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<document id="FCA0DDDA4AD8CF3ECC14B13DBC45C2A8" ID-CLB-Dataset="45651" ID-DOI="10.11646/zootaxa.3802.4.5" ID-GBIF-Dataset="71e8a84e-1a9b-4849-9a09-ec2023643364" ID-ISSN="1175-5326" ID-Zenodo-Dep="249605" ID-ZooBank="4095CA09-8EA4-4941-8286-32E95F0206AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460427480340" checkinUser="plazi" docAuthor="Carmona, Leila, Pola, Marta, Gosliner, Terrence M. &amp; Cervera, Juan Lucas" docDate="2014" docId="ED36FA51A03EFF84FF1BFC35E19EF886" docLanguage="en" docName="zt03802p514.pdf" docOrigin="Zootaxa 3802 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baeolidia moebii Bergh 1888" docType="treatment" docVersion="9" lastPageNumber="483" masterDocId="110F8229A03CFF82FF8CFFB7E275FFB7" masterDocTitle="Review of Baeolidia, the largest genus of Aeolidiidae (Mollusca: Nudibranchia), with the description of five new species" masterLastPageNumber="514" masterPageNumber="477" pageNumber="479" updateTime="1698330923557" updateUser="ExternalLinkService">
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<mods:title id="73331EC825AE66F262C26DAC4EF5B694">Review of Baeolidia, the largest genus of Aeolidiidae (Mollusca: Nudibranchia), with the description of five new species</mods:title>
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<mods:namePart id="19C523AC9834B71038EC987986DF1A6F">Carmona, Leila</mods:namePart>
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<mods:namePart id="4602E3E5F3B4B5806B0E9C954419DA5C">Cervera, Juan Lucas</mods:namePart>
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<mods:date id="593E764B9571E72CEAEDB7D6DC456A55">2014</mods:date>
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<treatment id="ED36FA51A03EFF84FF1BFC35E19EF886" ID-DOI="http://doi.org/10.5281/zenodo.6134190" ID-GBIF-Taxon="119429041" ID-Zenodo-Dep="6134190" LSID="urn:lsid:plazi:treatment:ED36FA51A03EFF84FF1BFC35E19EF886" httpUri="http://treatment.plazi.org/id/ED36FA51A03EFF84FF1BFC35E19EF886" lastPageId="6" lastPageNumber="483" pageId="2" pageNumber="479">
<subSubSection id="2D8518CCA03EFF80FF1BFC35E38DFC09" pageId="2" pageNumber="479" type="nomenclature">
<paragraph id="65204B47A03EFF80FF1BFC35E075FC2B" blockId="2.[151,512,898,958]" box="[151,512,898,924]" pageId="2" pageNumber="479">
<heading id="3E68FC2BA03EFF80FF1BFC35E075FC2B" bold="true" box="[151,512,898,924]" fontSize="11" level="1" pageId="2" pageNumber="479" reason="1">
<taxonomicName id="A29F30C4A03EFF80FF1BFC35E075FC2B" ID-CoL="KCLS" authority="Bergh, 1888" authorityName="Bergh" authorityYear="1888" box="[151,512,898,924]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="479" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03EFF80FF1BFC35E075FC2B" bold="true" box="[151,512,898,924]" pageId="2" pageNumber="479">
<emphasis id="57EB9755A03EFF80FF1BFC35E311FC2B" bold="true" box="[151,356,898,924]" italics="true" pageId="2" pageNumber="479">Baeolidia moebii</emphasis>
<bibRefCitation id="010E36B6A03EFF80FEE6FC35E075FC2B" author="Bergh" box="[362,512,898,924]" pageId="2" pageNumber="479" refString="Bergh, L. S. R. (1888) Malacologische Untersuchungen. In: Semper, C. (Ed.), Reisen im Archipel der Philippinen. Zweiter Theil, Wissenschaftliche Resultate. Band 2, Heft 16, pp. 755 - 814." type="journal article" year="1888">Bergh, 1888</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="65204B47A03EFF80FF1BFC12E38DFC09" blockId="2.[151,512,898,958]" box="[151,504,933,958]" pageId="2" pageNumber="479">
(
<figureCitation id="FDA457C2A03EFF80FF13FC12E299FC09" box="[159,236,933,958]" captionStart="FIGURE 2" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[151,1436,373,1863]" captionTargetId="figure@3.[151,1436,373,1865]" captionTargetPageId="3" captionText="FIGURE 2. Photographs of the living animals. A, B, Baeolidia moebii: (A) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177602; (B) specimen from Mozambique, photo by Yara Tibiriça, MNCN 15.05 / 63447; (C) Baeolidia macleayi, specimen from Australia, photo by Terrence M. Gosliner, CASIZ 71923; (D-E) Baeolidia japonica: (D) specimen from Japan, photo by Akira Kawahara, CASIZ 1814520; (E) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 186795; (F) Baeolidia ransoni, specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 186208." httpUri="https://zenodo.org/record/249607/files/figure.png" pageId="2" pageNumber="479">Figs. 2</figureCitation>
A–B, 3A, 4A–B, 5, 6A)
</paragraph>
</subSubSection>
<subSubSection id="2D8518CCA03EFF80FF1BFC5AE178FBC9" pageId="2" pageNumber="479" type="reference_group">
<paragraph id="65204B47A03EFF80FF1BFC5AE398FB95" blockId="2.[151,992,1005,1150]" pageId="2" pageNumber="479">
<treatmentCitationGroup id="458F6C69A03EFF80FF1BFC5AE398FB95" pageId="2" pageNumber="479">
<taxonomicName id="A29F30C4A03EFF80FF1BFC5AE388FBB4" ID-CoL="KCLS" authority="Bergh, 1888: 778" authorityName="Bergh" authorityPageNumber="778" authorityYear="1888" box="[151,509,1005,1027]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="479" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03EFF80FF1BFC5AE330FBB4" box="[151,325,1005,1027]" italics="true" pageId="2" pageNumber="479">Baeolidia moebii</emphasis>
<treatmentCitation id="E43E6D56A03EFF80FEC7FC5AE388FBB4" author="Bergh" box="[331,509,1005,1027]" page="778" pageId="2" pageNumber="479" year="1888">
<bibRefCitation id="010E36B6A03EFF80FEC7FC5AE3BEFBB4" author="Bergh" box="[331,459,1005,1027]" pageId="2" pageNumber="479" refString="Bergh, L. S. R. (1888) Malacologische Untersuchungen. In: Semper, C. (Ed.), Reisen im Archipel der Philippinen. Zweiter Theil, Wissenschaftliche Resultate. Band 2, Heft 16, pp. 755 - 814." type="journal article" year="1888">Bergh, 1888</bibRefCitation>
: 778
</treatmentCitation>
</taxonomicName>
, pl. LXXIX,
<figureCitation id="FDA457C2A03EFF80FD0FFC5AE088FBB4" box="[643,765,1005,1027]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStart-5="FIGURE 15" captionStart-6="FIGURE 16" captionStartId-0="18.[151,250,1811,1833]" captionStartId-1="19.[151,250,1706,1728]" captionStartId-2="22.[151,250,1811,1833]" captionStartId-3="24.[151,250,1384,1406]" captionStartId-4="25.[151,250,1198,1220]" captionStartId-5="27.[151,250,1869,1891]" captionStartId-6="28.[151,250,1201,1223]" captionTargetBox-0="[151,1436,193,1789]" captionTargetBox-1="[151,1436,193,1683]" captionTargetBox-2="[151,1436,193,1789]" captionTargetBox-3="[151,1436,373,1362]" captionTargetBox-4="[351,1236,512,1132]" captionTargetBox-5="[289,1324,452,1834]" captionTargetBox-6="[155,1432,193,1177]" captionTargetId-0="figure@18.[151,1436,193,1789]" captionTargetId-1="figure@19.[151,1436,193,1685]" captionTargetId-2="figure@22.[151,1436,193,1789]" captionTargetId-3="figure@24.[151,1436,373,1363]" captionTargetId-4="figure@25.[323,1263,483,1176]" captionTargetId-5="figure@27.[231,1355,423,1851]" captionTargetId-6="figure@28.[155,1432,193,1179]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="22" captionTargetPageId-3="24" captionTargetPageId-4="25" captionTargetPageId-5="27" captionTargetPageId-6="28" captionText-0="FIGURE 10. Photographs of the living animals. (A) Baeolidia rieae sp. nov., specimen from Japan, photo by Jum Imamoto, CASIZ 184525; (B – C) Baeolidia variabilis sp. nov.: (B) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 187741; (C) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177715." captionText-1="FIGURE 11. Scanning electron photographs. A, B, Baeolidia rieae sp. nov. (CASIZ 184525); (A) Detailed view of the jaw; (B) Radular teeth; (C – D) Baeolidia variabilis sp. nov. (CASIZ 177715); (C) Detailed view of the masticatory border; (D) Radular teeth; (E-F) Baeolidia lunaris sp. nov. (CASIZ 099221); (E) Detailed view of the masticatory border; (F) Radular teeth. Scale bars: A, 30 µm; B, 100 µm; C, 2 µm; D, 10 µm; E, 10 µm; F, 20 µm." captionText-2="FIGURE 12. Photographs of the living animals. (A) Baeolidia lunaris sp. nov., specimen from Tanzania, photo by Terrence M. Gosliner, CASIZ 099221; (B) Baeolidia gracilis sp. nov., specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 083766; (D) Baeolidia scottjohnsoni sp. nov., specimen from Marshall Islands, photo by Scott Johnson, CASIZ 184503." captionText-3="FIGURE 13. Scanning electron photographs. (A – B) Baeolidia gracilis sp. nov. (CASIZ 083766); (A) Detailed view of the masticatory border; (B) Radular teeth; (C – D) Baeolidia scottjohnsoni sp. nov. (CASIZ 184503); (C) Detailed view of the masticatory border; (D) Radular teeth. Scale bars: A, 3 µm; B, 10 µm; C, 50 µm; D, 100 µm." captionText-4="FIGURE 14. Reproductive system. (A) Baeolidia gracilis sp. nov., specimen from the Philippines (CASIZ 083766); (B) Baeolidia scottjohnsoni sp. nov., specimen from Marshall Islands (CASIZ 184503). Scale bars: A, 0.5 mm; B, 1 mm. Abbreviations: am, ampulla; fgm, female gland mass; ps, penial sac; rs, receptaculum seminis; v, vagina; vd, vas deferens." captionText-5="FIGURE 15. Schematic representation of differences of body shape among the genera Spurilla (A), drawing of Ascensión Cespedosa; Berghia (B), drawing of Ascensión Cespedosa; Limenandra (C), taken from Haefelfinger &amp; Stamm (1958); and Baeolidia (D), taken from Rudman (1982)." captionText-6="FIGURE 16. Comparison of the radular morphology among the genera Spurilla (A), Berghia (B), Limenandra (C) and Baeolidia (D)." httpUri-0="https://zenodo.org/record/249614/files/figure.png" httpUri-1="https://zenodo.org/record/249615/files/figure.png" httpUri-2="https://zenodo.org/record/249616/files/figure.png" httpUri-3="https://zenodo.org/record/249617/files/figure.png" httpUri-4="https://zenodo.org/record/249618/files/figure.png" httpUri-5="https://zenodo.org/record/249619/files/figure.png" httpUri-6="https://zenodo.org/record/249620/files/figure.png" pageId="2" pageNumber="479">Figs. 10–16</figureCitation>
, pl. LXXX,
<figureCitation id="FDA457C2A03EFF80FCF6FC5AE1AEFBB4" box="[890,987,1005,1027]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,984,1006]" captionStartId-1="3.[151,250,1886,1908]" captionStartId-2="4.[151,250,1459,1481]" captionStartId-3="5.[151,250,1706,1728]" captionTargetBox-0="[183,1403,519,961]" captionTargetBox-1="[151,1436,373,1863]" captionTargetBox-2="[354,1221,198,1422]" captionTargetBox-3="[151,1436,193,1683]" captionTargetId-0="figure@1.[183,1403,519,962]" captionTargetId-1="figure@3.[151,1436,373,1865]" captionTargetId-2="figure@4.[336,1251,193,1437]" captionTargetId-3="figure@5.[151,1436,193,1685]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Phylogeny of Baeolidia, based on Carmona et al. (2013). Names on right side of vertical bars refer to revised classification." captionText-1="FIGURE 2. Photographs of the living animals. A, B, Baeolidia moebii: (A) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177602; (B) specimen from Mozambique, photo by Yara Tibiriça, MNCN 15.05 / 63447; (C) Baeolidia macleayi, specimen from Australia, photo by Terrence M. Gosliner, CASIZ 71923; (D-E) Baeolidia japonica: (D) specimen from Japan, photo by Akira Kawahara, CASIZ 1814520; (E) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 186795; (F) Baeolidia ransoni, specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 186208." captionText-2="FIGURE 3. Different papillate rhinophores in Baeolidia: (A) from Rudman (1982); (B) from Gosliner (1985). Scale bars: A, 3 mm; B, 1 mm; C, 1.2 mm; D, 1.5 mm." captionText-3="FIGURE 4. Scanning electron photographs. (A – B) Baeolidia moebii (CASIZ 180327): (A) Detailed view of the masticatory border; (B) Radular teeth; (C – D) Baeolidia macleayi (CASIZ 195247): (C) Detailed view of the masticatory border; (D) Radular teeth; (E – F) Baeolidia japonica (CASIZ 181357); (E) Detailed view of the masticatory border; (F) Radular teeth. Scale bars: A, 10 µm; B, 30 µm; C-D, 10 µm; E, 100 µm; F, 150 µm." httpUri-0="https://zenodo.org/record/249606/files/figure.png" httpUri-1="https://zenodo.org/record/249607/files/figure.png" httpUri-2="https://zenodo.org/record/249608/files/figure.png" httpUri-3="https://zenodo.org/record/249609/files/figure.png" pageId="2" pageNumber="479">Figs. 1–4</figureCitation>
.
<taxonomicName id="A29F30C4A03EFF80FF1BFBBBE39CFB95" ID-CoL="68JS4" authority="Eliot, 1903: 252" authorityName="Eliot" authorityPageNumber="252" authorityYear="1903" box="[151,489,1036,1058]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="479" phylum="Mollusca" rank="species" species="major">
<emphasis id="57EB9755A03EFF80FF1BFBBBE349FB95" box="[151,316,1036,1058]" italics="true" pageId="2" pageNumber="479">Baeolidia major</emphasis>
<treatmentCitation id="E43E6D56A03EFF80FECEFBBBE39CFB95" author="Eliot" box="[322,489,1036,1058]" page="252" pageId="2" pageNumber="479" year="1903">
<bibRefCitation id="010E36B6A03EFF80FECEFBBBE3C3FB95" author="Eliot" box="[322,438,1036,1058]" pageId="2" pageNumber="479" refString="Eliot, C. N. E. (1903) On some nudibranchs from east Africa and Zanzibar, part II. Proceedings of the Zoological Society of London, 1, 250 - 257." type="journal article" year="1903">Eliot, 1903</bibRefCitation>
: 252
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="65204B47A03EFF80FF1BFB9DE01CFBF7" blockId="2.[151,992,1005,1150]" box="[151,617,1066,1088]" pageId="2" pageNumber="479">
<treatmentCitationGroup id="458F6C69A03EFF80FF1BFB9DE01CFBF7" box="[151,617,1066,1088]" pageId="2" pageNumber="479">
<taxonomicName id="A29F30C4A03EFF80FF1BFB9DE010FBF7" ID-CoL="5FW92" authority="Baba, 1937: 335" authorityName="Baba" authorityPageNumber="335" authorityYear="1937" box="[151,613,1066,1088]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="479" phylum="Mollusca" rank="subSpecies" species="major" subSpecies="amakusana">
<emphasis id="57EB9755A03EFF80FF1BFB9DE3C3FBF7" box="[151,438,1066,1088]" italics="true" pageId="2" pageNumber="479">Baeolidia major amakusana</emphasis>
<treatmentCitation id="E43E6D56A03EFF80FE30FB9DE010FBF7" author="Baba" box="[444,613,1066,1088]" page="335" pageId="2" pageNumber="479" year="1937">
<bibRefCitation id="010E36B6A03EFF80FE30FB9DE046FBF7" author="Baba" box="[444,563,1066,1088]" pageId="2" pageNumber="479" refString="Baba, K. (1937) Opisthobranchia of Japan (II). Journal of the Department of Agriculture, Kyushu Imperial University, 5, 289 - 344." type="journal article" year="1937">Baba, 1937</bibRefCitation>
: 335
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="65204B47A03EFF80FF1BFBFEE178FBC9" blockId="2.[151,992,1005,1150]" pageId="2" pageNumber="479">
<taxonomicName id="A29F30C4A03EFF80FF1BFBFEE3C3FBE8" ID-CoL="68JS4" authority="Eliot, 1903" authorityName="Eliot" authorityYear="1903" box="[151,438,1097,1119]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="479" phylum="Mollusca" rank="species" species="major">
<emphasis id="57EB9755A03EFF80FF1BFBFEE35EFBE8" box="[151,299,1097,1119]" italics="true" pageId="2" pageNumber="479">Berghia major</emphasis>
(
<bibRefCitation id="010E36B6A03EFF80FEBBFBFEE3D8FBE8" author="Eliot" box="[311,429,1097,1119]" pageId="2" pageNumber="479" refString="Eliot, C. N. E. (1903) On some nudibranchs from east Africa and Zanzibar, part II. Proceedings of the Zoological Society of London, 1, 250 - 257." type="journal article" year="1903">Eliot, 1903</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="010E36B6A03EFF80FE33FBFEE02FFBE8" author="Edmunds" box="[447,602,1097,1119]" pageId="2" pageNumber="479" refString="Edmunds, M. (1969) Opisthobranchiate Mollusca from Tanzania. I. Eolidacea (Eubranchidae and Aeolidiidae). Proceedings of the Malacological Society of London, 38, 451 - 469." type="journal article" year="1969">Edmunds 1969</bibRefCitation>
, 467,
<figureCitation id="FDA457C2A03EFF80FD1BFBFEE094FBE8" box="[663,737,1097,1119]" captionStart="FIGURE 10" captionStartId="18.[151,250,1811,1833]" captionTargetBox="[151,1436,193,1789]" captionTargetId="figure@18.[151,1436,193,1789]" captionTargetPageId="18" captionText="FIGURE 10. Photographs of the living animals. (A) Baeolidia rieae sp. nov., specimen from Japan, photo by Jum Imamoto, CASIZ 184525; (B – C) Baeolidia variabilis sp. nov.: (B) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 187741; (C) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177715." httpUri="https://zenodo.org/record/249614/files/figure.png" pageId="2" pageNumber="479">Fig. 10</figureCitation>
.
<taxonomicName id="A29F30C4A03EFF80FF1BFBDFE3C2FBC9" ID-CoL="4ZDW8" authority="Eliot, 1903" authorityName="Eliot" authorityYear="1903" box="[151,439,1128,1150]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="2" pageNumber="479" phylum="Mollusca" rank="species" species="major">
<emphasis id="57EB9755A03EFF80FF1BFBDFE35EFBC9" box="[151,299,1128,1150]" italics="true" pageId="2" pageNumber="479">Spurilla major</emphasis>
(
<bibRefCitation id="010E36B6A03EFF80FEB5FBDFE3DAFBC9" author="Eliot" box="[313,431,1128,1150]" pageId="2" pageNumber="479" refString="Eliot, C. N. E. (1903) On some nudibranchs from east Africa and Zanzibar, part II. Proceedings of the Zoological Society of London, 1, 250 - 257." type="journal article" year="1903">Eliot, 1903</bibRefCitation>
)
</taxonomicName>
:
<bibRefCitation id="010E36B6A03EFF80FE4CFBDFE024FBC9" author="Rudman" box="[448,593,1128,1150]" pageId="2" pageNumber="479" refString="Rudman, W. B. (1982) The taxonomy and biology of further aeolidacean and arminacean nudibranch molluscs with symbiotic zooxanthellae. Zoological Journal of the Linnean Society, 74, 147 - 196. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1982. tb 01146. x" type="journal article" year="1982">Rudman 1982</bibRefCitation>
, 169,
<figureCitation id="FDA457C2A03EFF80FD02FBDFE0AAFBC9" box="[654,735,1128,1150]" captionStart="FIGURE 18" captionStartId="30.[151,250,1519,1541]" captionTargetBox="[151,1427,195,1497]" captionTargetId="figure@30.[151,1436,195,1498]" captionTargetPageId="30" captionText="FIGURE 18. Map of the range of the different species of Spurilla, Berghia, Limenandra, and Baeolidia. A. Berghia and Limenandra. B. Spurilla and Baeolidia. Abbreviation: Isolated records of Baeolidia cryporos (star) and Spurilla braziliana (square)." httpUri="https://zenodo.org/record/249621/files/figure.png" pageId="2" pageNumber="479">Figs. 18</figureCitation>
–19.
</paragraph>
</subSubSection>
<subSubSection id="2D8518CCA03EFF87FF1BFB12E1F1F87F" lastPageId="5" lastPageNumber="482" pageId="2" pageNumber="479" type="materials_examined">
<paragraph id="65204B47A03EFF80FF1BFB12E176FB09" blockId="2.[151,1437,1189,2006]" box="[151,771,1189,1214]" pageId="2" pageNumber="479">
<emphasis id="57EB9755A03EFF80FF1BFB12E340FB09" bold="true" box="[151,309,1189,1214]" pageId="2" pageNumber="479">
<typeStatus id="BA24F5E5A03EFF80FF1BFB12E2A7FB09" box="[151,210,1189,1214]" pageId="2" pageNumber="479">Type</typeStatus>
locality.
</emphasis>
Grand Bay, Rodrigues
<collectingCountry id="1D880BD7A03EFF80FDB0FB12E0F6FB09" box="[572,643,1189,1214]" name="Iceland" pageId="2" pageNumber="479">Island</collectingCountry>
,
<collectingCountry id="1D880BD7A03EFF80FD03FB12E175FB09" box="[655,768,1189,1214]" name="Mauritius" pageId="2" pageNumber="479">Mauritius</collectingCountry>
.
</paragraph>
<paragraph id="65204B47A03EFF80FF4BFB7EE646FAB1" blockId="2.[151,1437,1189,2006]" pageId="2" pageNumber="479">
<emphasis id="57EB9755A03EFF80FF4BFB7EE303FB55" bold="true" box="[199,374,1225,1250]" pageId="2" pageNumber="479">
<typeStatus id="BA24F5E5A03EFF80FF4BFB7EE377FB55" box="[199,258,1225,1250]" pageId="2" pageNumber="479">Type</typeStatus>
material.
</emphasis>
To our knowledge no
<typeStatus id="BA24F5E5A03EFF80FDFAFB7DE0D3FB55" box="[630,678,1226,1250]" pageId="2" pageNumber="479">type</typeStatus>
material remains. We designate the specimen
<collectionCode id="038ED382A03EFF80FB38FB7DE77DFB55" LSID="urn:lsid:biocol.org:col:35275" box="[1204,1288,1226,1250]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="2" pageNumber="479">MNCN</collectionCode>
15.05/63447 from
<collectingCountry id="1D880BD7A03EFF80FF58FB5AE31DFAB1" box="[212,360,1261,1286]" name="Mozambique" pageId="2" pageNumber="479">Mozambique</collectingCountry>
as the
<typeStatus id="BA24F5E5A03EFF80FE3AFB59E07AFAB1" box="[438,527,1262,1286]" pageId="2" pageNumber="479" type="neotype">neotype</typeStatus>
in order to avoid confusion with similar species.
</paragraph>
<paragraph id="65204B47A03EFF80FF4BFAA6E07FF925" blockId="2.[151,1437,1189,2006]" pageId="2" pageNumber="479">
<emphasis id="57EB9755A03EFF80FF4BFAA6E3C1FA9D" bold="true" box="[199,436,1297,1322]" pageId="2" pageNumber="479">Material examined.</emphasis>
<typeStatus id="BA24F5E5A03EFF80FE33FAA5E057FA9D" box="[447,546,1298,1322]" pageId="2" pageNumber="479" type="neotype">Neotype</typeStatus>
:
<collectionCode id="038ED382A03EFF80FDBDFAA5E0F0FA9D" LSID="urn:lsid:biocol.org:col:35275" box="[561,645,1298,1322]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="2" pageNumber="479">MNCN</collectionCode>
15.05/63447, one specimen, dissected,
<quantity id="A267E6A2A03EFF80FBD4FAA5E6DEFA9E" box="[1112,1195,1298,1322]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="479" unit="mm" value="18.0">18 mm</quantity>
in length preserved,
<collectingCountry id="1D880BD7A03EFF80FF1BFA82E35BFAF9" box="[151,302,1333,1358]" name="Mozambique" pageId="2" pageNumber="479">Mozambique</collectingCountry>
, Zavora Bay, collected by Yara Tibiriça,
<date id="11216D87A03EFF80FCBEFA81E1A1FAF9" box="[818,980,1334,1358]" pageId="2" pageNumber="479" value="2012-01">January 2012</date>
. Other material:
<collectionCode id="038ED382A03EFF80FB25FA81E68FFAF9" box="[1193,1274,1334,1358]" pageId="2" pageNumber="479">CASIZ</collectionCode>
177602, one specimen, dissected,
<quantity id="A267E6A2A03EFF80FE04FAEDE3ACFAC6" box="[392,473,1370,1394]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="479" unit="mm" value="10.0">10 mm</quantity>
in length preserved,
<collectingCountry id="1D880BD7A03EFF80FD41FAEEE13BFAC5" box="[717,846,1369,1394]" name="Philippines" pageId="2" pageNumber="479">Philippines</collectingCountry>
, Luzon
<collectingCountry id="1D880BD7A03EFF80FC20FAEEE186FAC5" box="[940,1011,1369,1394]" name="Iceland" pageId="2" pageNumber="479">Island</collectingCountry>
, Batangas Bay, Calumpan Peninsula, collected by Terrence M. Gosliner,
<date id="11216D87A03EFF80FDBCFACAE0A3FA21" box="[560,726,1405,1430]" pageId="2" pageNumber="479" value="2008-04-16">16 April 2008</date>
;
<collectionCode id="038ED382A03EFF80FD68FAC9E140FA21" box="[740,821,1406,1430]" pageId="2" pageNumber="479">CASIZ</collectionCode>
157017, two specimens, dissected, 25 and
<quantity id="A267E6A2A03EFF80FAA7FAC9E708FA22" box="[1323,1405,1406,1430]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="479" unit="mm" value="24.0">24 mm</quantity>
in length preserved,
<collectingCountry id="1D880BD7A03EFF80FEEEFA16E396FA0D" box="[354,483,1441,1466]" name="Philippines" pageId="2" pageNumber="479">Philippines</collectingCountry>
, Luzon
<collectingCountry id="1D880BD7A03EFF80FDCCFA16E0F2FA0D" box="[576,647,1441,1466]" name="Iceland" pageId="2" pageNumber="479">Island</collectingCountry>
, Batangas Province, Balayan Bay, Seafari Beach, collected by Ángel Valdés and Mary James Adams,
<date id="11216D87A03EFF80FDA8FA71E0BFFA69" box="[548,714,1478,1502]" pageId="2" pageNumber="479" value="2001-05-12">12 May 2001</date>
;
<collectionCode id="038ED382A03EFF80FD50FA71E158FA69" box="[732,813,1478,1502]" pageId="2" pageNumber="479">CASIZ</collectionCode>
186211, one specimen, dissected,
<quantity id="A267E6A2A03EFF80FB5DFA71E753FA6A" box="[1233,1318,1478,1502]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="479" unit="mm" value="10.0">10 mm</quantity>
in length preserved,
<collectingCountry id="1D880BD7A03EFF80FE96FA5EE3EEF9B5" box="[282,411,1513,1538]" name="Philippines" pageId="2" pageNumber="479">Philippines</collectingCountry>
, Luzon, Batangas Province, Calumpan Peninsula, Batangas Bay, collected by Alicia Hermosillo, 0
<date id="11216D87A03EFF80FEB8F9B9E3B1F991" box="[308,452,1550,1574]" pageId="2" pageNumber="479" value="2011-05-04">4 May 2011</date>
;
<collectionCode id="038ED382A03EFF80FE58F9B9E050F991" box="[468,549,1550,1574]" pageId="2" pageNumber="479">CASIZ</collectionCode>
<date id="11216D87A03EFF80FDA3F9B9E0F3F991" box="[559,646,1550,1574]" pageId="2" pageNumber="479" value="27-03-18">180327</date>
, one specimen, dissected,
<quantity id="A267E6A2A03EFF80FC36F9BAE18AF992" box="[954,1023,1549,1574]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="479" unit="mm" value="6.0">6 mm</quantity>
in length preserved, Hawaii, Maui, Airport Beach, collected by Cory Pittman, 0
<date id="11216D87A03EFF80FD03F985E131F9FD" box="[655,836,1585,1610]" pageId="2" pageNumber="479" value="2008-10-04">4 October 2008</date>
;
<collectionCode id="038ED382A03EFF80FCDCF985E1D4F9FD" box="[848,929,1586,1610]" pageId="2" pageNumber="479">CASIZ</collectionCode>
0 76055, one specimen, dissected,
<quantity id="A267E6A2A03EFF80FAA2F986E70BF9FE" box="[1326,1406,1585,1610]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="479" unit="mm" value="16.0">16 mm</quantity>
in length preserved,
<collectingCountry id="1D880BD7A03EFF80FEE5F9E2E3B4F9D9" box="[361,449,1621,1646]" name="Mexico" pageId="2" pageNumber="479">Mexico</collectingCountry>
, Baja California, Gulf of California, San Jose
<collectingCountry id="1D880BD7A03EFF80FC7CF9E2E642F9D9" box="[1008,1079,1621,1646]" name="Iceland" pageId="2" pageNumber="479">Island</collectingCountry>
, Punta Colorado, collected by Antonio J. Ferreira
<date id="11216D87A03EFF80FEFEF9CEE073F925" box="[370,518,1657,1682]" pageId="2" pageNumber="479" value="1971-07-15">15 July 1971</date>
.
</paragraph>
<paragraph id="65204B47A03EFF80FF4BF92AE01CF839" blockId="2.[151,1437,1189,2006]" pageId="2" pageNumber="479">
<emphasis id="57EB9755A03EFF80FF4BF92AE07CF901" bold="true" box="[199,521,1693,1718]" pageId="2" pageNumber="479">Geographical distribution.</emphasis>
Originally described from
<collectingCountry id="1D880BD7A03EFF80FCC7F92AE1CDF901" box="[843,952,1693,1718]" name="Mauritius" pageId="2" pageNumber="479">Mauritius</collectingCountry>
(
<bibRefCitation id="010E36B6A03EFF80FC46F92AE622F901" author="Bergh" box="[970,1111,1693,1718]" pageId="2" pageNumber="479" refString="Bergh, L. S. R. (1888) Malacologische Untersuchungen. In: Semper, C. (Ed.), Reisen im Archipel der Philippinen. Zweiter Theil, Wissenschaftliche Resultate. Band 2, Heft 16, pp. 755 - 814." type="journal article" year="1888">Bergh 1888</bibRefCitation>
), it is also known from the
<collectingCountry id="1D880BD7A03EFF80FF1BF976E37AF96D" box="[151,271,1729,1754]" name="Seychelles" pageId="2" pageNumber="479">Seychelles</collectingCountry>
(Gosliner
<emphasis id="57EB9755A03EFF80FE18F975E3A1F96D" box="[404,468,1729,1754]" italics="true" pageId="2" pageNumber="479">et al.</emphasis>
2008),
<collectingCountry id="1D880BD7A03EFF80FDB4F976E0E8F96D" box="[568,669,1729,1754]" name="Tanzania" pageId="2" pageNumber="479">Tanzania</collectingCountry>
(
<bibRefCitation id="010E36B6A03EFF80FD3EF976E141F96D" author="Eliot" box="[690,820,1729,1754]" pageId="2" pageNumber="479" refString="Eliot, C. N. E. (1903) On some nudibranchs from east Africa and Zanzibar, part II. Proceedings of the Zoological Society of London, 1, 250 - 257." type="journal article" year="1903">Eliot 1903</bibRefCitation>
;
<bibRefCitation id="010E36B6A03EFF80FCCAF976E18FF96D" author="Edmunds" box="[838,1018,1729,1754]" pageId="2" pageNumber="479" refString="Edmunds, M. (1970) Opisthobranchiate Mollusca from Tanzania. II. Eolidacea (Eubranchidae and Aeolidiidae). Proceedings of the Malacological Society of London, 39, 15 - 57." type="journal article" year="1970">Edmunds 1970</bibRefCitation>
),
<collectingCountry id="1D880BD7A03EFF80FB99F976E6DCF96D" box="[1045,1193,1729,1754]" name="Mozambique" pageId="2" pageNumber="479">Mozambique</collectingCountry>
(present study), the
<collectingCountry id="1D880BD7A03EFF80FF1BF952E283F949" box="[151,246,1765,1790]" name="Réunion" pageId="2" pageNumber="479">Reunion</collectingCountry>
<collectingCountry id="1D880BD7A03EFF80FE89F952E33CF949" box="[261,329,1765,1790]" name="Iceland" pageId="2" pageNumber="479">Island</collectingCountry>
(
<bibRefCitation id="010E36B6A03EFF80FEEDF952E065F949" author="Bidgrain" box="[353,528,1765,1790]" pageId="2" pageNumber="479" refString="Bidgrain, P. (2013) Spurilla major (Eliot, 1903). [Message in] South-west Indian Ocean Seaslug site. Available from: http: // seaslugs. free. fr / nudibranche / a _ intro. htm. (accessed 1 January 2013)" type="journal article" year="2013">Bidgrain 2013</bibRefCitation>
),
<collectingCountry id="1D880BD7A03EFF80FDA0F952E0E7F949" box="[556,658,1765,1790]" name="Australia" pageId="2" pageNumber="479">Australia</collectingCountry>
(
<bibRefCitation id="010E36B6A03EFF80FD26F952E1B5F949" author="Cobb" box="[682,960,1765,1790]" pageId="2" pageNumber="479" refString="Cobb, G. &amp; Mullins, D. (2013) Baeolidia major [Message in] Nudibranchs Sunshine Coast Australia. Available at: http: // www. nudibranch. com. au / pages / b 2879 WPw. htm (accessed 1 January 2013)" type="journal article" year="2013">Cobb &amp; Mullins 2013</bibRefCitation>
),
<collectingCountry id="1D880BD7A03EFF80FC50F952E63CF949" box="[988,1097,1765,1790]" name="Indonesia" pageId="2" pageNumber="479">Indonesia</collectingCountry>
(Gosliner
<emphasis id="57EB9755A03EFF80FB5EF951E766F949" box="[1234,1299,1765,1790]" italics="true" pageId="2" pageNumber="479">et al.</emphasis>
2008), the
<collectingCountry id="1D880BD7A03EFF80FF1BF8BEE360F895" box="[151,277,1801,1826]" name="Philippines" pageId="2" pageNumber="479">Philippines</collectingCountry>
(Gosliner
<emphasis id="57EB9755A03EFF80FE1BF8BDE3A1F895" box="[407,468,1801,1826]" italics="true" pageId="2" pageNumber="479">et al.</emphasis>
2008),
<collectingCountry id="1D880BD7A03EFF80FDB9F8BDE000F896" box="[565,629,1802,1825]" name="Japan" pageId="2" pageNumber="479">Japan</collectingCountry>
(
<bibRefCitation id="010E36B6A03EFF80FD05F8BEE16CF895" author="Baba" box="[649,793,1801,1826]" pageId="2" pageNumber="479" refString="Baba, K. (1933 b) Preliminary account on the Nudibranchia collected in the vicinity of Amakusa Marine Biological Laboratory. Annotationes Zoologicae Japonenses, 14, 165 - 179." type="journal article" year="1933" yearSuffix="b">Baba 1933b</bibRefCitation>
,
<bibRefCitation id="010E36B6A03EFF80FCA5F8BDE111F895" author="Baba" box="[809,868,1802,1826]" pageId="2" pageNumber="479" refString="Baba, K. (1937) Opisthobranchia of Japan (II). Journal of the Department of Agriculture, Kyushu Imperial University, 5, 289 - 344." type="journal article" year="1937">1937</bibRefCitation>
,
<bibRefCitation id="010E36B6A03EFF80FCF9F8BEE1C5F895" author="Baba" box="[885,944,1801,1826]" pageId="2" pageNumber="479" refString="Baba, K. (1955) Opisthobranchia of Sagami Bay. Suppl. Iwanami Shoten, Tokyo, 59 pp." type="book" year="1955">1955</bibRefCitation>
;
<bibRefCitation id="010E36B6A03EFF80FC4DF8BDE64DF895" author="Ono" box="[961,1080,1802,1826]" pageId="2" pageNumber="479" refString="Ono, A. (1999) Opisthobranchs of Kerama Islands. TBS-Britannica Co., Ltd, Tokyo, 184 pp." type="book" year="1999">Ono 1999</bibRefCitation>
,
<bibRefCitation id="010E36B6A03EFF80FBC4F8BDE6F6F895" author="Ono" box="[1096,1155,1802,1826]" pageId="2" pageNumber="479" refString="Ono, A. (2004) Opisthobranchs of Ryukyu Islands. Rutles, Inc, Tokyo, 304 pp." type="book" year="2004">2004</bibRefCitation>
;
<bibRefCitation id="010E36B6A03EFF80FB18F8BEE747F895" author="Nakano" box="[1172,1330,1801,1826]" pageId="2" pageNumber="479" refString="Nakano, R. (2004) Opisthobranchs of Japan Islands. Rutles, Inc, Tokyo, 304 pp." type="book" year="2004">Nakano 2004</bibRefCitation>
) and the
<collectingCountry id="1D880BD7A03EFF80FF1BF89AE326F8F1" box="[151,339,1837,1862]" name="Marshall Islands" pageId="2" pageNumber="479">Marshall Islands</collectingCountry>
(present study). This species is also found in the Eastern Pacific: Hawaii (
<bibRefCitation id="010E36B6A03EFF80FB34F899E758F8F1" author="Kay" box="[1208,1325,1838,1862]" pageId="2" pageNumber="479" refString="Kay, E. A. (1979) Hawaiian marine shells, reef and shore fauna of Hawaii, section 4: Mollusca. Bishop Museum Press, Honolulu, 653 pp." type="book" year="1979">Kay 1979</bibRefCitation>
;
<bibRefCitation id="010E36B6A03EFF80FAB0F89AE2A6F8DD" author="Gosliner" pageId="2" pageNumber="479" refString="Gosliner, T. M. (1980) The systematics of the Aeolidacea (Nudibranchia: Mollusca) of the Hawaiian Islands, with description of two new species. Pacific Science, 33, 37 - 77." type="journal article" year="1980">Gosliner 1980</bibRefCitation>
), tropical Pacific North
<collectingCountry id="1D880BD7A03EFF80FE6DF8E6E034F8DD" box="[481,577,1873,1898]" name="United States of America" pageId="2" pageNumber="479">America</collectingCountry>
(Gosliner
<emphasis id="57EB9755A03EFF80FD36F8E5E087F8DD" box="[698,754,1873,1898]" italics="true" pageId="2" pageNumber="479">et al.</emphasis>
2008) and
<collectingCountry id="1D880BD7A03EFF80FCFDF8E6E1B2F8DD" box="[881,967,1873,1898]" name="Mexico" pageId="2" pageNumber="479">Mexico</collectingCountry>
(
<bibRefCitation id="010E36B6A03EFF80FC59F8E5E61EF8DD" author="Farmer" box="[981,1131,1873,1898]" pageId="2" pageNumber="479" refString="Farmer, W. (1966) Range extension of Berghia amakusana (Baba) to the East Pacific. The Veliger, 9, 251." type="journal article" year="1966">Farmer 1966</bibRefCitation>
). Recently, it has also been reported in
<collectingCountry id="1D880BD7A03EFF80FE94F8C2E31DF839" box="[280,360,1909,1934]" name="Turkey" pageId="2" pageNumber="479">Turkey</collectingCountry>
(
<bibRefCitation id="010E36B6A03EFF80FEFAF8C2E028F839" author="Turk" box="[374,605,1909,1934]" pageId="2" pageNumber="479" refString="Turk, T. &amp; Furlan, B. (2011) New records of Indo-Pacific and Atlantic mollusc species (Opisthobranchia) in the Eastern Mediterranean and Adriatic Sea. Annales Series Historia Naturalis, 21, 5 - 10." type="journal article" year="2011">
<collectingCountry id="1D880BD7A03EFF80FEFAF8C2E3D8F839" box="[374,429,1909,1934]" name="Turks and Caicos Islands" pageId="2" pageNumber="479">Turk</collectingCountry>
&amp; Furlan 2011
</bibRefCitation>
).
</paragraph>
<paragraph id="65204B47A03EFF81FF4BF82EE093FEF7" blockId="2.[151,1437,1189,2006]" lastBlockId="3.[151,1437,151,320]" lastPageId="3" lastPageNumber="480" pageId="2" pageNumber="479">
<emphasis id="57EB9755A03EFF80FF4BF82EE3B2F805" bold="true" box="[199,455,1945,1970]" pageId="2" pageNumber="479">External morphology</emphasis>
(
<figureCitation id="FDA457C2A03EFF80FE5BF82EE051F805" box="[471,548,1945,1970]" captionStart="FIGURE 2" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[151,1436,373,1863]" captionTargetId="figure@3.[151,1436,373,1865]" captionTargetPageId="3" captionText="FIGURE 2. Photographs of the living animals. A, B, Baeolidia moebii: (A) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177602; (B) specimen from Mozambique, photo by Yara Tibiriça, MNCN 15.05 / 63447; (C) Baeolidia macleayi, specimen from Australia, photo by Terrence M. Gosliner, CASIZ 71923; (D-E) Baeolidia japonica: (D) specimen from Japan, photo by Akira Kawahara, CASIZ 1814520; (E) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 186795; (F) Baeolidia ransoni, specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 186208." httpUri="https://zenodo.org/record/249607/files/figure.png" pageId="2" pageNumber="479">Figs. 2</figureCitation>
A–B, 3A): Body large, broad, tapering gradually towards posterior end of foot. Foot corners tentaculiform. Body colour variable, commonly greyish white or brownish green (
<figureCitation id="FDA457C2A03EFF80FB64F80AE75BF861" box="[1256,1326,1981,2006]" captionStart="FIGURE 2" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[151,1436,373,1863]" captionTargetId="figure@3.[151,1436,373,1865]" captionTargetPageId="3" captionText="FIGURE 2. Photographs of the living animals. A, B, Baeolidia moebii: (A) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177602; (B) specimen from Mozambique, photo by Yara Tibiriça, MNCN 15.05 / 63447; (C) Baeolidia macleayi, specimen from Australia, photo by Terrence M. Gosliner, CASIZ 71923; (D-E) Baeolidia japonica: (D) specimen from Japan, photo by Akira Kawahara, CASIZ 1814520; (E) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 186795; (F) Baeolidia ransoni, specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 186208." httpUri="https://zenodo.org/record/249607/files/figure.png" pageId="2" pageNumber="479">Fig. 2</figureCitation>
A). Some specimens reddish (
<figureCitation id="FDA457C2A03FFF81FEF4FF20E3CAFF07" box="[376,447,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1886,1908]" captionTargetBox="[151,1436,373,1863]" captionTargetId="figure@3.[151,1436,373,1865]" captionTargetPageId="3" captionText="FIGURE 2. Photographs of the living animals. A, B, Baeolidia moebii: (A) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177602; (B) specimen from Mozambique, photo by Yara Tibiriça, MNCN 15.05 / 63447; (C) Baeolidia macleayi, specimen from Australia, photo by Terrence M. Gosliner, CASIZ 71923; (D-E) Baeolidia japonica: (D) specimen from Japan, photo by Akira Kawahara, CASIZ 1814520; (E) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 186795; (F) Baeolidia ransoni, specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 186208." httpUri="https://zenodo.org/record/249607/files/figure.png" pageId="3" pageNumber="480">Fig. 2</figureCitation>
B). Background colour brownish green, overlaid with reticulate ochre pattern. Dorsum usually with bright white patches. Whitish or yellow ring on anteriormost edge of head, continuing towards oral tentacles and forming a distinct light band; both, ring and band, may be interrupted. Rhinophores, oral tentacles and foot corners same as body colour. Rhinophores shorter than oral tentacles. Rhinophores studded with minute knobs (
<figureCitation id="FDA457C2A03FFF81FF13FE90E294FEF7" box="[159,225,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1459,1481]" captionTargetBox="[354,1221,198,1422]" captionTargetId="figure@4.[336,1251,193,1437]" captionTargetPageId="4" captionText="FIGURE 3. Different papillate rhinophores in Baeolidia: (A) from Rudman (1982); (B) from Gosliner (1985). Scale bars: A, 3 mm; B, 1 mm; C, 1.2 mm; D, 1.5 mm." httpUri="https://zenodo.org/record/249608/files/figure.png" pageId="3" pageNumber="480">Fig. 3</figureCitation>
A); apex white. Oral tentacles with white tips.
</paragraph>
<caption id="31E01BCFA03FFF81FF1BF8E9E775F858" httpUri="https://zenodo.org/record/249607/files/figure.png" pageId="3" pageNumber="480" targetBox="[151,1436,373,1863]" targetPageId="3">
<paragraph id="65204B47A03FFF81FF1BF8E9E775F858" blockId="3.[151,1436,1886,2031]" pageId="3" pageNumber="480">
<emphasis id="57EB9755A03FFF81FF1BF8E9E366F8C4" bold="true" box="[151,275,1886,1908]" pageId="3" pageNumber="480">FIGURE 2.</emphasis>
Photographs of the living animals. A, B,
<taxonomicName id="A29F30C4A03FFF81FD37F8E9E11CF8C3" box="[699,873,1886,1908]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="480" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03FFF81FD37F8E9E11CF8C3" box="[699,873,1886,1908]" italics="true" pageId="3" pageNumber="480">Baeolidia moebii</emphasis>
</taxonomicName>
: (A) specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 177602; (B) specimen from Mozambique, photo by Yara Tibiriça, MNCN 15.05/63447; (C)
<emphasis id="57EB9755A03FFF81FAB5F8CAE28CF805" italics="true" pageId="3" pageNumber="480">
<taxonomicName id="A29F30C4A03FFF81FAB5F8CAE280F805" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="480" phylum="Mollusca" rank="species" species="macleayi">Baeolidia macleayi</taxonomicName>
,
</emphasis>
specimen from Australia, photo by Terrence M. Gosliner, CASIZ 71923; (D-E)
<taxonomicName id="A29F30C4A03FFF81FBB2F82BE777F805" box="[1086,1282,1948,1970]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="480" phylum="Mollusca" rank="species" species="japonica">
<emphasis id="57EB9755A03FFF81FBB2F82BE777F805" box="[1086,1282,1948,1970]" italics="true" pageId="3" pageNumber="480">Baeolidia japonica</emphasis>
</taxonomicName>
: (D) specimen from Japan, photo by Akira Kawahara, CASIZ 1814520; (E) specimen from Marshall Islands, photo by Scott Johnson, CASIZ 186795; (F)
<emphasis id="57EB9755A03FFF81FE99F86EE3A7F858" box="[277,466,2009,2031]" italics="true" pageId="3" pageNumber="480">
<taxonomicName id="A29F30C4A03FFF81FE99F86EE3BBF858" box="[277,462,2009,2031]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="3" pageNumber="480" phylum="Mollusca" rank="species" species="ransoni">Baeolidia ransoni</taxonomicName>
,
</emphasis>
specimen from the Philippines, photo by Terrence M. Gosliner, CASIZ 186208.
</paragraph>
</caption>
<caption id="31E01BCFA038FF86FF1BFA04E05CFA50" httpUri="https://zenodo.org/record/249608/files/figure.png" pageId="4" pageNumber="481" targetBox="[354,1221,198,1422]" targetPageId="4">
<paragraph id="65204B47A038FF86FF1BFA04E05CFA50" blockId="4.[151,1436,1459,1511]" pageId="4" pageNumber="481">
<emphasis id="57EB9755A038FF86FF1BFA04E366FA7F" bold="true" box="[151,275,1459,1481]" pageId="4" pageNumber="481">FIGURE 3.</emphasis>
Different papillate rhinophores in
<taxonomicName id="A29F30C4A038FF86FDF4FA04E0AEFA7E" box="[632,731,1459,1481]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="4" pageNumber="481" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A038FF86FDF4FA04E0AEFA7E" box="[632,731,1459,1481]" italics="true" pageId="4" pageNumber="481">Baeolidia</emphasis>
</taxonomicName>
: (A) from Rudman (1982); (B) from Gosliner (1985). Scale bars: A, 3mm; B, 1 mm; C, 1.2 mm; D, 1.5 mm.
</paragraph>
</caption>
<paragraph id="65204B47A038FF86FF4BF9A3E01AF956" blockId="4.[151,1437,1556,1977]" pageId="4" pageNumber="481">Cerata length variable. Cerata flattened, almost leaf-like. Branches of digestive gland brownish-green, shining through translucent body wall. Whitish patches on outer side of cerata, probably with bright yellow marks. Cerata usually with bright yellow subapical band and just below it a purplish area. Apex translucent white. Cerata composed of up to nine arches, leaving a distinct gap between pre and post-pericardial groups. Each arch with 4–15 cerata, decreasing in size towards foot. Anus cleioproctic, within second right arch. Genital opening placed among cerata of anteriormost group on right side.</paragraph>
<paragraph id="65204B47A038FF86FF4BF95BE0C1F80E" blockId="4.[151,1437,1556,1977]" pageId="4" pageNumber="481">
<emphasis id="57EB9755A038FF86FF4BF95BE34EF8B2" bold="true" box="[199,315,1772,1797]" pageId="4" pageNumber="481">Anatomy.</emphasis>
Masticatory border of jaws smooth (
<figureCitation id="FDA457C2A038FF86FD5AF95BE16DF8B3" box="[726,792,1772,1797]" captionStart="FIGURE 4" captionStartId="5.[151,250,1706,1728]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@5.[151,1436,193,1685]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron photographs. (A – B) Baeolidia moebii (CASIZ 180327): (A) Detailed view of the masticatory border; (B) Radular teeth; (C – D) Baeolidia macleayi (CASIZ 195247): (C) Detailed view of the masticatory border; (D) Radular teeth; (E – F) Baeolidia japonica (CASIZ 181357); (E) Detailed view of the masticatory border; (F) Radular teeth. Scale bars: A, 10 µm; B, 30 µm; C-D, 10 µm; E, 100 µm; F, 150 µm." httpUri="https://zenodo.org/record/249609/files/figure.png" pageId="4" pageNumber="481">Fig. 4</figureCitation>
A). Radular formulae
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0.1.0 (
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15.05/
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),
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),
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0.1.0 (
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) and
<date id="11216D87A038FF86FBDCF8A7E6F7F89F" box="[1104,1154,1808,1832]" pageId="4" pageNumber="481" value="1922-10">22 x</date>
0.1.0 (
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, 6 mm
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). Radular teeth bent upwards with up to 100 elongate, fine and acutely pointed denticles from side to side, without central notch or central cusp (
<figureCitation id="FDA457C2A038FF86FDB1F8EFE0F5F8C7" box="[573,640,1880,1905]" captionStart="FIGURE 4" captionStartId="5.[151,250,1706,1728]" captionTargetBox="[151,1436,193,1683]" captionTargetId="figure@5.[151,1436,193,1685]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron photographs. (A – B) Baeolidia moebii (CASIZ 180327): (A) Detailed view of the masticatory border; (B) Radular teeth; (C – D) Baeolidia macleayi (CASIZ 195247): (C) Detailed view of the masticatory border; (D) Radular teeth; (E – F) Baeolidia japonica (CASIZ 181357); (E) Detailed view of the masticatory border; (F) Radular teeth. Scale bars: A, 10 µm; B, 30 µm; C-D, 10 µm; E, 100 µm; F, 150 µm." httpUri="https://zenodo.org/record/249609/files/figure.png" pageId="4" pageNumber="481">Fig. 4</figureCitation>
B). Teeth progressively smaller towards posterior region of radula. Oral glands fragile, small and spongy, lying dorso-laterally to buccal bulb (
<figureCitation id="FDA457C2A038FF86FC49F8CBE679F822" box="[965,1036,1916,1941]" captionStart="FIGURE 5" captionStartId="6.[151,250,1251,1273]" captionText="FIGURE 5. Oral glands of Baeolidia. Scale bar: 0.5 mm." pageId="4" pageNumber="481">Fig. 5</figureCitation>
). Oral gland morphology variable, from rounded to elongate. Salivary glands large.
</paragraph>
<caption id="31E01BCFA039FF87FF1BF91DE162F8AB" httpUri="https://zenodo.org/record/249609/files/figure.png" pageId="5" pageNumber="482" targetBox="[151,1436,193,1683]" targetPageId="5">
<paragraph id="65204B47A039FF87FF1BF91DE162F8AB" blockId="5.[151,1436,1706,1820]" pageId="5" pageNumber="482">
<emphasis id="57EB9755A039FF87FF1BF91DE361F908" bold="true" box="[151,276,1706,1728]" pageId="5" pageNumber="482">FIGURE 4.</emphasis>
Scanning electron photographs. (A–B)
<taxonomicName id="A29F30C4A039FF87FD22F91DE128F977" box="[686,861,1706,1728]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="482" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A039FF87FD22F91DE128F977" box="[686,861,1706,1728]" italics="true" pageId="5" pageNumber="482">Baeolidia moebii</emphasis>
</taxonomicName>
(CASIZ 180327): (A) Detailed view of the masticatory border; (B) Radular teeth; (C–D)
<taxonomicName id="A29F30C4A039FF87FE73F97EE0B2F968" box="[511,711,1737,1759]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="482" phylum="Mollusca" rank="species" species="macleayi">
<emphasis id="57EB9755A039FF87FE73F97EE0B2F968" box="[511,711,1737,1759]" italics="true" pageId="5" pageNumber="482">Baeolidia macleayi</emphasis>
</taxonomicName>
(CASIZ 195247): (C) Detailed view of the masticatory border; (D) Radular teeth; (E–F)
<taxonomicName id="A29F30C4A039FF87FEE0F950E058F949" box="[364,557,1767,1790]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="5" pageNumber="482" phylum="Mollusca" rank="species" species="japonica">
<emphasis id="57EB9755A039FF87FEE0F950E058F949" box="[364,557,1767,1790]" italics="true" pageId="5" pageNumber="482">Baeolidia japonica</emphasis>
</taxonomicName>
(CASIZ 181357); (E) Detailed view of the masticatory border; (F) Radular teeth. Scale bars: A, 10 µm; B, 30 µm; C-D, 10 µm; E, 100 µm; F, 150 µm.
</paragraph>
</caption>
<paragraph id="65204B47A039FF87FF4BF8F4E1F1F87F" blockId="5.[151,1436,1859,1992]" pageId="5" pageNumber="482">
Reproductive system diaulic (
<figureCitation id="FDA457C2A039FF87FD98F8F4E022F8EB" box="[532,599,1859,1884]" captionStart="FIGURE 6" captionStartId="7.[151,250,1532,1554]" captionTargetBox="[337,1251,227,1481]" captionTargetId="figure@7.[321,1265,199,1494]" captionTargetPageId="7" captionText="FIGURE 6. Reproductive system. (A) Baeolidia moebii, specimen from Mozambique (MNCN 15.05 / 63447); (B) Baeolidia macleayi, specimen from Australia (CASIZ 195247); (C) Baeolidia japonica, specimen from Marshall Islands (CASIZ 181357); (D) Baeolidia ransoni, specimen from the Philippines (CASIZ 186209). Scale bars: A – D, 0.5 mm. Abbreviations: am, ampulla; fgm, female gland mass; ps, penial sac; rs, receptaculum seminis; va, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/249610/files/figure.png" pageId="5" pageNumber="482">Fig. 6</figureCitation>
A). Preampullary duct widening into elongate ampulla. Postampullary duct dividing into oviduct and vas deferens. Vas deferens elongate, moderately wide, penetrating into wider proximal portion of penial sac with unarmed penial papilla. Receptaculum seminis ovoid, short stalk connecting to short oviduct, before latter forms female glands. Vagina ventral to penis.
</paragraph>
</subSubSection>
<subSubSection id="2D8518CCA03AFF84FF4BFF20E19EF886" pageId="6" pageNumber="483" type="discussion">
<paragraph id="65204B47A03AFF84FF4BFF20E13AFDAF" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="483">
<emphasis id="57EB9755A03AFF84FF4BFF20E34EFF07" bold="true" box="[199,315,151,176]" pageId="6" pageNumber="483">Remarks.</emphasis>
This species has not been studied under the name of
<taxonomicName id="A29F30C4A03AFF84FC19FF20E623FF07" box="[917,1110,151,176]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FC19FF20E623FF07" box="[917,1110,151,176]" italics="true" pageId="6" pageNumber="483">Baeolidia moebii</emphasis>
</taxonomicName>
since its original description (
<bibRefCitation id="010E36B6A03AFF84FF12FF0BE35CFF63" author="Bergh" box="[158,297,188,213]" pageId="6" pageNumber="483" refString="Bergh, L. S. R. (1888) Malacologische Untersuchungen. In: Semper, C. (Ed.), Reisen im Archipel der Philippinen. Zweiter Theil, Wissenschaftliche Resultate. Band 2, Heft 16, pp. 755 - 814." type="journal article" year="1888">Bergh 1888</bibRefCitation>
). When
<bibRefCitation id="010E36B6A03AFF84FE05FF0BE066FF63" author="Eliot" box="[393,531,188,213]" pageId="6" pageNumber="483" refString="Eliot, C. N. E. (1903) On some nudibranchs from east Africa and Zanzibar, part II. Proceedings of the Zoological Society of London, 1, 250 - 257." type="journal article" year="1903">Eliot (1903)</bibRefCitation>
described
<taxonomicName id="A29F30C4A03AFF84FD1CFF0AE087FF63" box="[656,754,189,212]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="major">
<emphasis id="57EB9755A03AFF84FD1CFF0AE087FF63" box="[656,754,189,212]" italics="true" pageId="6" pageNumber="483">B. major</emphasis>
</taxonomicName>
, he pointed out that the latter could be merely a full-grown individual of
<taxonomicName id="A29F30C4A03AFF84FEBFFF56E3D4FF4F" box="[307,417,223,248]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FEBFFF56E3D4FF4F" box="[307,417,223,248]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
since the main difference between both species was the size. Nevertheless,
<bibRefCitation id="010E36B6A03AFF84FA9EFF68E7E9FF4F" author="Eliot" box="[1298,1436,223,248]" pageId="6" pageNumber="483" refString="Eliot, C. N. E. (1903) On some nudibranchs from east Africa and Zanzibar, part II. Proceedings of the Zoological Society of London, 1, 250 - 257." type="journal article" year="1903">Eliot (1903)</bibRefCitation>
finally described
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<emphasis id="57EB9755A03AFF84FED3FEB2E3B1FEAB" box="[351,452,261,284]" italics="true" pageId="6" pageNumber="483">B. major</emphasis>
</taxonomicName>
as a valid species, owing to the ambiguities in the original description of
<taxonomicName id="A29F30C4A03AFF84FAAAFEB2E7E1FEAA" box="[1318,1428,260,285]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FAAAFEB2E7E1FEAA" box="[1318,1428,260,285]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
. Taking into consideration Eliot’s statements together with the proximity of both
<typeStatus id="BA24F5E5A03AFF84FBBFFE9FE616FEF7" box="[1075,1123,296,320]" pageId="6" pageNumber="483">type</typeStatus>
localities (
<collectingCountry id="1D880BD7A03AFF84FB6EFE90E724FEF7" box="[1250,1361,295,320]" name="Mauritius" pageId="6" pageNumber="483">Mauritius</collectingCountry>
for
<taxonomicName id="A29F30C4A03AFF84FA08FE9EE296FED2" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FA08FE9EE296FED2" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
and
<collectingCountry id="1D880BD7A03AFF84FE95FEFBE308FED2" box="[281,381,332,357]" name="Tanzania" pageId="6" pageNumber="483">Zanzibar</collectingCountry>
for
<taxonomicName id="A29F30C4A03AFF84FE20FEFAE07BFED3" box="[428,526,333,356]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="major">
<emphasis id="57EB9755A03AFF84FE20FEFAE07BFED3" box="[428,526,333,356]" italics="true" pageId="6" pageNumber="483">B. major</emphasis>
</taxonomicName>
),
<bibRefCitation id="010E36B6A03AFF84FDA9FEFBE169FED3" box="[549,796,332,357]" pageId="6" pageNumber="483" refString="Carmona, L., Pola, M., Gosliner, T. M. &amp; Cervera, L. (2013) A tale that morphology fails to tell: a molecular phylogeny of Aeolidiidae (Aeolidida, Nudibranchia, Gastropoda). PloS One, 8, 5. http: // dx. doi. org / 10.1371 / journal. pone. 0063000" type="journal article">
Carmona
<emphasis id="57EB9755A03AFF84FD1FFEFAE0BEFED2" box="[659,715,332,357]" italics="true" pageId="6" pageNumber="483">et al.</emphasis>
(2013)
</bibRefCitation>
considered
<taxonomicName id="A29F30C4A03AFF84FC2FFEFAE670FED3" box="[931,1029,333,356]" class="Gastropoda" family="Aeolidiidae" genus="Spurilla" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="major">
<emphasis id="57EB9755A03AFF84FC2FFEFAE670FED3" box="[931,1029,333,356]" italics="true" pageId="6" pageNumber="483">B. major</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="A29F30C4A03AFF84FA98FEFAE70AFED2" box="[1300,1407,332,357]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FA98FEFAE70AFED2" box="[1300,1407,332,357]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
. Neither
<bibRefCitation id="010E36B6A03AFF84FF78FED8E3FFFE3F" author="Bergh" box="[244,394,367,392]" pageId="6" pageNumber="483" refString="Bergh, L. S. R. (1888) Malacologische Untersuchungen. In: Semper, C. (Ed.), Reisen im Archipel der Philippinen. Zweiter Theil, Wissenschaftliche Resultate. Band 2, Heft 16, pp. 755 - 814." type="journal article" year="1888">Bergh (1888)</bibRefCitation>
nor
<bibRefCitation id="010E36B6A03AFF84FE4DFED8E03EFE3F" author="Eliot" box="[449,587,367,392]" pageId="6" pageNumber="483" refString="Eliot, C. N. E. (1903) On some nudibranchs from east Africa and Zanzibar, part II. Proceedings of the Zoological Society of London, 1, 250 - 257." type="journal article" year="1903">Eliot (1903)</bibRefCitation>
mentioned the presence of oral glands in
<taxonomicName id="A29F30C4A03AFF84FBA0FEC6E6ECFE3F" box="[1068,1177,367,392]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FBA0FEC6E6ECFE3F" box="[1068,1177,367,392]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
. Only
<bibRefCitation id="010E36B6A03AFF84FB66FED8E7E9FE3F" author="Rudman" box="[1258,1436,367,392]" pageId="6" pageNumber="483" refString="Rudman, W. B. (1982) The taxonomy and biology of further aeolidacean and arminacean nudibranch molluscs with symbiotic zooxanthellae. Zoological Journal of the Linnean Society, 74, 147 - 196. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1982. tb 01146. x" type="journal article" year="1982">Rudman (1982)</bibRefCitation>
pointed out the existence of salivary glands in this species. Two of our specimens possessed oral glands, and salivary glands were found twice. This variability could be an artefact of the state of preservation of these specimens. An alternative hypothesis could be an intraspecific variation for these two characters, which has also been observed in
<taxonomicName id="A29F30C4A03AFF84FED0FE48E049FDAF" box="[348,572,511,536]" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="nodosa">
<emphasis id="57EB9755A03AFF84FED0FE48E049FDAF" box="[348,572,511,536]" italics="true" pageId="6" pageNumber="483">Limenandra nodosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="010E36B6A03AFF84FDC7FDB7E137FDAF" author="Carmona" box="[587,834,511,536]" pageId="6" pageNumber="483" refString="Carmona, L., Pola, M., Gosliner, T. M. &amp; Cervera, L. (2014 c) The end of a long controversy: systematics of the genus Limenandra (Mollusca: Nudibranchia: Aeolidiidae). Helgoland Marine Research, 68, 37 - 48. http: // dx. doi. org / 10.1007 / s 10152 - 013 - 0367 - y" type="journal article" year="2014" yearSuffix="c">
Carmona
<emphasis id="57EB9755A03AFF84FD36FDB6E09EFDAF" box="[698,747,511,536]" italics="true" pageId="6" pageNumber="483">et al</emphasis>
. 2014c
</bibRefCitation>
).
</paragraph>
<caption id="31E01BCFA03AFF84FF1BFB54E0ABFB4E" box="[151,734,1251,1273]" pageId="6" pageNumber="483">
<paragraph id="65204B47A03AFF84FF1BFB54E0ABFB4E" blockId="6.[151,734,1251,1273]" box="[151,734,1251,1273]" pageId="6" pageNumber="483">
<emphasis id="57EB9755A03AFF84FF1BFB54E367FB4E" bold="true" box="[151,274,1251,1273]" pageId="6" pageNumber="483">FIGURE 5.</emphasis>
Oral glands of
<taxonomicName id="A29F30C4A03AFF84FE3CFB54E067FB4E" box="[432,530,1251,1273]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="genus">
<emphasis id="57EB9755A03AFF84FE3CFB54E067FB4E" box="[432,530,1251,1273]" italics="true" pageId="6" pageNumber="483">Baeolidia</emphasis>
</taxonomicName>
. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="65204B47A03AFF84FF4BFA97E373F9A6" blockId="6.[151,1436,1312,1841]" pageId="6" pageNumber="483">
Despite the large variability in colouration, our specimens identified as
<taxonomicName id="A29F30C4A03AFF84FB96FA97E6AAFA8E" box="[1050,1247,1312,1337]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FB96FA97E6AAFA8E" box="[1050,1247,1312,1337]" italics="true" pageId="6" pageNumber="483">Baeolidia moebii</emphasis>
</taxonomicName>
match with the information given by
<bibRefCitation id="010E36B6A03AFF84FE18FAF3E058FAEA" author="Bergh" box="[404,557,1348,1373]" pageId="6" pageNumber="483" refString="Bergh, L. S. R. (1888) Malacologische Untersuchungen. In: Semper, C. (Ed.), Reisen im Archipel der Philippinen. Zweiter Theil, Wissenschaftliche Resultate. Band 2, Heft 16, pp. 755 - 814." type="journal article" year="1888">Bergh (1888)</bibRefCitation>
and
<bibRefCitation id="010E36B6A03AFF84FDE6FAF3E083FAEA" author="Eliot" box="[618,758,1348,1373]" pageId="6" pageNumber="483" refString="Eliot, C. N. E. (1903) On some nudibranchs from east Africa and Zanzibar, part II. Proceedings of the Zoological Society of London, 1, 250 - 257." type="journal article" year="1903">Eliot (1903)</bibRefCitation>
as well as with more recent descriptions (
<bibRefCitation id="010E36B6A03AFF84FB6AFAF3E7E2FAEA" author="Edmunds" box="[1254,1431,1348,1373]" pageId="6" pageNumber="483" refString="Edmunds, M. (1969) Opisthobranchiate Mollusca from Tanzania. I. Eolidacea (Eubranchidae and Aeolidiidae). Proceedings of the Malacological Society of London, 38, 451 - 469." type="journal article" year="1969">Edmunds 1969</bibRefCitation>
;
<bibRefCitation id="010E36B6A03AFF84FF1BFADFE34FFA36" author="Gosliner" box="[151,314,1384,1409]" pageId="6" pageNumber="483" refString="Gosliner, T. M. (1980) The systematics of the Aeolidacea (Nudibranchia: Mollusca) of the Hawaiian Islands, with description of two new species. Pacific Science, 33, 37 - 77." type="journal article" year="1980">Gosliner 1980</bibRefCitation>
;
<bibRefCitation id="010E36B6A03AFF84FEC9FADFE39DFA36" author="Rudman" box="[325,488,1384,1409]" pageId="6" pageNumber="483" refString="Rudman, W. B. (1982) The taxonomy and biology of further aeolidacean and arminacean nudibranch molluscs with symbiotic zooxanthellae. Zoological Journal of the Linnean Society, 74, 147 - 196. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1982. tb 01146. x" type="journal article" year="1982">Rudman 1982</bibRefCitation>
).
<bibRefCitation id="010E36B6A03AFF84FE70FADFE0DBFA36" author="Gosliner" box="[508,686,1384,1409]" pageId="6" pageNumber="483" refString="Gosliner, T. M. (1985) The aeolid nudibranchs family Aeolidiidae (Gastropoda, Opisthobranchia) from tropical southern Annals of the South African Museum, 95, 233 - 267." type="journal article" year="1985">Gosliner (1985)</bibRefCitation>
stated the presence of a penial gland in the reproductive system of
<taxonomicName id="A29F30C4A03AFF84FF1BFA39E376FA12" box="[151,259,1420,1445]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FF1BFA39E376FA12" box="[151,259,1420,1445]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
(identified as
<taxonomicName id="A29F30C4A03AFF84FE28FA3BE032FA12" box="[420,583,1420,1445]" class="Gastropoda" family="Aeolidiidae" genus="Berghia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="major">
<emphasis id="57EB9755A03AFF84FE28FA3BE032FA12" box="[420,583,1420,1445]" italics="true" pageId="6" pageNumber="483">Berghia major</emphasis>
</taxonomicName>
). The only aeolidiid where this condition has been described is
<taxonomicName id="A29F30C4A03AFF84FA9FFA3BE3C7FA7E" authority="Schmekel 1970" authorityName="Schmekel" authorityYear="1970" class="Gastropoda" family="Aeolidiidae" genus="Limenandra" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="nodosa">
<emphasis id="57EB9755A03AFF84FA9FFA3BE29DFA7E" italics="true" pageId="6" pageNumber="483">Limenandra nodosa</emphasis>
(
<bibRefCitation id="010E36B6A03AFF84FF7AFA07E3DFFA7E" author="Schmekel" box="[246,426,1456,1481]" pageId="6" pageNumber="483" refString="Schmekel, L. (1970) Anatomie der Genitalorgane von Nudibranchiern (Gastropoda Euthyneura). Pubblicazioni della Stazione Zoologica di Napoli, 38, 120 - 217." type="journal article" year="1970">Schmekel 1970</bibRefCitation>
)
</taxonomicName>
, although
<bibRefCitation id="010E36B6A03AFF84FDABFA06E159FA7E" box="[551,812,1456,1481]" pageId="6" pageNumber="483" refString="Carmona, L., Pola, M., Gosliner, T. M. &amp; Cervera, L. (2014 c) The end of a long controversy: systematics of the genus Limenandra (Mollusca: Nudibranchia: Aeolidiidae). Helgoland Marine Research, 68, 37 - 48. http: // dx. doi. org / 10.1007 / s 10152 - 013 - 0367 - y" type="journal article">
Carmona
<emphasis id="57EB9755A03AFF84FD19FA05E0BBFA7E" box="[661,718,1456,1481]" italics="true" pageId="6" pageNumber="483">et al.</emphasis>
(2014c)
</bibRefCitation>
did not find any traces of a penial gland. Since none of the specimens here examined exhibited this structure, the existence of a penial gland for
<taxonomicName id="A29F30C4A03AFF84FBFFFA61E6ABFA5A" box="[1139,1246,1492,1517]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FBFFFA61E6ABFA5A" box="[1139,1246,1492,1517]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
is not confirmed (
<figureCitation id="FDA457C2A03AFF84FF13FA4FE296F9A6" box="[159,227,1528,1553]" captionStart="FIGURE 6" captionStartId="7.[151,250,1532,1554]" captionTargetBox="[337,1251,227,1481]" captionTargetId="figure@7.[321,1265,199,1494]" captionTargetPageId="7" captionText="FIGURE 6. Reproductive system. (A) Baeolidia moebii, specimen from Mozambique (MNCN 15.05 / 63447); (B) Baeolidia macleayi, specimen from Australia (CASIZ 195247); (C) Baeolidia japonica, specimen from Marshall Islands (CASIZ 181357); (D) Baeolidia ransoni, specimen from the Philippines (CASIZ 186209). Scale bars: A – D, 0.5 mm. Abbreviations: am, ampulla; fgm, female gland mass; ps, penial sac; rs, receptaculum seminis; va, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/249610/files/figure.png" pageId="6" pageNumber="483">Fig. 6</figureCitation>
A).
</paragraph>
<paragraph id="65204B47A03AFF84FF4BF9ABE7F7F8B9" blockId="6.[151,1436,1312,1841]" pageId="6" pageNumber="483">
Finally, excluding
<taxonomicName id="A29F30C4A03AFF84FE3EF9ABE0E1F982" box="[434,660,1564,1589]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="australis">
<emphasis id="57EB9755A03AFF84FE3EF9ABE0E1F982" box="[434,660,1564,1589]" italics="true" pageId="6" pageNumber="483">Baeolidia australis</emphasis>
</taxonomicName>
(see
<taxonomicName id="A29F30C4A03AFF84FD65F9A9E102F982" box="[745,887,1564,1589]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="australis">
<emphasis id="57EB9755A03AFF84FD65F9A9E102F982" box="[745,887,1564,1589]" italics="true" pageId="6" pageNumber="483">B. australis</emphasis>
</taxonomicName>
remarks),
<taxonomicName id="A29F30C4A03AFF84FB86F9A9E6F4F982" box="[1034,1153,1564,1589]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FB86F9A9E6F4F982" box="[1034,1153,1564,1589]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
colouration is clearly distinguishable from the remaining species of this genus. In terms of internal anatomy, the radular morphology of
<taxonomicName id="A29F30C4A03AFF84FF1BF9D2E322F9C9" box="[151,343,1637,1662]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FF1BF9D2E322F9C9" box="[151,343,1637,1662]" italics="true" pageId="6" pageNumber="483">Baeolidia moebii</emphasis>
</taxonomicName>
is quite particular since teeth are bent and do not have any central notch or denticle. So far, only
<taxonomicName id="A29F30C4A03AFF84FF1BF93DE362F916" box="[151,279,1672,1697]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="australis">
<emphasis id="57EB9755A03AFF84FF1BF93DE362F916" box="[151,279,1672,1697]" italics="true" pageId="6" pageNumber="483">B. australis</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A03AFF84FEC0F93DE3CFF916" box="[332,442,1672,1697]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="lunaris" status="sp. nov.">
<emphasis id="57EB9755A03AFF84FEC0F93DE3CFF916" box="[332,442,1672,1697]" italics="true" pageId="6" pageNumber="483">B. lunaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CD82A2EA03AFF84FE4CF93FE06CF916" box="[448,537,1672,1697]" pageId="6" pageNumber="483" rank="species">
<emphasis id="57EB9755A03AFF84FE4CF93FE06CF916" bold="true" box="[448,537,1672,1697]" pageId="6" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
have similar radular teeth.
<taxonomicName id="A29F30C4A03AFF84FCC4F93FE67DF916" box="[840,1032,1672,1697]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FCC4F93FE67DF916" box="[840,1032,1672,1697]" italics="true" pageId="6" pageNumber="483">Baeolidia moebii</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A03AFF84FBB1F93DE6DEF916" box="[1085,1195,1672,1697]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="lunaris" status="sp. nov.">
<emphasis id="57EB9755A03AFF84FBB1F93DE6DEF916" box="[1085,1195,1672,1697]" italics="true" pageId="6" pageNumber="483">B. lunaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CD82A2EA03AFF84FB3DF93FE77FF916" box="[1201,1290,1672,1697]" pageId="6" pageNumber="483" rank="species">
<emphasis id="57EB9755A03AFF84FB3DF93FE77FF916" bold="true" box="[1201,1290,1672,1697]" pageId="6" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
differ in size (
<taxonomicName id="A29F30C4A03AFF84FF2CF919E37AF971" box="[160,271,1709,1734]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FF2CF919E37AF971" box="[160,271,1709,1734]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
is larger than
<taxonomicName id="A29F30C4A03AFF84FE37F919E059F971" box="[443,556,1709,1734]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="lunaris" status="sp. nov.">
<emphasis id="57EB9755A03AFF84FE37F919E059F971" box="[443,556,1709,1734]" italics="true" pageId="6" pageNumber="483">B. lunaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CD82A2EA03AFF84FDBBF91AE0E1F971" box="[567,660,1709,1734]" pageId="6" pageNumber="483" rank="species">
<emphasis id="57EB9755A03AFF84FDBBF91AE0E1F971" bold="true" box="[567,660,1709,1734]" pageId="6" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
), and rhinophorial ornamentation (
<taxonomicName id="A29F30C4A03AFF84FBA7F919E6E9F971" box="[1067,1180,1709,1734]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="lunaris" status="sp. nov.">
<emphasis id="57EB9755A03AFF84FBA7F919E6E9F971" box="[1067,1180,1709,1734]" italics="true" pageId="6" pageNumber="483">B. lunaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CD82A2EA03AFF84FB2AF91AE776F971" box="[1190,1283,1709,1734]" pageId="6" pageNumber="483" rank="species">
<emphasis id="57EB9755A03AFF84FB2AF91AE776F971" bold="true" box="[1190,1283,1709,1734]" pageId="6" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
has elongate papillae instead of minute knobs and colouration). Additionally, the ampulla of
<taxonomicName id="A29F30C4A03AFF84FB83F965E60FF95E" box="[1039,1146,1744,1769]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FB83F965E60FF95E" box="[1039,1146,1744,1769]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
is larger than the ampulla found in
<taxonomicName id="A29F30C4A03AFF84FF70F941E31FF8B9" box="[252,362,1781,1806]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="lunaris" status="sp. nov.">
<emphasis id="57EB9755A03AFF84FF70F941E31FF8B9" box="[252,362,1781,1806]" italics="true" pageId="6" pageNumber="483">B. lunaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CD82A2EA03AFF84FEFDF942E3BEF8B9" box="[369,459,1781,1806]" pageId="6" pageNumber="483" rank="species">
<emphasis id="57EB9755A03AFF84FEFDF942E3BEF8B9" bold="true" box="[369,459,1781,1806]" pageId="6" pageNumber="483">sp. nov.</emphasis>
</taxonomicNameLabel>
Differences between
<taxonomicName id="A29F30C4A03AFF84FD33F941E15FF8B9" box="[703,810,1781,1806]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="moebii">
<emphasis id="57EB9755A03AFF84FD33F941E15FF8B9" box="[703,810,1781,1806]" italics="true" pageId="6" pageNumber="483">B. moebii</emphasis>
</taxonomicName>
and
<taxonomicName id="A29F30C4A03AFF84FCEDF941E197F8B9" box="[865,994,1781,1806]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="australis">
<emphasis id="57EB9755A03AFF84FCEDF941E197F8B9" box="[865,994,1781,1806]" italics="true" pageId="6" pageNumber="483">B. australis</emphasis>
</taxonomicName>
are recorded in
<taxonomicName id="A29F30C4A03AFF84FB14F941E76CF8B9" box="[1176,1305,1781,1806]" class="Gastropoda" family="Aeolidiidae" genus="Baeolidia" kingdom="Animalia" order="Nudibranchia" pageId="6" pageNumber="483" phylum="Mollusca" rank="species" species="australis">
<emphasis id="57EB9755A03AFF84FB14F941E76CF8B9" box="[1176,1305,1781,1806]" italics="true" pageId="6" pageNumber="483">B. australis</emphasis>
</taxonomicName>
remarks.
</paragraph>
<paragraph id="65204B47A03AFF84FF4BF8AFE19EF886" blockId="6.[151,1436,1312,1841]" box="[199,1003,1816,1841]" pageId="6" pageNumber="483">
This species was studied molecularly by
<bibRefCitation id="010E36B6A03AFF84FD1EF8AEE1FCF886" box="[658,905,1816,1841]" pageId="6" pageNumber="483" refString="Carmona, L., Pola, M., Gosliner, T. M. &amp; Cervera, L. (2013) A tale that morphology fails to tell: a molecular phylogeny of Aeolidiidae (Aeolidida, Nudibranchia, Gastropoda). PloS One, 8, 5. http: // dx. doi. org / 10.1371 / journal. pone. 0063000" type="journal article">
Carmona
<emphasis id="57EB9755A03AFF84FD73F8ADE14DF886" box="[767,824,1816,1841]" italics="true" pageId="6" pageNumber="483">et al.</emphasis>
(2013)
</bibRefCitation>
(
<figureCitation id="FDA457C2A03AFF84FC14F8AFE1ACF886" box="[920,985,1816,1841]" captionStart="FIGURE 1" captionStartId="1.[151,250,984,1006]" captionTargetBox="[183,1403,519,961]" captionTargetId="figure@1.[183,1403,519,962]" captionTargetPageId="1" captionText="FIGURE 1. Phylogeny of Baeolidia, based on Carmona et al. (2013). Names on right side of vertical bars refer to revised classification." httpUri="https://zenodo.org/record/249606/files/figure.png" pageId="6" pageNumber="483">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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