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<mods:title id="EE5D42DB5FF8F934C58EA56C7DDFE6BD">Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam</mods:title>
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<mods:namePart id="945E920BEF67CB2B883EBAFF01C3C242">Tran, A. D.</mods:namePart>
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<mods:namePart id="F5B86C083D81B7B716433DE9997CF745">Nguyen, X. Q.</mods:namePart>
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<treatment id="4F67DC7C68699F6DFF00F8A318F6FD2F" ID-DOI="http://doi.org/10.5281/zenodo.4509714" ID-GBIF-Taxon="120827705" ID-Zenodo-Dep="4509714" LSID="urn:lsid:plazi:treatment:4F67DC7C68699F6DFF00F8A318F6FD2F" httpUri="http://treatment.plazi.org/id/4F67DC7C68699F6DFF00F8A318F6FD2F" lastPageId="12" lastPageNumber="513" pageId="11" pageNumber="512">
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<paragraph id="C7716D6A68699F6AFF00F8A318B9F8B5" blockId="11.[151,674,1796,1822]" box="[151,674,1796,1822]" pageId="11" pageNumber="512">
<heading id="9C39DA0668699F6AFF00F8A318B9F8B5" bold="true" box="[151,674,1796,1822]" fontSize="11" level="1" pageId="11" pageNumber="512" reason="1">
<taxonomicName id="00CE16E968699F6AFF00F8A318B9F8B5" ID-CoL="4SNHH" authority="Tran &amp; Yang, 2006" authorityName="Tran &amp; Yang" authorityYear="2006" box="[151,674,1796,1822]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="anderseni">
<emphasis id="F5BAB17868699F6AFF00F8A318B9F8B5" bold="true" box="[151,674,1796,1822]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF00F8A31BB5F8B5" bold="true" box="[151,430,1796,1822]" italics="true" pageId="11" pageNumber="512">Rhyacobates anderseni</emphasis>
<bibRefCitation id="A35F109B68699F6AFE22F8A318B9F8B5" author="Tran" box="[437,674,1796,1822]" pageId="11" pageNumber="512" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="C7716D6A68699F6AFF00F8ED1F58F8CB" blockId="11.[151,1347,1866,1888]" box="[151,1347,1866,1888]" pageId="11" pageNumber="512">
<treatmentCitationGroup id="E7DE4A4468699F6AFF00F8ED1F58F8CB" box="[151,1347,1866,1888]" pageId="11" pageNumber="512">
<taxonomicName id="00CE16E968699F6AFF00F8ED1869F8CB" ID-CoL="4SNHH" authority="Tran &amp; Yang, 2006: 14" authorityName="Tran &amp; Yang" authorityPageNumber="14" authorityYear="2006" box="[151,626,1866,1888]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="anderseni">
<emphasis id="F5BAB17868699F6AFF00F8ED1B99F8CB" box="[151,386,1866,1888]" italics="true" pageId="11" pageNumber="512">Rhyacobates anderseni</emphasis>
<treatmentCitation id="466F4B7B68699F6AFE1EF8EC1869F8CB" author="Tran" box="[393,626,1866,1888]" httpUri="http://treatment.plazi.org/id/CB74DA5DFFC2FFC96FFD56FD97C0FEE6" page="14" pageId="11" pageNumber="512" year="2006">
<bibRefCitation id="A35F109B68699F6AFE1EF8EC184BF8CB" author="Tran" box="[393,592,1866,1888]" pageId="11" pageNumber="512" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
: 14
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16,
<figureCitation id="5FF571EF68699F6AFD32F8ED1909F8CB" box="[677,786,1866,1888]" captionStart-0="FIGURES 1 10" captionStart-1="FIGURES 11 13" captionStart-2="FIGURES 14 23" captionStartId-0="3.[151,264,1911,1934]" captionStartId-1="5.[151,264,1937,1960]" captionStartId-2="6.[151,264,1776,1799]" captionTargetBox-0="[162,1434,207,1867]" captionTargetBox-1="[292,1295,337,1916]" captionTargetBox-2="[160,1424,209,1742]" captionTargetId-0="figure@3.[151,1436,195,1890]" captionTargetId-1="figure@5.[292,1296,337,1917]" captionTargetId-2="figure@6.[151,1436,193,1756]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 1 10. Morphological features of Rhyacobates angustus sp. n. (1) right fore leg of female. (2) basal part of right middle leg of male, ventral view. (3) proctiger of male. (4, 5) left paramere, from different views. (6) endosomal sclerites. (7) body of female, lateral view. (8) abdomen of female, dorsal view, (9, 10) apex of abdomen of female, ventral and lateral views." captionText-1="FIGURES 11 13. (11) Habitat of type locality of Rhyacobates zetteli sp. n. (12, 13) Habitus of R. zetteli sp. n. (12) apterous male. (13) apterous female (photographed by Tran Anh Duc)." captionText-2="FIGURES 14 23. Morphological features of Rhyacobates zetteli sp. n. (14) right fore leg of female. (15) basal part of right middle leg of male, ventral view. (16) proctiger. (17, 18) left paramere, two different views. (19) endosomal sclerites. (20) body of female, lateral view. (21 23) apex of abdomen of female, dorsal, ventral, and lateral views respectively." httpUri-0="https://zenodo.org/record/271685/files/figure.png" httpUri-1="https://zenodo.org/record/271686/files/figure.png" httpUri-2="https://zenodo.org/record/271687/files/figure.png" pageId="11" pageNumber="512">Figs. 716</figureCitation>
,
<figureCitation id="5FF571EF68699F6AFC8BF8ED192EF8CB" box="[796,821,1866,1888]" captionStart="FIGURES 27 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="11" pageNumber="512">27</figureCitation>
(
<typeStatus id="1875D3C868699F6AFCD4F8ED196BF8CB" box="[835,880,1866,1888]" pageId="11" pageNumber="512">type</typeStatus>
locality: Vu Quang, Ha Tinh Prov.,
<collectingCountry id="BFD92DFA68699F6AFB49F8ED1F22F8CB" box="[1246,1337,1866,1888]" name="Vietnam" pageId="11" pageNumber="512">Vietnam</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8FD43EE168699F6AFF00F82A1F77F845" pageId="11" pageNumber="512" type="materials_examined">
<paragraph id="C7716D6A68699F6AFF00F82A19C7F80D" blockId="11.[151,1436,1933,2030]" box="[151,988,1933,1958]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF00F82A1B9BF80D" bold="true" box="[151,384,1933,1958]" pageId="11" pageNumber="512">Material examined.</emphasis>
For
<typeStatus id="1875D3C868699F6AFE22F82A180CF80D" box="[437,535,1933,1958]" pageId="11" pageNumber="512" type="holotype">holotype</typeStatus>
and
<typeStatus id="1875D3C868699F6AFDDAF82918A0F80D" box="[589,699,1934,1958]" pageId="11" pageNumber="512" type="paratype">paratypes</typeStatus>
, see
<bibRefCitation id="A35F109B68699F6AFD66F82919C3F80D" author="Tran" box="[753,984,1933,1958]" pageId="11" pageNumber="512" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang (2006)</bibRefCitation>
.
</paragraph>
<paragraph id="C7716D6A68699F6AFF50F8161F77F845" blockId="11.[151,1436,1933,2030]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF50F8161AE5F861" bold="true" box="[199,254,1969,1994]" pageId="11" pageNumber="512">Size.</emphasis>
Males, length 6.06.1 (allotype 6.0), width 1.831.85 (allotype 1.83) (apterous), length 6.16.2, width 1.851.88 (macropterous); females, length 6.87.0 (
<typeStatus id="1875D3C868699F6AFD4BF872195BF845" box="[732,832,2005,2030]" pageId="11" pageNumber="512" type="holotype">holotype</typeStatus>
6.8), width 2.522.57 (
<typeStatus id="1875D3C868699F6AFBD0F8721EB0F845" box="[1095,1195,2005,2030]" pageId="11" pageNumber="512" type="holotype">holotype</typeStatus>
2.52) (apterous).
</paragraph>
</subSubSection>
<subSubSection id="8FD43EE1686E9F6DFF50FF30189BFDCB" pageId="12" pageNumber="513" type="discussion">
<paragraph id="C7716D6A686E9F6DFF50FF301885FD96" blockId="12.[151,1436,151,645]" pageId="12" pageNumber="513">
<emphasis id="F5BAB178686E9F6DFF50FF301B20FF1B" bold="true" box="[199,315,151,176]" pageId="12" pageNumber="513">Remarks.</emphasis>
The followings are diagnostic characteristics of
<taxonomicName id="00CE16E9686E9F6DFCE3FF3E1E1FFF1B" box="[884,1028,152,176]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="anderseni">
<emphasis id="F5BAB178686E9F6DFCE3FF3E1E1FFF1B" box="[884,1028,152,176]" italics="true" pageId="12" pageNumber="513">R. anderseni</emphasis>
</taxonomicName>
: the mesonotum of the male has a slender yellow stripe on the posterior half; the mesonotum of the female has a broader yellow median marking on the posterior three-fifths; the abdomen of the male is relatively short; in lateral view, abdominal segment 8 of the male has a concave ventral surface; the male proctiger has round angular projections on each side (see
<bibRefCitation id="A35F109B686E9F6DFA9BFEA21ACFFEEB" author="Tran" pageId="12" pageNumber="513" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang 2006</bibRefCitation>
:
<figureCitation id="5FF571EF686E9F6DFF4AFE801B34FEEB" box="[221,303,295,320]" captionStart="FIGURES 11 13" captionStartId="5.[151,264,1937,1960]" captionTargetBox="[292,1295,337,1916]" captionTargetId="figure@5.[292,1296,337,1917]" captionTargetPageId="5" captionText="FIGURES 11 13. (11) Habitat of type locality of Rhyacobates zetteli sp. n. (12, 13) Habitus of R. zetteli sp. n. (12) apterous male. (13) apterous female (photographed by Tran Anh Duc)." httpUri="https://zenodo.org/record/271686/files/figure.png" pageId="12" pageNumber="513">Fig. 12</figureCitation>
); the male paramere is falciform, slightly broad, long, and not conspicuously setose (see
<bibRefCitation id="A35F109B686E9F6DFA9BFE8F1ACFFECE" author="Tran" pageId="12" pageNumber="513" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang 2006</bibRefCitation>
:
<figureCitation id="5FF571EF686E9F6DFF4AFEEB1B22FECF" box="[221,313,332,357]" captionStart="FIGURES 11 13" captionStartId="5.[151,264,1937,1960]" captionTargetBox="[292,1295,337,1916]" captionTargetId="figure@5.[292,1296,337,1917]" captionTargetPageId="5" captionText="FIGURES 11 13. (11) Habitat of type locality of Rhyacobates zetteli sp. n. (12, 13) Habitus of R. zetteli sp. n. (12) apterous male. (13) apterous female (photographed by Tran Anh Duc)." httpUri="https://zenodo.org/record/271686/files/figure.png" pageId="12" pageNumber="513">Figs. 13</figureCitation>
,
<figureCitation id="5FF571EF686E9F6DFED2FEEB1B7FFECF" box="[325,356,332,356]" captionStart="FIGURES 14 23" captionStartId="6.[151,264,1776,1799]" captionTargetBox="[160,1424,209,1742]" captionTargetId="figure@6.[151,1436,193,1756]" captionTargetPageId="6" captionText="FIGURES 14 23. Morphological features of Rhyacobates zetteli sp. n. (14) right fore leg of female. (15) basal part of right middle leg of male, ventral view. (16) proctiger. (17, 18) left paramere, two different views. (19) endosomal sclerites. (20) body of female, lateral view. (21 23) apex of abdomen of female, dorsal, ventral, and lateral views respectively." httpUri="https://zenodo.org/record/271687/files/figure.png" pageId="12" pageNumber="513">14</figureCitation>
); the metanotum of the female has a pointed median process on the posterior margin (see
<bibRefCitation id="A35F109B686E9F6DFADBFEEA1B0DFE23" author="Tran" pageId="12" pageNumber="513" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang 2006</bibRefCitation>
:
<figureCitation id="5FF571EF686E9F6DFEB4FEC81B73FE23" box="[291,360,367,392]" captionStart="FIGURES 1 10" captionStartId="3.[151,264,1911,1934]" captionTargetBox="[162,1434,207,1867]" captionTargetId="figure@3.[151,1436,195,1890]" captionTargetPageId="3" captionText="FIGURES 1 10. Morphological features of Rhyacobates angustus sp. n. (1) right fore leg of female. (2) basal part of right middle leg of male, ventral view. (3) proctiger of male. (4, 5) left paramere, from different views. (6) endosomal sclerites. (7) body of female, lateral view. (8) abdomen of female, dorsal view, (9, 10) apex of abdomen of female, ventral and lateral views." httpUri="https://zenodo.org/record/271685/files/figure.png" pageId="12" pageNumber="513">Fig. 7</figureCitation>
); the abdomen of the female is short (length about 0.2 times body length), sternum 7 is long, almost enclosing the genital segments with its connexival margin raised slightly upwards, with a pair of long posterior projections pointing outwards and downwards (see
<bibRefCitation id="A35F109B686E9F6DFCF6FE1F1E58FE7B" author="Tran" box="[865,1091,439,464]" pageId="12" pageNumber="513" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang 2006</bibRefCitation>
:
<figureCitation id="5FF571EF686E9F6DFBC5FE101EDEFE7B" box="[1106,1221,439,464]" captionStart="FIGURES 1 10" captionStartId="3.[151,264,1911,1934]" captionTargetBox="[162,1434,207,1867]" captionTargetId="figure@3.[151,1436,195,1890]" captionTargetPageId="3" captionText="FIGURES 1 10. Morphological features of Rhyacobates angustus sp. n. (1) right fore leg of female. (2) basal part of right middle leg of male, ventral view. (3) proctiger of male. (4, 5) left paramere, from different views. (6) endosomal sclerites. (7) body of female, lateral view. (8) abdomen of female, dorsal view, (9, 10) apex of abdomen of female, ventral and lateral views." httpUri="https://zenodo.org/record/271685/files/figure.png" pageId="12" pageNumber="513">Figs. 8, 9</figureCitation>
), and its posterior margin is almost straight, bearing two short lateral processes (see
<bibRefCitation id="A35F109B686E9F6DFC3BFE7A1E8BFE5E" author="Tran" box="[940,1168,476,501]" pageId="12" pageNumber="513" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang 2006</bibRefCitation>
:
<figureCitation id="5FF571EF686E9F6DFB08FE7B1EECFE5F" box="[1183,1271,476,501]" captionStart="FIGURES 1 10" captionStartId="3.[151,264,1911,1934]" captionTargetBox="[162,1434,207,1867]" captionTargetId="figure@3.[151,1436,195,1890]" captionTargetPageId="3" captionText="FIGURES 1 10. Morphological features of Rhyacobates angustus sp. n. (1) right fore leg of female. (2) basal part of right middle leg of male, ventral view. (3) proctiger of male. (4, 5) left paramere, from different views. (6) endosomal sclerites. (7) body of female, lateral view. (8) abdomen of female, dorsal view, (9, 10) apex of abdomen of female, ventral and lateral views." httpUri="https://zenodo.org/record/271685/files/figure.png" pageId="12" pageNumber="513">Fig. 10</figureCitation>
). For detailed comparisons of
<taxonomicName id="00CE16E9686E9F6DFECCFDA71873FDB3" box="[347,616,512,536]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="anderseni">
<emphasis id="F5BAB178686E9F6DFECCFDA71873FDB3" box="[347,616,512,536]" italics="true" pageId="12" pageNumber="513">Rhyacobates anderseni</emphasis>
</taxonomicName>
with its congeners and other ptilomerine genera (
<emphasis id="F5BAB178686E9F6DFB4EFDA71F7FFDB3" box="[1241,1380,512,536]" italics="true" pageId="12" pageNumber="513">Andersenius</emphasis>
and
<taxonomicName id="00CE16E9686E9F6DFF00FD831B01FD97" box="[151,282,548,572]" class="Insecta" family="Gerridae" genus="Pleciobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="genus">
<emphasis id="F5BAB178686E9F6DFF00FD831B01FD97" box="[151,282,548,572]" italics="true" pageId="12" pageNumber="513">Pleciobates</emphasis>
</taxonomicName>
), refer to
<bibRefCitation id="A35F109B686E9F6DFE1EFD821881FD96" author="Tran" box="[393,666,548,573]" pageId="12" pageNumber="513" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang (2006: 16)</bibRefCitation>
.
</paragraph>
<paragraph id="C7716D6A686E9F6DFF50FDE0189BFDCB" blockId="12.[151,1436,151,645]" box="[199,640,583,608]" pageId="12" pageNumber="513">
<emphasis id="F5BAB178686E9F6DFF50FDE01B2EFDCB" bold="true" box="[199,309,583,608]" pageId="12" pageNumber="513">Habitats.</emphasis>
See
<bibRefCitation id="A35F109B686E9F6DFEFCFDEF1860FDCB" author="Tran" box="[363,635,583,608]" pageId="12" pageNumber="513" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang (2006: 16)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8FD43EE1686E9F6DFF50FDCB18F6FD2F" box="[199,749,620,645]" pageId="12" pageNumber="513" type="distribution">
<paragraph id="C7716D6A686E9F6DFF50FDCB18F6FD2F" blockId="12.[151,1436,151,645]" box="[199,749,620,645]" pageId="12" pageNumber="513">
<emphasis id="F5BAB178686E9F6DFF50FDCB1B7BFD2E" bold="true" box="[199,352,620,645]" pageId="12" pageNumber="513">Distribution.</emphasis>
<collectingCountry id="BFD92DFA686E9F6DFEF0FDCB1BD1FD2E" box="[359,458,620,645]" name="Vietnam" pageId="12" pageNumber="513">Vietnam</collectingCountry>
: Ha Tinh.
<collectingCountry id="BFD92DFA686E9F6DFDA9FDCB189EFD2E" box="[574,645,620,645]" name="China" pageId="12" pageNumber="513">China</collectingCountry>
: Yunnan.
</paragraph>
</subSubSection>
</treatment>
</document>