560" ModsDocID="1313-2970-366-1" checkinTime="1451246487361" checkinUser="pensoft" docAuthor="Kolibac, Jiri" docDate="2013" docId="5368CD864BA15A4691858B647A50E114" docLanguage="en" docName="ZooKeys 366: 1-194" docOrigin="ZooKeys 366" docPubDate="2013-12-31" docSource="http://dx.doi.org/10.3897/zookeys.366.6172" docTitle="Paracalanthis Crowson 1970" docType="treatment" docVersion="3" id="FFD8DC462108382BCB68FFC9FF97F235" lastPageNumber="21" masterDocId="FFD8DC462108382BCB68FFC9FF97F235" masterDocTitle="Trogossitidae: A review of the beetle family, with a catalogue and keys" masterLastPageNumber="194" masterPageNumber="1" pageNumber="20" updateTime="1668157348086" updateUser="ExternalLinkService">
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<mods:title>Trogossitidae: A review of the beetle family, with a catalogue and keys</mods:title>
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<mods:namePart>Kolibac, Jiri</mods:namePart>
<mods:affiliation>Moravian Museum, Department of Entomology, Hviezdoslavova 29 a, 627 00 Brno, Czech Republic</mods:affiliation>
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<mods:date>2013</mods:date>
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<mods:number>2013-12-31</mods:number>
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<treatment ID-GBIF-Taxon="182235266" LSID="urn:lsid:plazi:treatment:5368CD864BA15A4691858B647A50E114" httpUri="http://treatment.plazi.org/id/5368CD864BA15A4691858B647A50E114" lastPageId="21" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection pageId="19" pageNumber="20" type="nomenclature">
<paragraph pageId="19" pageNumber="20">
Tribe
<taxonomicName LSID="5368CD86-4BA1-5A46-9185-8B647A50E114" authority="Lacordaire, 1854" authorityName="Crowson" authorityYear="1970" class="Insecta" family="Trogossitidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="tribe" tribe="Gymnochilini">Gymnochilini Lacordaire, 1854</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="reference_group">
<paragraph pageId="19" pageNumber="20">
<taxonomicName authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="19" pageNumber="20" rank="tribe" tribe="Gymnochilini">Gymnochilini</taxonomicName>
Lacordaire, J. T. 1854: 344.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="type genus">
<paragraph pageId="19" pageNumber="20">Type genus.</paragraph>
<paragraph pageId="19" pageNumber="20">
<taxonomicName class="Insecta" family="Trogossitidae" genus="Gymnochila" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="19" pageNumber="20">Gymnochila</emphasis>
</taxonomicName>
Erichson, 1844 (=
<taxonomicName authorityName="Dejean" authorityYear="1835" class="Insecta" family="Trogossitidae" genus="Gymnocheilis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="19" pageNumber="20">Gymnocheilis</emphasis>
</taxonomicName>
Dejean, 1835)
</paragraph>
<paragraph pageId="19" pageNumber="20">
Bouchard, P. et al. 2011: 57. Burakowski, B. et al. 1986: 118 (
<taxonomicName lsidName="" pageId="19" pageNumber="20" rank="subFamily" subFamily="Gymnochilinae">Gymnochilinae</taxonomicName>
).
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
, J. 2006: 119 (diagnosis, stat. n.).
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
, J. 2007a: 364.
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
, J. 2008: 118-119 (phylogeny). Leschen, R. A. B. &amp; Lackner, T. 2013: 283
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="remarks">
<paragraph pageId="19" pageNumber="20">Remarks.</paragraph>
<paragraph lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Five genera of
<taxonomicName authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="19" pageNumber="20" rank="tribe" tribe="Gymnochilini">Gymnochilini</taxonomicName>
(namely
<taxonomicName authorityName="Illiger" authorityYear="1807" class="Insecta" family="Trogossitidae" genus="Anacypta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">
<pageBreakToken pageId="20" pageNumber="21" start="start">Anacypta</pageBreakToken>
</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Dejean" authorityYear="1835" class="Insecta" family="Trogossitidae" genus="Gymnocheilis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Gymnocheilis</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Pascoe" authorityYear="1863" class="Insecta" family="Trogossitidae" genus="Narcisa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Narcisa</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Reitter" authorityYear="1876" class="Insecta" family="Trogossitidae" genus="Xenoglena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Xenoglena</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Hexapoda" family="Trogossitidae" genus="Leperina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Leperina</emphasis>
</taxonomicName>
) form, beyond doubt, a monophyletic group. Gymnochilins constitute an advanced group of
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="20" pageNumber="21" rank="subFamily" subFamily="Trogossitinae">Trogossitinae</taxonomicName>
, adapted to a predatory way of life. They are rapid flyers, dwelling on fallen logs and hunting for bostrichids, scolytids and other insects, strongly resembling the jewel beetles (
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Buprestidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="family">Buprestidae</taxonomicName>
) in their body shape and movement. Two distinctly separated pairs of eyes in most of them and their ability to jump (
<taxonomicName authorityName="Illiger" authorityYear="1807" class="Insecta" family="Trogossitidae" genus="Anacypta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Anacypta</emphasis>
</taxonomicName>
) characterize the tribe as one of the most advanced of all trogossitids. Some remarks about the independent status of the tribe
<taxonomicName authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="20" pageNumber="21" rank="tribe" tribe="Gymnochilini">Gymnochilini</taxonomicName>
with regard to
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="20" pageNumber="21" rank="tribe" tribe="Trogossitini">Trogossitini</taxonomicName>
are made below, in the section relating the latter tribe.
</paragraph>
<paragraph pageId="20" pageNumber="21">
The inclusion of
<taxonomicName authorityName="Reitter" authorityYear="1876" class="Hexapoda" family="Trogossitidae" genus="Phanodesta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Phanodesta</emphasis>
</taxonomicName>
from Juan Fernandez Isl. was more or less confirmed by two separate character analyses (
<bibRefCitation author="Kolibac, J" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="155" pageNumber="156" pagination="105 - 153" refId="B146" refString="Kolibac, J, 2006. A review of the Trogossitidae. Part 2: Larval morphology, phylogeny and taxonomy (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey 28: 105 - 153" title="A review of the Trogossitidae. Part 2: Larval morphology, phylogeny and taxonomy (Coleoptera, Cleroidea)." volume="28" year="2006">
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
2006
</bibRefCitation>
,
<bibRefCitation author="Kolibac, J" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="155" pageNumber="156" pagination="105 - 133" refId="B149" refString="Kolibac, J, 2008. Morphology, taxonomy and phylogeny of Phloiophilus edwardsi Stephens, 1830 (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey 30: 105 - 133" title="Morphology, taxonomy and phylogeny of Phloiophilus edwardsi Stephens, 1830 (Coleoptera, Cleroidea)." volume="30" year="2008">2008</bibRefCitation>
). However, the phylogeny of the genus was rendered unclear by the number of autapomorphies (e.g. winglessness, characteristic elytral structure) and separated distribution. I postulated a a sister group of
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Hexapoda" family="Trogossitidae" genus="Leperina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Leperina</emphasis>
</taxonomicName>
with a
<normalizedToken originalValue="“Gondwanan”">&quot;Gondwanan&quot;</normalizedToken>
distribution (Australia-Chile). Recently, a phylogeny and distribution of
<taxonomicName authorityName="Reitter" authorityYear="1876" class="Hexapoda" family="Trogossitidae" genus="Phanodesta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Phanodesta</emphasis>
</taxonomicName>
, together with descriptions and combinations of some more species from New Zealand and its vicinity have been addressed by
<bibRefCitation author="Leschen, RAB" journalOrPublisher="Systematic Entomology" pageId="156" pageNumber="157" pagination="278 - 304" publicationUrl="10.1111/j.1365-3113.2012.00661.x" refId="B184" refString="Leschen, RAB, Lackner, T, 2013. Gondwanan Gymnochilini (Coleoptera: Trogossitidae): generic concepts, review of New Zealand species and long-range Pacific dispersal. Systematic Entomology 38: 278 - 304, 10.1111/j.1365-3113.2012.00661.x" title="Gondwanan Gymnochilini (Coleoptera: Trogossitidae): generic concepts, review of New Zealand species and long-range Pacific dispersal." url="10.1111/j.1365-3113.2012.00661.x" volume="38" year="2013">Leschen and Lackner (2013)</bibRefCitation>
in detail. Two more genera occured in the gymnochiline clade in my second analysis (
<bibRefCitation author="Kolibac, J" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="155" pageNumber="156" pagination="105 - 133" refId="B149" refString="Kolibac, J, 2008. Morphology, taxonomy and phylogeny of Phloiophilus edwardsi Stephens, 1830 (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey 30: 105 - 133" title="Morphology, taxonomy and phylogeny of Phloiophilus edwardsi Stephens, 1830 (Coleoptera, Cleroidea)." volume="30" year="2008">
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
2008
</bibRefCitation>
):
<taxonomicName authorityName="Jakobson" authorityYear="1915" class="Insecta" family="Trogositidae" genus="Seidlitzella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Seidlitzella</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Insecta" family="Trogossitidae" genus="Melambia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Melambia</emphasis>
</taxonomicName>
. The first genus,
<taxonomicName authorityName="Jakobson" authorityYear="1915" class="Insecta" family="Trogositidae" genus="Seidlitzella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Seidlitzella</emphasis>
</taxonomicName>
, was considered related to the Palaearctic species of
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Hexapoda" family="Trogossitidae" genus="Leperina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Leperina</emphasis>
</taxonomicName>
, as also pointed out by
<bibRefCitation author="Schawaller, W" journalOrPublisher="Stuttgarter Beitraege zur Naturkunde, Serie A (Biologie)" pageId="161" pageNumber="162" pagination="1 - 9" refId="B272" refString="Schawaller, W, 1993. Taxonomie und Larvalmorphologie palaarktischer Leperina (Coleoptera: Trogossitidae). Stuttgarter Beitraege zur Naturkunde, Serie A (Biologie) 500: 1 - 9" title="Taxonomie und Larvalmorphologie palaarktischer Leperina (Coleoptera: Trogossitidae)." volume="500" year="1993">Schawaller (1993)</bibRefCitation>
. However, his formal synonymization of
<taxonomicName authorityName="Jakobson" authorityYear="1915" class="Insecta" family="Trogositidae" genus="Seidlitzella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Seidlitzella</emphasis>
</taxonomicName>
was not confirmed in a recent study by
<bibRefCitation author="Leschen, RAB" journalOrPublisher="Systematic Entomology" pageId="156" pageNumber="157" pagination="278 - 304" publicationUrl="10.1111/j.1365-3113.2012.00661.x" refId="B184" refString="Leschen, RAB, Lackner, T, 2013. Gondwanan Gymnochilini (Coleoptera: Trogossitidae): generic concepts, review of New Zealand species and long-range Pacific dispersal. Systematic Entomology 38: 278 - 304, 10.1111/j.1365-3113.2012.00661.x" title="Gondwanan Gymnochilini (Coleoptera: Trogossitidae): generic concepts, review of New Zealand species and long-range Pacific dispersal." url="10.1111/j.1365-3113.2012.00661.x" volume="38" year="2013">Leschen and Lackner (2013)</bibRefCitation>
who established the new genus
<taxonomicName authorityName="Leschen &amp; Lackner" authorityYear="2013" class="Insecta" family="Trogossitidae" genus="Kolibacia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Kolibacia</emphasis>
</taxonomicName>
for
<taxonomicName authorityName="Reitter" authorityYear="1889" class="Hexapoda" family="Trogossitidae" genus="Leperina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis italics="true" pageId="20" pageNumber="21">Leperina tibialis</emphasis>
</taxonomicName>
and
<taxonomicName class="Hexapoda" family="Trogossitidae" genus="Leperina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="squamulata">
<emphasis italics="true" pageId="20" pageNumber="21">Leperina squamulata</emphasis>
</taxonomicName>
instead.
</paragraph>
<paragraph pageId="20" pageNumber="21">
In both analyses by
<bibRefCitation author="Kolibac, J" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="155" pageNumber="156" pagination="105 - 153" refId="B146" refString="Kolibac, J, 2006. A review of the Trogossitidae. Part 2: Larval morphology, phylogeny and taxonomy (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey 28: 105 - 153" title="A review of the Trogossitidae. Part 2: Larval morphology, phylogeny and taxonomy (Coleoptera, Cleroidea)." volume="28" year="2006">
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
(2006
</bibRefCitation>
,
<bibRefCitation author="Kolibac, J" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="155" pageNumber="156" pagination="105 - 133" refId="B149" refString="Kolibac, J, 2008. Morphology, taxonomy and phylogeny of Phloiophilus edwardsi Stephens, 1830 (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey 30: 105 - 133" title="Morphology, taxonomy and phylogeny of Phloiophilus edwardsi Stephens, 1830 (Coleoptera, Cleroidea)." volume="30" year="2008">2008</bibRefCitation>
), the genera
<taxonomicName authorityName="Jakobson" authorityYear="1915" class="Insecta" family="Trogositidae" genus="Seidlitzella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Seidlitzella</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Insecta" family="Trogossitidae" genus="Melambia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Melambia</emphasis>
</taxonomicName>
were considered primitive or basal among the
<taxonomicName authorityName="Lacordaire" authorityYear="1854" lsidName="" pageId="20" pageNumber="21" rank="tribe" tribe="Gymnochilini">Gymnochilini</taxonomicName>
or
<taxonomicName authorityName="Latreille" authorityYear="1802" lsidName="" pageId="20" pageNumber="21" rank="tribe" tribe="Trogossitini">Trogossitini</taxonomicName>
. They were included in the trogossitins in my original tribe definition (
<bibRefCitation author="Kolibac, J" journalOrPublisher="Entomologica Basiliensia et Collectionis Frey" pageId="155" pageNumber="156" pagination="105 - 153" refId="B146" refString="Kolibac, J, 2006. A review of the Trogossitidae. Part 2: Larval morphology, phylogeny and taxonomy (Coleoptera, Cleroidea). Entomologica Basiliensia et Collectionis Frey 28: 105 - 153" title="A review of the Trogossitidae. Part 2: Larval morphology, phylogeny and taxonomy (Coleoptera, Cleroidea)." volume="28" year="2006">
<normalizedToken originalValue="Kolibáč">Kolibac</normalizedToken>
2006
</bibRefCitation>
). A comparison made specifically for the current shows the heterogenity of
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Insecta" family="Trogossitidae" genus="Melambia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Melambia</emphasis>
</taxonomicName>
species and the need for revision of the genus with respect to the systematic position of particular species. It cannot be excluded that there are some species of
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Insecta" family="Trogossitidae" genus="Melambia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Melambia</emphasis>
</taxonomicName>
congeneric with
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Insecta" family="Trogossitidae" genus="Alindria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Alindria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="key to genera">
<paragraph pageId="20" pageNumber="21">Key to genera</paragraph>
<paragraph lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<table inLine="true" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<tr pageId="20" pageNumber="21">
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">1</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">Head with 1 pair of eyes. Ventral part of cranium with long setae at sides. Radial cell triangular and moved down, or reduced. Elytra with conspicuous carinae</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">2</td>
</tr>
<tr pageId="20" pageNumber="21">
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">-</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">Head with 2 pairs of eyes. Ventral part of cranium without long setae at sides. Radial cell mostly oblong. Elytral carinae reduced or inconspicuous</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">5</td>
</tr>
<tr pageId="20" pageNumber="21">
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">2</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">Body bare, without setae. Window punctures absent. Unicolorous, black species</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">
<taxonomicName authorityName="Jakobson" authorityYear="1915" class="Insecta" family="Trogositidae" genus="Seidlitzella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Seidlitzella</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="20" pageNumber="21">
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">-</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">Body surface with vestiture consisting of scales or setae or both. Window punctures absent or present. Dorsal side brown or black-brown, with colour patterns formed by vestiture</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">3</td>
</tr>
<tr pageId="20" pageNumber="21">
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">3</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">Dorsal vestiture consisting of scales. Elytral carinae not beaded; intercarinal space punctate; window punctures present. Mucro absent</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">4</td>
</tr>
<tr pageId="20" pageNumber="21">
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">-</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">Dorsal vestiture consisting of setae, scales or both. Elytral carinae usually beaded; intercarinal space apunctate; window punctures absent. Mucro absent or weakly developed</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">
<taxonomicName authorityName="Reitter" authorityYear="1876" class="Hexapoda" family="Trogossitidae" genus="Phanodesta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Phanodesta</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="20" pageNumber="21">
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">4</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">Elytral intercarinal space bipunctate; window punctures simple. Protibial edge smooth. Mandibles with mola</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">
<taxonomicName authorityName="Leschen &amp; Lackner" authorityYear="2013" class="Insecta" family="Trogossitidae" genus="Kolibacia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Kolibacia</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="20" pageNumber="21">
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">-</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">Elytral intercarinal space multipunctate; window punctures tuberculate. Protibial edge spinate. Mandibles without mola</td>
<td colspan="1" pageId="20" pageNumber="21" rowspan="1">
<taxonomicName authorityName="Erichson" authorityYear="1844" class="Hexapoda" family="Trogossitidae" genus="Leperina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="20" pageNumber="21">Leperina</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="21" pageNumber="22">
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">
<pageBreakToken pageId="21" pageNumber="22" start="start">5</pageBreakToken>
</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">Body rather flat, compact; smaller species (about 4-8 mm)</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">6</td>
</tr>
<tr pageId="21" pageNumber="22">
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">-</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">Body rather cylindrical, elongate; larger species (about 6-20 mm)</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">7</td>
</tr>
<tr pageId="21" pageNumber="22">
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">6</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">Body perfectly covered with large scales</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">
<taxonomicName authorityName="Pascoe" authorityYear="1863" class="Insecta" family="Trogossitidae" genus="Narcisa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Narcisa</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="21" pageNumber="22">
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">-</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">Body without scales or pubescence, with metallic lustre</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">
<taxonomicName authorityName="Illiger" authorityYear="1807" class="Insecta" family="Trogossitidae" genus="Anacypta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anacypta</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="21" pageNumber="22">
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">7</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">Body with colour pattern composed of scales and short thick setae; antennal club large</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">
<taxonomicName authorityName="Dejean" authorityYear="1835" class="Insecta" family="Trogossitidae" genus="Gymnocheilis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Gymnocheilis</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="21" pageNumber="22">
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">-</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">Body without scales, spots on pronotum and elytra formed by short setae; antennal club smaller</td>
<td colspan="1" pageId="21" pageNumber="22" rowspan="1">
<taxonomicName authorityName="Reitter" authorityYear="1876" class="Insecta" family="Trogossitidae" genus="Xenoglena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Xenoglena</emphasis>
</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>