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<document id="E66E0430EF0E1EBA05DC52A9B94CA3D2" ID="10.11646/zootaxa.4121.5.1" ID-CLB-Dataset="38884" ID-DOI="10.11646/zootaxa.4121.5.1" ID-GBIF-Dataset="50bb9c19-8df5-4fbf-a78b-f8ba2d6355bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="271684" ID-ZooBank="BCADF85B-CB84-4694-963A-AF9F0E3F40D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465798066874" checkinUser="plazi" docAuthor="Tran, A. D. &amp; Nguyen, X. Q." docDate="2016" docId="4F67DC7C68699F6AFF00FC081902F91B" docLanguage="en" docName="zootaxa.4121.5.1.pdf" docOrigin="Zootaxa 4121 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacobates abdominalis Andersen &amp; Chen 1995" docType="treatment" docVersion="10" lastPageNumber="512" masterDocId="B35EA40468629F61FF97FFA71A1BFFAB" masterDocTitle="Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam" masterLastPageNumber="516" masterPageNumber="501" pageNumber="512" updateTime="1698662823377" updateUser="ExternalLinkService">
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<mods:title id="7AE0A4B684E9B3AB082B0B4D52D25F6A">Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam</mods:title>
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<treatment id="4F67DC7C68699F6AFF00FC081902F91B" ID-DOI="http://doi.org/10.5281/zenodo.4509712" ID-GBIF-Taxon="120827706" ID-Zenodo-Dep="4509712" LSID="urn:lsid:plazi:treatment:4F67DC7C68699F6AFF00FC081902F91B" httpUri="http://treatment.plazi.org/id/4F67DC7C68699F6AFF00FC081902F91B" lastPageNumber="512" pageId="11" pageNumber="512">
<subSubSection id="8FD43EE168699F6AFF00FC0818E4FC61" box="[151,767,943,970]" pageId="11" pageNumber="512" type="nomenclature">
<paragraph id="C7716D6A68699F6AFF00FC0818E4FC61" blockId="11.[151,767,943,970]" box="[151,767,943,970]" pageId="11" pageNumber="512">
<heading id="9C39DA0668699F6AFF00FC0818E4FC61" bold="true" box="[151,767,943,970]" fontSize="11" level="1" pageId="11" pageNumber="512" reason="1">
<taxonomicName id="00CE16E968699F6AFF00FC0818E4FC61" ID-CoL="4SNHG" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[151,767,943,970]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="F5BAB17868699F6AFF00FC0818E4FC61" bold="true" box="[151,767,943,970]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF00FC081BD0FC62" bold="true" box="[151,459,943,969]" italics="true" pageId="11" pageNumber="512">Rhyacobates abdominalis</emphasis>
<bibRefCitation id="A35F109B68699F6AFE44FC1718E4FC61" author="Andersen" box="[467,767,944,970]" pageId="11" pageNumber="512" refString="Andersen, N. M. &amp; Chen, P. - P. (1995) A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie, 138, 51 - 67. Chen, P-P., Nieser, N. &amp; Zettel, H. (2005) The aquatic and semi-aquatic bugs (Heteroptera: Nepomorpha &amp; Gerromorpha) of Malesia. Fauna Malesiana Handbooks 5. Brill, Leiden, / Boston, 546 pp." type="book chapter" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8FD43EE168699F6AFF00FC5118A5FB80" pageId="11" pageNumber="512" type="reference_group">
<paragraph id="C7716D6A68699F6AFF00FC5118A5FB80" blockId="11.[151,1436,1014,1067]" pageId="11" pageNumber="512">
<treatmentCitationGroup id="E7DE4A4468699F6AFF00FC5118A5FB80" pageId="11" pageNumber="512">
<taxonomicName id="00CE16E968699F6AFF00FC5118D6FBA7" ID-CoL="4SNHG" authority="Andersen &amp; Chen, 1995: 58" authorityName="Andersen &amp; Chen" authorityPageNumber="58" authorityYear="1995" box="[151,717,1014,1036]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="F5BAB17868699F6AFF00FC511B85FBA7" box="[151,414,1014,1036]" italics="true" pageId="11" pageNumber="512">Rhyacobates abdominalis</emphasis>
<treatmentCitation id="466F4B7B68699F6AFE31FC5118D6FBA7" author="Andersen" box="[422,717,1014,1036]" page="58" pageId="11" pageNumber="512" year="1995">
<bibRefCitation id="A35F109B68699F6AFE31FC5118BCFBA7" author="Andersen" box="[422,679,1014,1036]" pageId="11" pageNumber="512" refString="Andersen, N. M. &amp; Chen, P. - P. (1995) A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie, 138, 51 - 67. Chen, P-P., Nieser, N. &amp; Zettel, H. (2005) The aquatic and semi-aquatic bugs (Heteroptera: Nepomorpha &amp; Gerromorpha) of Malesia. Fauna Malesiana Handbooks 5. Brill, Leiden, / Boston, 546 pp." type="book chapter" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
: 58
</treatmentCitation>
</taxonomicName>
59,
<figureCitation id="5FF571EF68699F6AFC96FC511960FBA7" box="[769,891,1014,1036]" captionStart-0="FIGURES 11 13" captionStart-1="FIGURES 14 23" captionStartId-0="5.[151,264,1937,1960]" captionStartId-1="6.[151,264,1776,1799]" captionTargetBox-0="[292,1295,337,1916]" captionTargetBox-1="[160,1424,209,1742]" captionTargetId-0="figure@5.[292,1296,337,1917]" captionTargetId-1="figure@6.[151,1436,193,1756]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 11 13. (11) Habitat of type locality of Rhyacobates zetteli sp. n. (12, 13) Habitus of R. zetteli sp. n. (12) apterous male. (13) apterous female (photographed by Tran Anh Duc)." captionText-1="FIGURES 14 23. Morphological features of Rhyacobates zetteli sp. n. (14) right fore leg of female. (15) basal part of right middle leg of male, ventral view. (16) proctiger. (17, 18) left paramere, two different views. (19) endosomal sclerites. (20) body of female, lateral view. (21 23) apex of abdomen of female, dorsal, ventral, and lateral views respectively." httpUri-0="https://zenodo.org/record/271686/files/figure.png" httpUri-1="https://zenodo.org/record/271687/files/figure.png" pageId="11" pageNumber="512">Figs. 1215</figureCitation>
(
<typeStatus id="1875D3C868699F6AFC1CFC5119A3FBA7" box="[907,952,1014,1036]" pageId="11" pageNumber="512">type</typeStatus>
locality: Ruyang Nat. Res., Guangdong Prov.,
<collectingCountry id="BFD92DFA68699F6AFF50FBB21B13FB80" box="[199,264,1045,1067]" name="China" pageId="11" pageNumber="512">China</collectingCountry>
);
<treatmentCitation id="466F4B7B68699F6AFE8EFBB11819FB80" author="Tran" box="[281,514,1045,1067]" page="19" pageId="11" pageNumber="512" year="2006">
<bibRefCitation id="A35F109B68699F6AFE8EFBB11BFBFB80" author="Tran" box="[281,480,1045,1067]" pageId="11" pageNumber="512" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
: 19
</treatmentCitation>
(record
<collectingCountry id="BFD92DFA68699F6AFDCFFBB218A8FB80" box="[600,691,1045,1067]" name="Vietnam" pageId="11" pageNumber="512">Vietnam</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="8FD43EE168699F6AFF00FBF01AF1FA8E" pageId="11" pageNumber="512" type="materials_examined">
<paragraph id="C7716D6A68699F6AFF00FBF01839FB13" blockId="11.[151,1437,1111,1712]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF00FBF01B98FBDB" bold="true" box="[151,387,1111,1136]" pageId="11" pageNumber="512">Material examined.</emphasis>
<typeStatus id="1875D3C868699F6AFE1AFBFF1BE5FBDB" box="[397,510,1112,1136]" pageId="11" pageNumber="512" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="D1C8A6E368699F6AFD9BFBFF1846FBDB" box="[524,605,1111,1136]" pageId="11" pageNumber="512" type="male">1 male</specimenCount>
,
<specimenCount id="D1C8A6E368699F6AFDFCFBFF18D4FBDB" box="[619,719,1111,1136]" pageId="11" pageNumber="512" type="female">1 female</specimenCount>
(apterous), “
<collectingCountry id="BFD92DFA68699F6AFCF1FBF019B4FBDB" box="[870,943,1111,1136]" name="China" pageId="11" pageNumber="512">China</collectingCountry>
: Guangdong Prov., Ruyang Nat. Res., Lao Peng Keng stream,
<quantity id="0036C08F68699F6AFEE5FBDB1BD9FB3F" box="[370,450,1148,1172]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="512" unit="m" value="1100.0">1100m</quantity>
,
<date id="B3704BAA68699F6AFE5AFBDB185BFB3F" box="[461,576,1148,1172]" pageId="11" pageNumber="512" value="1990-08-14">14.8.1990</date>
, coll. P.P. Chen, C9012” (
<collectionCode id="A1DFF5AF68699F6AFCF6FBDB19DFFB3F" box="[865,964,1148,1172]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="11" pageNumber="512">NHMW</collectionCode>
). For formerly examined specimens (from
<collectingCountry id="BFD92DFA68699F6AFF00FB381AE1FB13" box="[151,250,1183,1208]" name="Vietnam" pageId="11" pageNumber="512">Vietnam</collectingCountry>
), see
<bibRefCitation id="A35F109B68699F6AFEAFFB071805FB13" author="Tran" box="[312,542,1183,1208]" pageId="11" pageNumber="512" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang (2006)</bibRefCitation>
.
</paragraph>
<paragraph id="C7716D6A68699F6AFF50FB631AF1FA8E" blockId="11.[151,1437,1111,1712]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF50FB631AE5FB76" bold="true" box="[199,254,1220,1245]" pageId="11" pageNumber="512">Size.</emphasis>
Males, length 6.87.0, width 2.102.20 (apterous), length 6.46.6, width 2.062.11 (macropterous, dealated); females, length 9.910.0, width 3.50 (apterous), length 9.49.6, width 3.143.30 (macropterous, dealated).
</paragraph>
</subSubSection>
<subSubSection id="8FD43EE168699F6AFF50FA8818B1F927" pageId="11" pageNumber="512" type="discussion">
<paragraph id="C7716D6A68699F6AFF50FA881842F9C3" blockId="11.[151,1437,1111,1712]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF50FA881B20FAE3" bold="true" box="[199,315,1327,1352]" pageId="11" pageNumber="512">Remarks.</emphasis>
<taxonomicName id="00CE16E968699F6AFED1FA971870FAE3" box="[326,619,1328,1352]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="F5BAB17868699F6AFED1FA971870FAE3" box="[326,619,1328,1352]" italics="true" pageId="11" pageNumber="512">Rhyacobates abdominalis</emphasis>
</taxonomicName>
can be recognised by the following diagnostic characters: abdominal segment 8 of the male is ventrolaterally impressed; the proctiger of the male has a small angular projection on each side; the male paramere is slender with a broad basal part and hook-shaped apex (see
<bibRefCitation id="A35F109B68699F6AFBC0FAD01F7EFA3B" author="Andersen" box="[1111,1381,1399,1424]" pageId="11" pageNumber="512" refString="Andersen, N. M. &amp; Chen, P. - P. (1995) A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie, 138, 51 - 67. Chen, P-P., Nieser, N. &amp; Zettel, H. (2005) The aquatic and semi-aquatic bugs (Heteroptera: Nepomorpha &amp; Gerromorpha) of Malesia. Fauna Malesiana Handbooks 5. Brill, Leiden, / Boston, 546 pp." type="book chapter" year="1995">Andersen &amp; Chen 1995</bibRefCitation>
:
<figureCitation id="5FF571EF68699F6AFAE7FAD01AADFA1E" captionStart="FIGURES 14 23" captionStartId="6.[151,264,1776,1799]" captionTargetBox="[160,1424,209,1742]" captionTargetId="figure@6.[151,1436,193,1756]" captionTargetPageId="6" captionText="FIGURES 14 23. Morphological features of Rhyacobates zetteli sp. n. (14) right fore leg of female. (15) basal part of right middle leg of male, ventral view. (16) proctiger. (17, 18) left paramere, two different views. (19) endosomal sclerites. (20) body of female, lateral view. (21 23) apex of abdomen of female, dorsal, ventral, and lateral views respectively." httpUri="https://zenodo.org/record/271687/files/figure.png" pageId="11" pageNumber="512">Fig. 15</figureCitation>
); the abdomen of the female is distinctly curved dorsad, abdominal segment 7 possesses a short projection on each connexivum, and the posterior margin of sternum 7 has a sharp, small median process (see
<bibRefCitation id="A35F109B68699F6AFB45FA181AC8FA56" author="Andersen" pageId="11" pageNumber="512" refString="Andersen, N. M. &amp; Chen, P. - P. (1995) A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie, 138, 51 - 67. Chen, P-P., Nieser, N. &amp; Zettel, H. (2005) The aquatic and semi-aquatic bugs (Heteroptera: Nepomorpha &amp; Gerromorpha) of Malesia. Fauna Malesiana Handbooks 5. Brill, Leiden, / Boston, 546 pp." type="book chapter" year="1995">Andersen &amp; Chen 1995</bibRefCitation>
:
<figureCitation id="5FF571EF68699F6AFF48FA431B72FA57" box="[223,361,1508,1533]" captionStart-0="FIGURES 11 13" captionStart-1="FIGURES 14 23" captionStartId-0="5.[151,264,1937,1960]" captionStartId-1="6.[151,264,1776,1799]" captionTargetBox-0="[292,1295,337,1916]" captionTargetBox-1="[160,1424,209,1742]" captionTargetId-0="figure@5.[292,1296,337,1917]" captionTargetId-1="figure@6.[151,1436,193,1756]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 11 13. (11) Habitat of type locality of Rhyacobates zetteli sp. n. (12, 13) Habitus of R. zetteli sp. n. (12) apterous male. (13) apterous female (photographed by Tran Anh Duc)." captionText-1="FIGURES 14 23. Morphological features of Rhyacobates zetteli sp. n. (14) right fore leg of female. (15) basal part of right middle leg of male, ventral view. (16) proctiger. (17, 18) left paramere, two different views. (19) endosomal sclerites. (20) body of female, lateral view. (21 23) apex of abdomen of female, dorsal, ventral, and lateral views respectively." httpUri-0="https://zenodo.org/record/271686/files/figure.png" httpUri-1="https://zenodo.org/record/271687/files/figure.png" pageId="11" pageNumber="512">Figs. 1214</figureCitation>
). The specimens of
<taxonomicName id="00CE16E968699F6AFDD9FA4218E2FA57" box="[590,761,1508,1532]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="F5BAB17868699F6AFDD9FA4218E2FA57" box="[590,761,1508,1532]" italics="true" pageId="11" pageNumber="512">R. abdominalis</emphasis>
</taxonomicName>
from
<collectingCountry id="BFD92DFA68699F6AFCD7FA431984FA56" box="[832,927,1508,1533]" name="Vietnam" pageId="11" pageNumber="512">Vietnam</collectingCountry>
were not significantly different compared to the
<typeStatus id="1875D3C868699F6AFF55F9AF1B36F98B" box="[194,301,1544,1568]" pageId="11" pageNumber="512" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="BFD92DFA68699F6AFEE2F9A01BA0F98B" box="[373,443,1543,1568]" name="China" pageId="11" pageNumber="512">China</collectingCountry>
, except for the larger size and less pointed median process of sternum
<quantity id="0036C08F68699F6AFB7EF9AF1F0EF98B" box="[1257,1301,1543,1568]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="11" pageNumber="512" unit="in" value="7.0">7 in</quantity>
the female. They were collected from the same habitat as
<taxonomicName id="00CE16E968699F6AFD49F98B19D4F9EF" box="[734,975,1580,1604]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="gongvo">
<emphasis id="F5BAB17868699F6AFD49F98B19D4F9EF" box="[734,975,1580,1604]" italics="true" pageId="11" pageNumber="512">Rhyacobates gongvo</emphasis>
</taxonomicName>
. For description notes of Vietnamese specimens, see
<bibRefCitation id="A35F109B68699F6AFED3F9F7184EF9C3" author="Tran" box="[324,597,1615,1640]" pageId="11" pageNumber="512" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang (2006: 19)</bibRefCitation>
.
</paragraph>
<paragraph id="C7716D6A68699F6AFF50F9D318B1F927" blockId="11.[151,1437,1111,1712]" box="[199,682,1652,1677]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF50F9D31B2EF926" bold="true" box="[199,309,1652,1677]" pageId="11" pageNumber="512">Habitats.</emphasis>
See
<bibRefCitation id="A35F109B68699F6AFEFCF9D218BDF927" author="Tran" box="[363,678,1652,1677]" pageId="11" pageNumber="512" refString="Tran, A. D. &amp; Yang, C. M. (2006) New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology, 54 (1), 11 - 20." type="journal article" year="2006">Tran &amp; Yang (2006: 18, 19)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8FD43EE168699F6AFF50F9301902F91B" box="[199,793,1687,1712]" pageId="11" pageNumber="512" type="distribution">
<paragraph id="C7716D6A68699F6AFF50F9301902F91B" blockId="11.[151,1437,1111,1712]" box="[199,793,1687,1712]" pageId="11" pageNumber="512">
<emphasis id="F5BAB17868699F6AFF50F9301B7BF91B" bold="true" box="[199,352,1687,1712]" pageId="11" pageNumber="512">Distribution.</emphasis>
<collectingCountry id="BFD92DFA68699F6AFEF0F9301BD1F91B" box="[359,458,1687,1712]" name="Vietnam" pageId="11" pageNumber="512">Vietnam</collectingCountry>
: Lao Cai.
<collectingCountry id="BFD92DFA68699F6AFDABF9301898F91B" box="[572,643,1687,1712]" name="China" pageId="11" pageNumber="512">China</collectingCountry>
: Guangdong.
</paragraph>
</subSubSection>
</treatment>
</document>