429 lines
62 KiB
XML
429 lines
62 KiB
XML
<document id="7F9CCC4698EDAB79FEA50C32F2584D45" ID="10.11646/zootaxa.4121.5.1" ID-CLB-Dataset="38884" ID-DOI="10.11646/zootaxa.4121.5.1" ID-GBIF-Dataset="50bb9c19-8df5-4fbf-a78b-f8ba2d6355bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="271684" ID-ZooBank="BCADF85B-CB84-4694-963A-AF9F0E3F40D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465798066874" checkinUser="plazi" docAuthor="Tran, A. D. & Nguyen, X. Q." docDate="2016" docId="4F67DC7C68659F6AFF00FA481827FCF6" docLanguage="en" docName="zootaxa.4121.5.1.pdf" docOrigin="Zootaxa 4121 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacobates constrictus Tran & Nguyen, 2016, sp.n." docType="treatment" docVersion="10" lastPageNumber="512" masterDocId="B35EA40468629F61FF97FFA71A1BFFAB" masterDocTitle="Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam" masterLastPageNumber="516" masterPageNumber="501" pageNumber="508" updateTime="1698662823377" updateUser="ExternalLinkService">
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<mods:title id="6BC10FBCA25CEF1FFAEBBFA3E6B00EC0">Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam</mods:title>
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<mods:namePart id="72B864C3A8928DDFF5C3B20371277734">Tran, A. D.</mods:namePart>
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<mods:namePart id="F4F9A98C349C88C7A999D22DC623F36D">Nguyen, X. Q.</mods:namePart>
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<mods:title id="1B1CED2AFBFA30099EE2A27C667A2CC5">Zootaxa</mods:title>
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<mods:part id="1BAACED9AD5BA28C19FF606827C8A498">
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<mods:date id="A578A851A793C69E5073E595CC2F714F">2016</mods:date>
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<mods:number id="9791DE6615D9EDC2C30FD83AA04D663B">4121</mods:number>
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<treatment id="4F67DC7C68659F6AFF00FA481827FCF6" ID-DOI="http://doi.org/10.5281/zenodo.4509710" ID-GBIF-Taxon="120827708" ID-Zenodo-Dep="4509710" LSID="urn:lsid:plazi:treatment:4F67DC7C68659F6AFF00FA481827FCF6" httpUri="http://treatment.plazi.org/id/4F67DC7C68659F6AFF00FA481827FCF6" lastPageId="11" lastPageNumber="512" pageId="7" pageNumber="508">
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<subSubSection id="8FD43EE168659F66FF00FA481B35F980" pageId="7" pageNumber="508" type="nomenclature">
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<paragraph id="C7716D6A68659F66FF00FA481BE6F9A1" blockId="7.[151,509,1519,1579]" box="[151,509,1519,1546]" pageId="7" pageNumber="508">
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<heading id="9C39DA0668659F66FF00FA481BE6F9A1" bold="true" box="[151,509,1519,1546]" fontSize="11" level="1" pageId="7" pageNumber="508" reason="1">
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<emphasis id="F5BAB17868659F66FF00FA481BE6F9A1" bold="true" box="[151,509,1519,1546]" pageId="7" pageNumber="508">
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<taxonomicName id="00CE16E968659F66FF00FA481BA0F9A2" ID-CoL="4SNHL" box="[151,443,1519,1545]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="508" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
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<emphasis id="F5BAB17868659F66FF00FA481BA0F9A2" bold="true" box="[151,443,1519,1545]" italics="true" pageId="7" pageNumber="508">Rhyacobates constrictus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="EE890C0368659F66FE55FA571BE6F9A1" box="[450,509,1520,1546]" pageId="7" pageNumber="508" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C7716D6A68659F66FF00F9B51B35F980" blockId="7.[151,509,1519,1579]" box="[151,302,1554,1579]" pageId="7" pageNumber="508">
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(
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<figureCitation id="5FF571EF68659F66FF08F9B51B3DF980" box="[159,294,1554,1579]" captionStart-0="FIGURES 24 – 26" captionStart-1="FIGURES 27 – 39" captionStartId-0="9.[151,264,1633,1656]" captionStartId-1="10.[151,264,1801,1824]" captionTargetBox-0="[151,1436,193,1610]" captionTargetBox-1="[168,1417,201,1747]" captionTargetId-0="figure@9.[151,1436,193,1612]" captionTargetId-1="figure@10.[151,1436,193,1781]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 24 – 26. (24) Habitat of type locality of Rhyacobates constrictus sp. n. (25) A copulating pair of R. constrictus sp. n., skating on the fast current of the stream. (26) A copulating pair of R. constrictus sp. n., resting on the emerged rocks in the middle of the stream (photographed by Tran Anh Duc)." captionText-1="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri-0="https://zenodo.org/record/271688/files/figure.png" httpUri-1="https://zenodo.org/record/271689/files/figure.png" pageId="7" pageNumber="508">Figs. 25–39</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="8FD43EE168659F69FF00F9FD1F9BFDB3" lastPageId="8" lastPageNumber="509" pageId="7" pageNumber="508" type="materials_examined">
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<paragraph id="C7716D6A68659F66FF00F9FD1BE1F910" blockId="7.[151,1437,1626,2011]" pageId="7" pageNumber="508">
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<emphasis id="F5BAB17868659F66FF00F9FD1B9AF9D8" bold="true" box="[151,385,1626,1651]" pageId="7" pageNumber="508">Material examined.</emphasis>
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<typeStatus id="1875D3C868659F66FE1DF9FD1BE9F9D8" box="[394,498,1626,1651]" pageId="7" pageNumber="508" type="holotype">Holotype</typeStatus>
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||
(apterous female) and allotype (apterous male),
|
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<collectingCountry id="BFD92DFA68659F66FB8BF9FC1EBBF9D8" box="[1052,1184,1627,1651]" name="Vietnam" pageId="7" pageNumber="508">VIETNAM</collectingCountry>
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, Phu Tho Prov., Xuan Son N’Park, Lap stream, site 1, at Ngoc waterfall,
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<geoCoordinate id="A2FA0BAD68659F66FD47F9D81979F93C" box="[720,866,1663,1687]" direction="north" orientation="latitude" pageId="7" pageNumber="508" precision="1" value="21.130283">21°07.817’N</geoCoordinate>
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<geoCoordinate id="A2FA0BAD68659F66FCFEF9D91E13F93C" box="[873,1032,1662,1687]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="1" value="104.9251">104°55.506’E</geoCoordinate>
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,
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<quantity id="0036C08F68659F66FB83F9D81E40F93D" box="[1044,1115,1663,1687]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="7" pageNumber="508" unit="m" value="390.0">390 m</quantity>
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||
asl., coll. Tran A.D.
|
||
<emphasis id="F5BAB17868659F66FAD0F9D81F9BF93D" box="[1351,1408,1662,1686]" italics="true" pageId="7" pageNumber="508">et al.</emphasis>
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,
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<date id="B3704BAA68659F66FA19F9D91B17F910" pageId="7" pageNumber="508" value="2013-06-05">5 June 2013</date>
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, TAD1331 (
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).
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</paragraph>
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<paragraph id="C7716D6A68659F69FF50F9601F9BFDB3" blockId="7.[151,1437,1626,2011]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="509" pageId="7" pageNumber="508">
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<typeStatus id="1875D3C868659F66FF50F9601B22F974" box="[199,313,1735,1759]" pageId="7" pageNumber="508" type="paratype">Paratypes</typeStatus>
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:
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<collectingCountry id="BFD92DFA68659F66FED2F9601BD0F974" box="[325,459,1735,1759]" name="Vietnam" pageId="7" pageNumber="508">VIETNAM</collectingCountry>
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:
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<emphasis id="F5BAB17868659F66FE4FF9611890F974" bold="true" box="[472,651,1734,1759]" pageId="7" pageNumber="508">Phu Tho Prov.</emphasis>
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:
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,
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(apterous),
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<specimenCount id="D1C8A6E368659F66FB93F9601E73F974" box="[1028,1128,1734,1759]" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
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(macropterous, de-alated), same locality data as
|
||
<typeStatus id="1875D3C868659F66FE12F94D1BFCF8A8" box="[389,487,1770,1795]" pageId="7" pageNumber="508" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="A1DFF5AF68659F66FE61F94C1856F8A8" box="[502,589,1771,1795]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="7" pageNumber="508">ZMHU</collectionCode>
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,
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<collectionCode id="A1DFF5AF68659F66FDC0F94C1889F8A8" box="[599,658,1771,1795]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="508">ZRC</collectionCode>
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);
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<specimenCount id="D1C8A6E368659F66FD31F94C18E5F8A8" box="[678,766,1770,1795]" pageId="7" pageNumber="508" type="male">2 males</specimenCount>
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,
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<specimenCount id="D1C8A6E368659F66FC9EF94C196EF8A8" box="[777,885,1770,1795]" pageId="7" pageNumber="508" type="female">3 females</specimenCount>
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(apterous), Xuan Son N’Park, Lap stream, site 2, first concrete bridge from Ngoc waterfall,
|
||
<geoCoordinate id="A2FA0BAD68659F66FDE8F8A8190AF88C" box="[639,785,1807,1831]" direction="north" orientation="latitude" pageId="7" pageNumber="508" precision="1" value="21.133467">21°08.008’N</geoCoordinate>
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<geoCoordinate id="A2FA0BAD68659F66FC8DF8A919A2F88C" box="[794,953,1806,1831]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="1" value="104.92947">104°55.768’E</geoCoordinate>
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,
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<quantity id="0036C08F68659F66FC50F8A81E0BF88D" box="[967,1040,1807,1831]" metricMagnitude="2" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="508" unit="m" value="330.0">330 m</quantity>
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||
asl., coll. Tran A.D.
|
||
<emphasis id="F5BAB17868659F66FA90F8A81F5AF88D" box="[1287,1345,1806,1830]" italics="true" pageId="7" pageNumber="508">et al.</emphasis>
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,
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<date id="B3704BAA68659F66FAC6F8A91AC9F8E0" pageId="7" pageNumber="508" value="2013-06-05">5 June 2013</date>
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, TAD1332 (
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<collectionCode id="A1DFF5AF68659F66FECCF8941BAFF8E0" box="[347,436,1843,1867]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="7" pageNumber="508">ZMHU</collectionCode>
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);
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<specimenCount id="D1C8A6E368659F66FE5FF8941832F8E0" box="[456,553,1842,1867]" pageId="7" pageNumber="508" type="female">1 female</specimenCount>
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(apterous), Xuan Son N’Park, Lap stream, site 3, second concrete bridge from Ngoc waterfall,
|
||
<geoCoordinate id="A2FA0BAD68659F66FED9F8F01BFBF8C4" box="[334,480,1879,1903]" direction="north" orientation="latitude" pageId="7" pageNumber="508" precision="1" value="21.138317">21°08.299’N</geoCoordinate>
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<geoCoordinate id="A2FA0BAD68659F66FE7FF8F11861F8C4" box="[488,634,1878,1903]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="9" value="104.93767">104°56.26’E</geoCoordinate>
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,
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<quantity id="0036C08F68659F66FD11F8F118D5F8C5" box="[646,718,1878,1903]" metricMagnitude="2" metricUnit="m" metricValue="2.68" pageId="7" pageNumber="508" unit="m" value="268.0">268 m</quantity>
|
||
asl., coll. Tran A.D.
|
||
<emphasis id="F5BAB17868659F66FC57F8F019E2F8C5" box="[960,1017,1878,1902]" italics="true" pageId="7" pageNumber="508">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA68659F66FB9FF8F11E8EF8C4" box="[1032,1173,1878,1903]" pageId="7" pageNumber="508" value="2013-06-05">5 June 2013</date>
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, TAD1333 (
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<collectionCode id="A1DFF5AF68659F66FAB6F8F01F61F8C4" box="[1313,1402,1879,1903]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="7" pageNumber="508">ZMHU</collectionCode>
|
||
);
|
||
<specimenCount id="D1C8A6E368659F66FA19F8F01AD6F838" pageId="7" pageNumber="508" type="male">1 male</specimenCount>
|
||
(apterous), Xuan Son N’Park, Kim Thuong, Tan Ong stream, site 2, ca.
|
||
<quantity id="0036C08F68659F66FBB8F8DD1E7BF838" box="[1071,1120,1914,1939]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="508" unit="km" value="2.0">2km</quantity>
|
||
from
|
||
<collectingCountry id="BFD92DFA68659F66FB3AF8DD1EF8F838" box="[1197,1251,1914,1939]" name="China" pageId="7" pageNumber="508">Chin</collectingCountry>
|
||
Tang waterfall,
|
||
<geoCoordinate id="A2FA0BAD68659F66FF00F8381B32F81C" box="[151,297,1951,1975]" direction="north" orientation="latitude" pageId="7" pageNumber="508" precision="1" value="21.070017">21°04.201’N</geoCoordinate>
|
||
<geoCoordinate id="A2FA0BAD68659F66FEA4F8391BC9F81C" box="[307,466,1950,1975]" direction="east" orientation="longitude" pageId="7" pageNumber="508" precision="1" value="104.94865">104°56.919’E</geoCoordinate>
|
||
,
|
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<quantity id="0036C08F68659F66FE77F8381830F81D" box="[480,555,1951,1975]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="508" unit="m" value="340.0">340 m</quantity>
|
||
asl., coll. Tran A.D.
|
||
<emphasis id="F5BAB17868659F66FCB2F838197AF81D" box="[805,865,1950,1974]" italics="true" pageId="7" pageNumber="508">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA68659F66FCE5F8381E36F81C" box="[882,1069,1951,1975]" pageId="7" pageNumber="508" value="2013-08-28">28 August 2013</date>
|
||
, TAD1341 (
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<collectionCode id="A1DFF5AF68659F66FB29F8381F0CF81C" box="[1214,1303,1951,1975]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="7" pageNumber="508">ZMHU</collectionCode>
|
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);
|
||
<specimenCount id="D1C8A6E368659F66FABAF8381F87F81C" box="[1325,1436,1950,1975]" pageId="7" pageNumber="508" type="female">2 females</specimenCount>
|
||
(apterous), Xuan Son N’Park, Lap stream, site 1, at Ngoc waterfall, coll. Tran A.D.
|
||
<emphasis id="F5BAB17868659F66FB1FF8641EDEF870" box="[1160,1221,1987,2011]" italics="true" pageId="7" pageNumber="508">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA68659F66FB4FF8641F83F870" box="[1240,1432,1987,2011]" pageId="7" pageNumber="508" value="2013-08-30">30 August 2013</date>
|
||
, TAD1352 (GPS data same as TAD1331) (
|
||
<collectionCode id="A1DFF5AF686A9F69FDE3FF3F18D6FF1B" box="[628,717,152,176]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="509">ZMHU</collectionCode>
|
||
);
|
||
<specimenCount id="D1C8A6E3686A9F69FD76FF3F1920FF1B" box="[737,827,151,176]" pageId="8" pageNumber="509" type="male">2 males</specimenCount>
|
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,
|
||
<specimenCount id="D1C8A6E3686A9F69FCDFFF3F19AEFF1B" box="[840,949,151,176]" pageId="8" pageNumber="509" type="female">2 females</specimenCount>
|
||
(apterous), Xuan Son N’Park, Lap stream, site 2, first concrete bridge from Ngoc waterfall, coll. Tran A.D.
|
||
<emphasis id="F5BAB178686A9F69FCFBFF1A19BFFF7F" box="[876,932,188,212]" italics="true" pageId="8" pageNumber="509">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA686A9F69FC25FF1B1E73FF7F" box="[946,1128,188,212]" pageId="8" pageNumber="509" value="2013-08-30">30 August 2013</date>
|
||
, TAD1353 (GPS data same as TAD1332) (
|
||
<collectionCode id="A1DFF5AF686A9F69FED3FF471B86FF53" box="[324,413,224,248]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="509">ZMHU</collectionCode>
|
||
);
|
||
<specimenCount id="D1C8A6E3686A9F69FE23FF78180AFF53" box="[436,529,223,248]" pageId="8" pageNumber="509" type="male">5 males</specimenCount>
|
||
,
|
||
<specimenCount id="D1C8A6E3686A9F69FDB6FF47188BFF53" box="[545,656,223,248]" pageId="8" pageNumber="509" type="female">2 females</specimenCount>
|
||
(apterous), Xuan Son N’Park, Lap stream, site 3, second concrete bridge from Ngoc waterfall, coll. Tran A.D.
|
||
<emphasis id="F5BAB178686A9F69FD32FEA218FAFEB7" box="[677,737,260,284]" italics="true" pageId="8" pageNumber="509">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA686A9F69FD64FEA319AAFEB7" box="[755,945,260,284]" pageId="8" pageNumber="509" value="2013-08-30">30 August 2013</date>
|
||
, TAD1354 (GPS data same as TAD1333) (
|
||
<collectionCode id="A1DFF5AF686A9F69FF09FE8F1AEEFEEB" box="[158,245,296,320]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="509">ZMHU</collectionCode>
|
||
,
|
||
<collectionCode id="A1DFF5AF686A9F69FE94FE8F1B7FFEEB" box="[259,356,296,320]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="509">NHMW</collectionCode>
|
||
);
|
||
<specimenCount id="D1C8A6E3686A9F69FEEDFE8F1BCDFEEB" box="[378,470,295,320]" pageId="8" pageNumber="509" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="D1C8A6E3686A9F69FE73FE8F1853FEEB" box="[484,584,295,320]" pageId="8" pageNumber="509" type="female">1 female</specimenCount>
|
||
(apterous), Xuan Son N’Park, Lap stream, site 1, at Ngoc waterfall, coll. Nguyen T.S.
|
||
<emphasis id="F5BAB178686A9F69FEA0FEEA1B6DFECF" box="[311,374,332,356]" italics="true" pageId="8" pageNumber="509">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA686A9F69FE1DFEEB1856FECF" box="[394,589,332,356]" pageId="8" pageNumber="509" value="2014-08-27">27 August 2014</date>
|
||
, TS1409 (GPS data same as TAD1331) (
|
||
<collectionCode id="A1DFF5AF686A9F69FBD2FEEB1E85FECF" box="[1093,1182,332,356]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="509">ZMHU</collectionCode>
|
||
);
|
||
<specimenCount id="D1C8A6E3686A9F69FB2FFEEB1F03FECE" box="[1208,1304,332,357]" pageId="8" pageNumber="509" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="D1C8A6E3686A9F69FABEFEEB1F87FECE" box="[1321,1436,332,357]" pageId="8" pageNumber="509" type="female">2 females</specimenCount>
|
||
(apterous), Xuan Son N’Park, Lap stream, site 2, first concrete bridge from Ngoc waterfall, coll. Nguyen T.S.
|
||
<emphasis id="F5BAB178686A9F69FACAFED61F8EFE23" box="[1373,1429,368,392]" italics="true" pageId="8" pageNumber="509">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA686A9F69FF00FE331B56FE07" box="[151,333,404,428]" pageId="8" pageNumber="509" value="2014-08-27">27 August 2014</date>
|
||
, TS1410 (GPS data same as TAD1332) (
|
||
<collectionCode id="A1DFF5AF686A9F69FC81FE331974FE07" box="[790,879,404,428]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="509">ZMHU</collectionCode>
|
||
);
|
||
<specimenCount id="D1C8A6E3686A9F69FC15FE3319CAFE06" box="[898,977,404,429]" pageId="8" pageNumber="509" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="D1C8A6E3686A9F69FC4BFE331E53FE06" box="[988,1096,404,429]" pageId="8" pageNumber="509" type="female">2 females</specimenCount>
|
||
(apterous), Xuan Son N’Park, Lap stream, site 2, first concrete bridge from Ngoc waterfall, coll. Nguyen T.S.
|
||
<emphasis id="F5BAB178686A9F69FB82FE1E1E56FE7B" box="[1045,1101,440,464]" italics="true" pageId="8" pageNumber="509">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA686A9F69FBCBFE1F1EEFFE7B" box="[1116,1268,439,464]" pageId="8" pageNumber="509" value="2015-05-23">23 May 2015</date>
|
||
, TS1512 (GPS data same as TAD1332) (
|
||
<collectionCode id="A1DFF5AF686A9F69FE2FFE7B180AFE5F" box="[440,529,476,500]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="509">ZMHU</collectionCode>
|
||
);
|
||
<specimenCount id="D1C8A6E3686A9F69FDB2FE7B1867FE5E" box="[549,636,476,501]" pageId="8" pageNumber="509" type="male">2 males</specimenCount>
|
||
(apterous), Xuan Son N’Park, Lap stream, site 2, first concrete bridge from Ngoc waterfall, coll. Nguyen T.S.
|
||
<emphasis id="F5BAB178686A9F69FDC3FDA61896FDB3" box="[596,653,512,536]" italics="true" pageId="8" pageNumber="509">et al.</emphasis>
|
||
,
|
||
<date id="B3704BAA686A9F69FD0CFE58194BFDB3" box="[667,848,511,536]" pageId="8" pageNumber="509" value="2015-08-26">26 August 2015</date>
|
||
, TS8512 (GPS data same as TAD1332) (
|
||
<collectionCode id="A1DFF5AF686A9F69FA8CFDA71F6FFDB3" box="[1307,1396,512,536]" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="509">ZMHU</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8FD43EE1686A9F68FF50FD831804F805" lastPageId="9" lastPageNumber="510" pageId="8" pageNumber="509" type="description">
|
||
<paragraph id="C7716D6A686A9F69FF50FD831BB9FD2F" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
|
||
<emphasis id="F5BAB178686A9F69FF50FD831B42FD96" bold="true" box="[199,345,548,573]" pageId="8" pageNumber="509">Description.</emphasis>
|
||
Size: apterous morph: males, length 6.90–7.10 (allotype 6.95), width 2.20–2.30 (allotype 2.24); females, length 8.7–9.4 (
|
||
<typeStatus id="1875D3C8686A9F69FE3AFDE0180AFDCB" box="[429,529,583,608]" pageId="8" pageNumber="509" type="holotype">holotype</typeStatus>
|
||
9.0), width 3.08–3.20 (
|
||
<typeStatus id="1875D3C8686A9F69FC8DFDE01965FDCB" box="[794,894,583,608]" pageId="8" pageNumber="509" type="holotype">holotype</typeStatus>
|
||
3.20); macropterous morph (de-alated): female, length 9.10, width 3.23.
|
||
</paragraph>
|
||
<paragraph id="C7716D6A686A9F69FF50FD281905FB9E" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
|
||
Colour (
|
||
<figureCitation id="5FF571EF686A9F69FEB0FD281BAEFD03" box="[295,437,655,680]" captionStart="FIGURES 24 – 26" captionStartId="9.[151,264,1633,1656]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@9.[151,1436,193,1612]" captionTargetPageId="9" captionText="FIGURES 24 – 26. (24) Habitat of type locality of Rhyacobates constrictus sp. n. (25) A copulating pair of R. constrictus sp. n., skating on the fast current of the stream. (26) A copulating pair of R. constrictus sp. n., resting on the emerged rocks in the middle of the stream (photographed by Tran Anh Duc)." httpUri="https://zenodo.org/record/271688/files/figure.png" pageId="8" pageNumber="509">Figs. 25, 26</figureCitation>
|
||
): Dorsum of body mainly black, covered with silvery pubescence, especially dense on thoracic pleura, metacoxa and abdomen. Dorsum of head yellow with median, longitudinal brown mark on anterior three quarters. Antennae mainly dark brown, segment 4 with whitish area in distal two-fifths. Pronotum mainly black with median yellow mark (in female, width of mark about one third of pronotum mark; in male, yellow mark narrower), anterolateral corners of pronotum yellow. Mesonotum and metanotum black, without yellow mark. Abdominal terga chiefly black, except abdominal apex yellowish and brown (in proctiger and pygophore of male; and in abdominal segment 7 and genitalia of female). In both sexes, venter of body covered with dense silvery pubescence. Venter of female mostly yellow, except mesosternum, mesopleuron and anterior third of metasternum black. Venter of male with mesosternum, mesopleuron and metasternum black, abdominal sternum 2 black or brown, the remaining parts yellow. All acetabulae, coxae and trochanters yellow. Fore femur yellow with brown longitudinal stripe dorsally. Middle and hind femora yellow at basal part, dark brown at distal part. Fore tibia and tarsus dark brown; other tibiae and tarsi yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="C7716D6A686A9F69FF50FBE71F3AF986" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
|
||
Apterous female (
|
||
<typeStatus id="1875D3C8686A9F69FE05FB981BE0FBF3" box="[402,507,1087,1112]" pageId="8" pageNumber="509" type="holotype">holotype</typeStatus>
|
||
): Head width 1.43, interocular width 0.57, eye length (dorsal view) 0.63. Lengths of antennal segments 1–4: 4.30: 1.21: 1.50: 1.09. Pronotum broader than long, length 0.81, width 1.53; mesonotum length 2.56, metanotum length 1.07. Mesopleuron with scattered long erect bristle-like brown hairs. All coxae with rings of dark bristle-like hairs at apical margins. Fore trochanter with long hairs on ventral side. Lengths of leg segments (femur: tibia: tarsus 1: tarsus 2): fore leg: 4.35: 3.55: 2.00: 0.94; middle leg: 11.50: 6.70: 4.10: 0.44; hind leg: 11.80: 5.19: 0.14: 0.22. Fore femur slender (width 0.42), venter of femur with row of 11 dark bristle-like hairs, with scattered soft long hairs, sub-apical part with small tooth-like elevation, sometimes indistinct (
|
||
<figureCitation id="5FF571EF686A9F69FAA1FAB01F8BFA9B" box="[1334,1424,1303,1328]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Fig. 27</figureCitation>
|
||
). Abdomen relatively short, ventral length 3.85; abdominal apex directed dorsocaudad (
|
||
<figureCitation id="5FF571EF686A9F69FB27FA9B1F16FAFF" box="[1200,1293,1340,1365]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Fig. 34</figureCitation>
|
||
). Tergum 1 posteromedially swollen (
|
||
<figureCitation id="5FF571EF686A9F69FE5EFAF8182AFAD3" box="[457,561,1375,1400]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Figs. 36</figureCitation>
|
||
). Connexival margin of segments 1–5 narrow, of segment 6 thickened. Abdominal segment 7 prolonged, completely enclosing genital segments, tergum 7 short (length 0.45), posterior part thickened, medially notched on posterior margin; sternum 7 (length 1.79) about 4.0 times as long as tergum 7; connexival and posterolateral margins expanded dorsad, folded inwards, meeting in middle of abdominal dorsum (
|
||
<typeStatus id="1875D3C8686A9F69FF09FA481B1FF9A3" box="[158,260,1519,1544]" pageId="8" pageNumber="509" type="holotype">holotype</typeStatus>
|
||
,
|
||
<figureCitation id="5FF571EF686A9F69FE87FA481B7AF9A3" box="[272,353,1519,1544]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Fig. 36</figureCitation>
|
||
) or overlapping each other (
|
||
<figureCitation id="5FF571EF686A9F69FD09FA4818EAF9A3" box="[670,753,1519,1544]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Fig. 38</figureCitation>
|
||
), thus covering most of genital segments; sternum 7 tapering caudad, posterior margin with narrow median process, apex of process narrowly rounded (
|
||
<figureCitation id="5FF571EF686A9F69FB1CF9B31F0EF986" box="[1163,1301,1556,1581]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Figs. 36–39</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C7716D6A686A9F69FF50F99F1EE4F877" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="509">
|
||
Apterous male (allotype): Head width 1.32, interocular width 0.49, eye length 0.65. Lengths of antennal segments 1–4: 3.65: 1.10: 1.50: 1.06. Pronotum broader than long, length 0.72, width 1.41; mesonotum length 2.13, metanotum length 0.79. Mesopleuron with scattered long erect bristle-like brown hairs. Lengths of leg segments: fore leg: 3.90: 3.20: 0.98: 0.69; middle leg: 10.30: 5.81: 3.32: 0.40; hind leg: 10.40: 4.00: 0.12: 0.18. Fore femur slender (width 0.45), with row of seven bristle-like hairs and few scattered soft long hairs on venter, sub-apical part with small tooth-like elevation. Middle trochanter without spines, middle femur with scattered small spines, not arranged in distinct row (
|
||
<figureCitation id="5FF571EF686A9F69FD18F8A818FCF883" box="[655,743,1807,1832]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Fig. 28</figureCitation>
|
||
). Other leg structures similar to those of female. Abdomen relatively short, ventral length 2.15. Length of sternum 7: 0.30, posterior margin slightly concave. Genitalia: slightly directed downwards; venter of abdominal segment 8 length (in-situ) 0.15; proctiger laterally widened forming broad, angular lobe on each side (
|
||
<figureCitation id="5FF571EF686A9F69FDE1F8DB18D0F83F" box="[630,715,1916,1941]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Fig. 29</figureCitation>
|
||
); pygophore of moderate size, apical margin rounded; paramere relatively long, curved at basal fourth, strongly thickened at middle, then tapering towards narrow apex, apex slightly directed inwards, setae mostly on distal part (
|
||
<figureCitation id="5FF571EF686A9F69FD79F863196CF877" box="[750,887,1988,2013]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Figs. 30–32</figureCitation>
|
||
); endosomal sclerites as in
|
||
<figureCitation id="5FF571EF686A9F69FB3EF8631EE0F877" box="[1193,1275,1988,2013]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="8" pageNumber="509">Fig. 33</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="93B13DE2686B9F68FF00F9C618D3F91F" httpUri="https://zenodo.org/record/271688/files/figure.png" pageId="9" pageNumber="510" targetBox="[151,1436,193,1610]" targetPageId="9">
|
||
<paragraph id="C7716D6A686B9F68FF00F9C618D3F91F" blockId="9.[151,1436,1633,1716]" pageId="9" pageNumber="510">
|
||
<emphasis id="F5BAB178686B9F68FF00F9C61B4FF9DD" bold="true" box="[151,340,1633,1656]" pageId="9" pageNumber="510">FIGURES 24–26.</emphasis>
|
||
(24) Habitat of type locality of
|
||
<taxonomicName id="00CE16E9686B9F68FD00F9C61995F9DC" box="[663,910,1633,1655]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB178686B9F68FD00F9C61995F9DC" box="[663,910,1633,1655]" italics="true" pageId="9" pageNumber="510">Rhyacobates constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C03686B9F68FC03F9C519DCF9DC" box="[916,967,1634,1655]" pageId="9" pageNumber="510" rank="species">
|
||
<emphasis id="F5BAB178686B9F68FC03F9C519DCF9DC" bold="true" box="[916,967,1634,1655]" pageId="9" pageNumber="510">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(25) A copulating pair of
|
||
<taxonomicName id="00CE16E9686B9F68FB46F9C51F47F9DC" box="[1233,1372,1634,1655]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB178686B9F68FB46F9C51F47F9DC" box="[1233,1372,1634,1655]" italics="true" pageId="9" pageNumber="510">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C03686B9F68FAF4F9C51F8DF9DC" box="[1379,1430,1634,1655]" pageId="9" pageNumber="510" rank="species">
|
||
<emphasis id="F5BAB178686B9F68FAF4F9C51F8DF9DC" bold="true" box="[1379,1430,1634,1655]" pageId="9" pageNumber="510">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, skating on the fast current of the stream. (26) A copulating pair of
|
||
<taxonomicName id="00CE16E9686B9F68FCC3F92619FAF93D" box="[852,993,1665,1686]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB178686B9F68FCC3F92619FAF93D" box="[852,993,1665,1686]" italics="true" pageId="9" pageNumber="510">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C03686B9F68FC7DF9271E06F93E" box="[1002,1053,1664,1685]" pageId="9" pageNumber="510" rank="species">
|
||
<emphasis id="F5BAB178686B9F68FC7DF9271E06F93E" bold="true" box="[1002,1053,1664,1685]" pageId="9" pageNumber="510">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, resting on the emerged rocks in the middle of the stream (photographed by Tran Anh Duc).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C7716D6A686B9F68FF50F9461B47F821" blockId="9.[151,1437,1761,2038]" pageId="9" pageNumber="510">De-alated macropterous female: Head width 1.46, interocular width 0.56, eye length 0.67. Lengths of antennal segments 1–4: 4.25: 1.17: 1.42: 1.08. Pronotum covering most of mesonotum, length 3.35, width (across humeri) 2.56; metanotum length 0.83. Lengths of leg segments: fore leg: 4.35: 3.60: 1.91: 0.95; middle leg: 11.50: 6.80: 4.15: 0.46; hind leg: 12.10: 5.30: 0.14: 0.23; width of fore femur 0.44. Other structural characteristics similar to apterous females.</paragraph>
|
||
<paragraph id="C7716D6A686B9F68FF50F8311804F805" blockId="9.[151,1437,1761,2038]" box="[199,543,1941,1966]" pageId="9" pageNumber="510">Macropterous male: unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8FD43EE1686B9F68FF50F81E1889F85D" pageId="9" pageNumber="510" type="etymology">
|
||
<paragraph id="C7716D6A686B9F68FF50F81E1889F85D" blockId="9.[151,1437,1761,2038]" pageId="9" pageNumber="510">
|
||
<emphasis id="F5BAB178686B9F68FF50F81E1B55F879" bold="true" box="[199,334,1977,2002]" pageId="9" pageNumber="510">Etymology.</emphasis>
|
||
The species epithet
|
||
<taxonomicName id="00CE16E9686B9F68FDABF81D18ACF87A" box="[572,695,1978,2001]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="510" phylum="Arthropoda" rank="species" species="constrictus">
|
||
<emphasis id="F5BAB178686B9F68FDABF81D18ACF87A" box="[572,695,1978,2001]" italics="true" pageId="9" pageNumber="510">constrictus</emphasis>
|
||
</taxonomicName>
|
||
refers to the posteriorly constricted abdominal sternum 7 of the female, which holds genital segments within.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="93B13DE268689F6BFF00F8AE1B01F831" httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="10" pageNumber="511" targetBox="[168,1417,201,1747]" targetPageId="10">
|
||
<paragraph id="C7716D6A68689F6BFF00F8AE1B01F831" blockId="10.[151,1436,1801,1946]" pageId="10" pageNumber="511">
|
||
<emphasis id="F5BAB17868689F6BFF00F8AE1B4DF8B4" bold="true" box="[151,342,1801,1824]" pageId="10" pageNumber="511">FIGURES 27–39.</emphasis>
|
||
Morphological features of
|
||
<taxonomicName id="00CE16E968689F6BFDE3F8AD1976F88B" box="[628,877,1802,1824]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB17868689F6BFDE3F8AD1976F88B" box="[628,877,1802,1824]" italics="true" pageId="10" pageNumber="511">Rhyacobates constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C0368689F6BFCE1F8AD19B2F8B4" box="[886,937,1802,1823]" pageId="10" pageNumber="511" rank="species">
|
||
<emphasis id="F5BAB17868689F6BFCE1F8AD19B2F8B4" bold="true" box="[886,937,1802,1823]" pageId="10" pageNumber="511">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30–32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="8FD43EE168699F6AFF50FF301BCBFC93" pageId="11" pageNumber="512" type="discussion">
|
||
<paragraph id="C7716D6A68699F6AFF50FF30187CFD0C" blockId="11.[151,1436,151,861]" pageId="11" pageNumber="512">
|
||
<emphasis id="F5BAB17868699F6AFF50FF301B20FF1B" bold="true" box="[199,315,151,176]" pageId="11" pageNumber="512">Remarks.</emphasis>
|
||
<taxonomicName id="00CE16E968699F6AFED3FF3F1843FF04" box="[324,600,152,176]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB17868699F6AFED3FF3F1843FF04" box="[324,600,152,176]" italics="true" pageId="11" pageNumber="512">Rhyacobates constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C0368699F6AFDF5FF301882FF1B" box="[610,665,151,176]" pageId="11" pageNumber="512" rank="species">
|
||
<emphasis id="F5BAB17868699F6AFDF5FF301882FF1B" bold="true" box="[610,665,151,176]" pageId="11" pageNumber="512">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be easily separated from all of its congeners by the following characteristics: in both sexes, the meso- and metanota are totally black, without yellow marking (
|
||
<figureCitation id="5FF571EF68699F6AFB78FF1B1F5EFF7E" box="[1263,1349,188,213]" captionStart="FIGURES 24 – 26" captionStartId="9.[151,264,1633,1656]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@9.[151,1436,193,1612]" captionTargetPageId="9" captionText="FIGURES 24 – 26. (24) Habitat of type locality of Rhyacobates constrictus sp. n. (25) A copulating pair of R. constrictus sp. n., skating on the fast current of the stream. (26) A copulating pair of R. constrictus sp. n., resting on the emerged rocks in the middle of the stream (photographed by Tran Anh Duc)." httpUri="https://zenodo.org/record/271688/files/figure.png" pageId="11" pageNumber="512">Fig. 26</figureCitation>
|
||
); in the female, the abdomen is curved dorsad (
|
||
<figureCitation id="5FF571EF68699F6AFDCAFF7818A8FF53" box="[605,691,223,248]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="11" pageNumber="512">Fig. 34</figureCitation>
|
||
), the connexival and posterolateral margins of sternum 7 expand dorsad and fold inwards; thus, sternum 7 covers most of the genital segments (
|
||
<figureCitation id="5FF571EF68699F6AFBA0FEA31EDFFEB6" box="[1079,1220,260,285]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="11" pageNumber="512">Figs. 35–39</figureCitation>
|
||
), sternum 7 tapers caudad, with a narrow median process on the posterior margin, the apex of the process is narrowly rounded (
|
||
<figureCitation id="5FF571EF68699F6AFAF3FE801AFAFECE" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="11" pageNumber="512">Figs. 36–39</figureCitation>
|
||
); the male paramere is strongly thickened at the middle (
|
||
<figureCitation id="5FF571EF68699F6AFCF2FEEB19EBFECF" box="[869,1008,332,357]" captionStart="FIGURES 27 – 39" captionStartId="10.[151,264,1801,1824]" captionTargetBox="[168,1417,201,1747]" captionTargetId="figure@10.[151,1436,193,1781]" captionTargetPageId="10" captionText="FIGURES 27 – 39. Morphological features of Rhyacobates constrictus sp. n. (27) right fore leg of female. (28) basal part of right middle leg of male, ventral view. (29) proctiger. (30 – 32) left paramere from three different views. (33) endosomal sclerites. (34) body of female, lateral view. (35) apex of abdomen of female, lateral view. (36) abdomen of female holotype, dorsal view. (37) apex of abdomen of female holotype, ventral view. (38, 39) apex of abdomen of female paratype, dorsal and ventral view." httpUri="https://zenodo.org/record/271689/files/figure.png" pageId="11" pageNumber="512">Figs. 30, 32</figureCitation>
|
||
). This new species is the only species of
|
||
<taxonomicName id="00CE16E968699F6AFF20FED71B5DFE23" box="[183,326,368,392]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F5BAB17868699F6AFF20FED71B5DFE23" box="[183,326,368,392]" italics="true" pageId="11" pageNumber="512">Rhyacobates</emphasis>
|
||
</taxonomicName>
|
||
that has no yellow marking on the meso- and metanota (in other species of
|
||
<taxonomicName id="00CE16E968699F6AFB3AFED71F27FE23" box="[1197,1340,368,392]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F5BAB17868699F6AFB3AFED71F27FE23" box="[1197,1340,368,392]" italics="true" pageId="11" pageNumber="512">Rhyacobates</emphasis>
|
||
</taxonomicName>
|
||
, at least the mesonotum has yellow marking). Compared with other species of
|
||
<taxonomicName id="00CE16E968699F6AFC5AFE331E47FE07" box="[973,1116,404,428]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F5BAB17868699F6AFC5AFE331E47FE07" box="[973,1116,404,428]" italics="true" pageId="11" pageNumber="512">Rhyacobates</emphasis>
|
||
</taxonomicName>
|
||
, the shape and structure of sternum 7 of the female is unique for
|
||
<taxonomicName id="00CE16E968699F6AFDF1FE1E191DFE64" box="[614,774,440,464]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB17868699F6AFDF1FE1E191DFE64" box="[614,774,440,464]" italics="true" pageId="11" pageNumber="512">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C0368699F6AFC85FE101952FE7B" box="[786,841,439,464]" pageId="11" pageNumber="512" rank="species">
|
||
<emphasis id="F5BAB17868699F6AFC85FE101952FE7B" bold="true" box="[786,841,439,464]" pageId="11" pageNumber="512">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
The median process on sternum 7 is found in
|
||
<taxonomicName id="00CE16E968699F6AFA13FE1E1821FE5E" authority="Andersen & Chen, 1995" authorityName="Andersen & Chen" authorityYear="1995" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="abdominalis">
|
||
<emphasis id="F5BAB17868699F6AFA13FE1E1B39FE5F" italics="true" pageId="11" pageNumber="512">R. abdominalis</emphasis>
|
||
<bibRefCitation id="A35F109B68699F6AFEBEFE7B1821FE5E" author="Andersen" box="[297,570,476,501]" pageId="11" pageNumber="512" refString="Andersen, N. M. & Chen, P. - P. (1995) A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie, 138, 51 - 67. Chen, P-P., Nieser, N. & Zettel, H. (2005) The aquatic and semi-aquatic bugs (Heteroptera: Nepomorpha & Gerromorpha) of Malesia. Fauna Malesiana Handbooks 5. Brill, Leiden, / Boston, 546 pp." type="book chapter" year="1995">Andersen & Chen, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="00CE16E968699F6AFDE7FE7A1E10FE5E" authority="Andersen & Chen, 1995" authorityName="Andersen & Chen" authorityYear="1995" box="[624,1035,476,501]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="recurvus">
|
||
<emphasis id="F5BAB17868699F6AFDE7FE7A18EAFE5F" box="[624,753,477,500]" italics="true" pageId="11" pageNumber="512">R. recurvus</emphasis>
|
||
<bibRefCitation id="A35F109B68699F6AFD6FFE7B1E10FE5E" author="Andersen" box="[760,1035,476,501]" pageId="11" pageNumber="512" refString="Andersen, N. M. & Chen, P. - P. (1995) A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie, 138, 51 - 67. Chen, P-P., Nieser, N. & Zettel, H. (2005) The aquatic and semi-aquatic bugs (Heteroptera: Nepomorpha & Gerromorpha) of Malesia. Fauna Malesiana Handbooks 5. Brill, Leiden, / Boston, 546 pp." type="book chapter" year="1995">Andersen & Chen, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
, but they are not as prominent as in
|
||
<taxonomicName id="00CE16E968699F6AFF00FDA61B29FDBC" box="[151,306,512,536]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB17868699F6AFF00FDA61B29FDBC" box="[151,306,512,536]" italics="true" pageId="11" pageNumber="512">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C0368699F6AFEADFE581B6AFDB3" box="[314,369,511,536]" pageId="11" pageNumber="512" rank="species">
|
||
<emphasis id="F5BAB17868699F6AFEADFE581B6AFDB3" bold="true" box="[314,369,511,536]" pageId="11" pageNumber="512">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; and the connexival margin of sternum 7 without a process also separates
|
||
<taxonomicName id="00CE16E968699F6AFB56FDA61F47FDBC" box="[1217,1372,512,536]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB17868699F6AFB56FDA61F47FDBC" box="[1217,1372,512,536]" italics="true" pageId="11" pageNumber="512">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C0368699F6AFAF3FE581F80FDB3" box="[1380,1435,511,536]" pageId="11" pageNumber="512" rank="species">
|
||
<emphasis id="F5BAB17868699F6AFAF3FE581F80FDB3" bold="true" box="[1380,1435,511,536]" pageId="11" pageNumber="512">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from these two taxa. Males of
|
||
<taxonomicName id="00CE16E968699F6AFE64FD821894FD97" box="[499,655,549,572]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB17868699F6AFE64FD821894FD97" box="[499,655,549,572]" italics="true" pageId="11" pageNumber="512">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C0368699F6AFD01FD8318D5FD96" box="[662,718,548,573]" pageId="11" pageNumber="512" rank="species">
|
||
<emphasis id="F5BAB17868699F6AFD01FD8318D5FD96" bold="true" box="[662,718,548,573]" pageId="11" pageNumber="512">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be easily separated from
|
||
<taxonomicName id="00CE16E968699F6AFBB5FD821ED5FD97" box="[1058,1230,548,572]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="abdominalis">
|
||
<emphasis id="F5BAB17868699F6AFBB5FD821ED5FD97" box="[1058,1230,548,572]" italics="true" pageId="11" pageNumber="512">R. abdominalis</emphasis>
|
||
</taxonomicName>
|
||
by the absence of yellow marking on the meso- and metanota and the shape of the paramere. To date, only the female of
|
||
<taxonomicName id="00CE16E968699F6AFA8BFDEE1F87FDCB" box="[1308,1436,585,608]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="recurvus">
|
||
<emphasis id="F5BAB17868699F6AFA8BFDEE1F87FDCB" box="[1308,1436,585,608]" italics="true" pageId="11" pageNumber="512">R. recurvus</emphasis>
|
||
</taxonomicName>
|
||
has been described; the description of the male is not available. Thus, we are unable to compare the male of
|
||
<taxonomicName id="00CE16E968699F6AFA13FDCA1B09FD0C" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB17868699F6AFA13FDCA1B09FD0C" italics="true" pageId="11" pageNumber="512">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C0368699F6AFE8EFD281B4BFD03" box="[281,336,655,680]" pageId="11" pageNumber="512" rank="species">
|
||
<emphasis id="F5BAB17868699F6AFE8EFD281B4BFD03" bold="true" box="[281,336,655,680]" pageId="11" pageNumber="512">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
with that of
|
||
<taxonomicName id="00CE16E968699F6AFE77FD36187BFD03" box="[480,608,657,680]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="recurvus">
|
||
<emphasis id="F5BAB17868699F6AFE77FD36187BFD03" box="[480,608,657,680]" italics="true" pageId="11" pageNumber="512">R. recurvus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C7716D6A68699F6AFF50FD131BCBFC93" blockId="11.[151,1436,151,861]" pageId="11" pageNumber="512">
|
||
<emphasis id="F5BAB17868699F6AFF50FD131B2EFD66" bold="true" box="[199,309,692,717]" pageId="11" pageNumber="512">Habitats.</emphasis>
|
||
Specimens of
|
||
<taxonomicName id="00CE16E968699F6AFE70FD12189DFD67" box="[487,646,693,716]" class="Insecta" family="Gerridae" genus="Rhyacobates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="512" phylum="Arthropoda" rank="species" species="constrictus" status="sp. nov.">
|
||
<emphasis id="F5BAB17868699F6AFE70FD12189DFD67" box="[487,646,693,716]" italics="true" pageId="11" pageNumber="512">R. constrictus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="EE890C0368699F6AFD06FD1318D2FD66" box="[657,713,692,717]" pageId="11" pageNumber="512" rank="species">
|
||
<emphasis id="F5BAB17868699F6AFD06FD1318D2FD66" bold="true" box="[657,713,692,717]" pageId="11" pageNumber="512">sp.n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
were collected at fast flowing sections of streams shaded by pristine forest at elevation above
|
||
<quantity id="0036C08F68699F6AFD8EFD701878FD44" box="[537,611,727,752]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="512" unit="m" value="250.0">250 m</quantity>
|
||
(
|
||
<figureCitation id="5FF571EF68699F6AFDE2FD701918FD5B" box="[629,771,727,752]" captionStart="FIGURES 24 – 26" captionStartId="9.[151,264,1633,1656]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@9.[151,1436,193,1612]" captionTargetPageId="9" captionText="FIGURES 24 – 26. (24) Habitat of type locality of Rhyacobates constrictus sp. n. (25) A copulating pair of R. constrictus sp. n., skating on the fast current of the stream. (26) A copulating pair of R. constrictus sp. n., resting on the emerged rocks in the middle of the stream (photographed by Tran Anh Duc)." httpUri="https://zenodo.org/record/271688/files/figure.png" pageId="11" pageNumber="512">Figs. 24, 25</figureCitation>
|
||
). They were found skating along streams with adults and nymphs in a mixture. Individuals of this species were also found often resting on partially submerged rocks in the middle the stream (
|
||
<figureCitation id="5FF571EF68699F6AFEF8FCB81BD8FC93" box="[367,451,799,824]" captionStart="FIGURES 24 – 26" captionStartId="9.[151,264,1633,1656]" captionTargetBox="[151,1436,193,1610]" captionTargetId="figure@9.[151,1436,193,1612]" captionTargetPageId="9" captionText="FIGURES 24 – 26. (24) Habitat of type locality of Rhyacobates constrictus sp. n. (25) A copulating pair of R. constrictus sp. n., skating on the fast current of the stream. (26) A copulating pair of R. constrictus sp. n., resting on the emerged rocks in the middle of the stream (photographed by Tran Anh Duc)." httpUri="https://zenodo.org/record/271688/files/figure.png" pageId="11" pageNumber="512">Fig. 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8FD43EE168699F6AFF50FCE31827FCF6" box="[199,572,836,861]" pageId="11" pageNumber="512" type="distribution">
|
||
<paragraph id="C7716D6A68699F6AFF50FCE31827FCF6" blockId="11.[151,1436,151,861]" box="[199,572,836,861]" pageId="11" pageNumber="512">
|
||
<emphasis id="F5BAB17868699F6AFF50FCE31B7BFCF6" bold="true" box="[199,352,836,861]" pageId="11" pageNumber="512">Distribution.</emphasis>
|
||
<collectingCountry id="BFD92DFA68699F6AFEF0FCE31BD1FCF6" box="[359,458,836,861]" name="Vietnam" pageId="11" pageNumber="512">Vietnam</collectingCountry>
|
||
: Phu Tho.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |