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<document id="5F3EB945100EC4D5CE8F21C1B80F4906" ID-CLB-Dataset="29418" ID-DOI="10.11646/zootaxa.4482.3.2" ID-GBIF-Dataset="fd24ef18-76ac-46f1-aec2-2faf5b0ff325" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440707" ID-ZooBank="3FBE11E2-9F97-4A29-92A9-67780C782E0D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538375301724" checkinUser="plazi" docAuthor="Li, Jheng-Jhang, Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2018" docId="376A1B4DFFBDFF9DD1CDFA1944CBFD8A" docLanguage="en" docName="zootaxa.4482.3.2.pdf" docOrigin="Zootaxa 4482 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parasesarma tarantula Li &amp; Rahayu &amp; Ng 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="478" masterDocId="CB536335FFA4FF86D15AFFF74432FFDA" masterDocTitle="Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific" masterLastPageNumber="490" masterPageNumber="451" pageNumber="476" updateTime="1698559507815" updateUser="ExternalLinkService">
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<mods:title id="2CA198D756950B93E15750FF819E985A">Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific</mods:title>
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<mods:namePart id="29D9EEB3BADF294833A9DDC0E231AD59">Li, Jheng-Jhang</mods:namePart>
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<mods:namePart id="A39860532EAF0CB575D0051E179A02B9">Rahayu, Dwi Listyo</mods:namePart>
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<mods:namePart id="A2E239BDE1A3DE50D696DD7771B63A4A">Ng, Peter K. L.</mods:namePart>
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<mods:title id="DA8D86915100A8EF2E6C8EE2E5C7086E">Zootaxa</mods:title>
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<mods:date id="4598653705BE13640D2B3D95959FB67A">2018</mods:date>
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<mods:number id="81A84D200120EEBBD676DE75A59AE627">2018-09-19</mods:number>
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<mods:number id="E2C0F4F027AF62D78F94E61DF7C403B0">4482</mods:number>
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<treatment id="376A1B4DFFBDFF9DD1CDFA1944CBFD8A" ID-DOI="http://doi.org/10.5281/zenodo.5986799" ID-GBIF-Taxon="148421720" ID-Zenodo-Dep="5986799" LSID="urn:lsid:plazi:treatment:376A1B4DFFBDFF9DD1CDFA1944CBFD8A" httpUri="http://treatment.plazi.org/id/376A1B4DFFBDFF9DD1CDFA1944CBFD8A" lastPageId="27" lastPageNumber="478" pageId="25" pageNumber="476">
<subSubSection id="F7D9F9D0FFBDFF9FD1CDFA1946A7F9F0" pageId="25" pageNumber="476" type="nomenclature">
<paragraph id="BF7CAA5BFFBDFF9FD1CDFA1945C4F9D2" blockId="25.[151,661,1518,1578]" box="[151,502,1518,1544]" pageId="25" pageNumber="476">
<heading id="E4341D37FFBDFF9FD1CDFA1945C4F9D2" bold="true" box="[151,502,1518,1544]" fontSize="11" level="1" pageId="25" pageNumber="476" reason="1">
<emphasis id="8DB77649FFBDFF9FD1CDFA1945C4F9D2" bold="true" box="[151,502,1518,1544]" pageId="25" pageNumber="476">
<taxonomicName id="78C3D1D8FFBDFF9FD1CDFA19459FF9D2" ID-CoL="84G4H" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[151,429,1518,1544]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="tarantula" status="sp. nov.">
<emphasis id="8DB77649FFBDFF9FD1CDFA19459FF9D2" bold="true" box="[151,429,1518,1544]" italics="true" pageId="25" pageNumber="476">Parasesarma tarantula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFBDFF9FD0E9FA1945C4F9D2" box="[435,502,1518,1544]" pageId="25" pageNumber="476" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="BF7CAA5BFFBDFF9FD1CDF9E646A7F9F0" blockId="25.[151,661,1518,1578]" box="[151,661,1553,1578]" pageId="25" pageNumber="476">
(
<figureCitation id="27F8B6DEFFBDFF9FD1C5F9E644CEF9F0" box="[159,252,1553,1578]" captionStart="FIGURE 17" captionStartId="26.[151,250,1781,1803]" captionTargetBox="[152,1435,449,1769]" captionTargetId="figure@26.[152,1435,449,1769]" captionTargetPageId="26" captionText="FIGURE 17. Parasesarma tarantula n. sp.. AE, paratype male (12.6×10.9 mm) (ZRC 2018.0801), Sulawesi, Indonesia; F, G, paratype female (11.2×9.6 mm) (ZRC 2018.0802), same locality. A, F, dorsal views; B, E, G, ventral views; C, D, frontal views; E, close-up of gonopods." httpUri="https://zenodo.org/record/1440743/files/figure.png" pageId="25" pageNumber="476">Figs. 17</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD052F9E54514F9F0" box="[264,294,1554,1578]" captionStart="FIGURE 18" captionStartId="27.[151,250,1907,1929]" captionTargetBox="[192,1347,669,1874]" captionTargetId="figure@27.[160,1426,637,1891]" captionTargetPageId="27" captionText="FIGURE 18. Parasesarma tarantula n. sp., holotype male (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia. A, dorsal view of carapace; BD, tip of left cheliped; EH, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440745/files/figure.png" pageId="25" pageNumber="476">18</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD068F9E54557F9F0" box="[306,357,1554,1578]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="25" pageNumber="476">19H</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD02BF9E54593F9F0" box="[369,417,1554,1578]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="25" pageNumber="476">20G</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD0F6F9E545ECF9F0" box="[428,478,1554,1578]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="25" pageNumber="476">21H</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD0B0F9E5462BF9F0" box="[490,537,1554,1578]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="25" pageNumber="476">22E</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD37FF9E54667F9F0" box="[549,597,1554,1578]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="25" pageNumber="476">23E</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD33AF9E546BFF9F0" box="[608,653,1554,1578]" captionStart="FIGURE 24" captionStartId="34.[151,250,1531,1553]" captionTargetBox="[151,1436,218,1490]" captionTargetId="figure@34.[151,1436,218,1490]" captionTargetPageId="34" captionText="FIGURE 24. In situ photographs of Parasesarma leptosoma species-complex, specimens not collected. A, P. leptosoma (Hilgendorf, 1869), Kenya (Photograph: S. Cannicci); B, P. gecko n. sp., Okinawa, Japan; C, P. macaco n. sp., Pingtung, Taiwan; D, P. kui n. sp., Pingtung, Taiwan; E, P. purpureum n. sp., Borneo, Malaysia; F, P. tarantula n. sp., Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440757/files/figure.png" pageId="25" pageNumber="476">24F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFBDFF9FD1CDF9AE404EF904" pageId="25" pageNumber="476" type="materials_examined">
<paragraph id="BF7CAA5BFFBDFF9FD1CDF9AE404EF904" blockId="25.[151,1436,1625,2027]" pageId="25" pageNumber="476">
<emphasis id="8DB77649FFBDFF9FD1CDF9AE45B3F9A8" bold="true" box="[151,385,1625,1650]" pageId="25" pageNumber="476">Material examined.</emphasis>
HOLOTYPE: male (13.4×
<quantity id="783B07BEFFBDFF9FD3EFF9AE472EF9A8" box="[693,796,1625,1650]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="25" pageNumber="476" unit="mm" value="11.5">11.5 mm</quantity>
) (MZB Cru 4801), Sulawesi: Bitung,
<date id="CB7D8C9BFFBDFF9FD594F9AD4151F9A8" box="[1230,1379,1625,1650]" pageId="25" pageNumber="476" value="2017-04-02">2 April 2017</date>
, J.-J. Li
<emphasis id="8DB77649FFBDFF9FD1E6F98944C0F94C" box="[188,242,1661,1686]" italics="true" pageId="25" pageNumber="476">et al</emphasis>
.
<materialsCitation id="0FABA006FFBDFF9FD05FF98944CEF960" ID-GBIF-Occurrence="1913889806" collectionCode="ZRC" pageId="25" pageNumber="476" specimenCode="ZRC 2018.0801" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<typeStatus id="607814F9FFBDFF9FD05FF9894596F94C" box="[261,420,1662,1686]" pageId="25" pageNumber="476" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="A9C561D2FFBDFF9FD0EEF989463DF94C" box="[436,527,1661,1686]" pageId="25" pageNumber="476" type="male">3 males</specimenCount>
(12.6×
<quantity id="783B07BEFFBDFF9FD33FF98A46E3F94F" box="[613,721,1661,1686]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="25" pageNumber="476" unit="mm" value="10.9">10.9 mm</quantity>
, 8.6×
<quantity id="783B07BEFFBDFF9FD249F98A4743F94F" box="[787,881,1661,1686]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="25" pageNumber="476" unit="mm" value="7.6">7.6 mm</quantity>
, 8.2×7.0 mm) (
<specimenCode id="EF650220FFBDFF9FD576F98940DEF94C" box="[1068,1260,1662,1686]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="25" pageNumber="476">ZRC 2018.0801</specimenCode>
), same data as holotype
</materialsCitation>
.—2 males (9.6×
<quantity id="783B07BEFFBDFF9FD09BF9564613F963" box="[449,545,1697,1722]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="25" pageNumber="476" unit="mm" value="7.9">7.9 mm</quantity>
, 6.7×
<quantity id="783B07BEFFBDFF9FD33CF95646F5F960" box="[614,711,1697,1722]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="25" pageNumber="476" unit="mm" value="5.7">5.7 mm</quantity>
),
<materialsCitation id="0FABA006FFBDFF9FD3BAF95544CDF904" ID-GBIF-Occurrence="1913889810" collectionCode="MZB" collectorName="Cru" pageId="25" pageNumber="476" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A9C561D2FFBDFF9FD3BAF955477AF960" box="[736,840,1697,1722]" pageId="25" pageNumber="476" type="female">1 female</specimenCount>
(11.9×
<quantity id="783B07BEFFBDFF9FD2FAF9554039F960" box="[928,1035,1698,1722]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="25" pageNumber="476" unit="mm" value="10.1">10.1 mm</quantity>
) (
<collectionCode id="D9D2329EFFBDFF9FD572F9554054F960" box="[1064,1126,1698,1722]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="25" pageNumber="476">MZB</collectionCode>
<collectorName id="1236CF8DFFBDFF9FD52EF95540ACF960" box="[1140,1182,1698,1722]" pageId="25" pageNumber="476">Cru</collectorName>
4802), same data as holotype
</materialsCitation>
.—
<materialsCitation id="0FABA006FFBDFF9FD07BF932404AF904" ID-GBIF-Occurrence="1913889809" box="[289,1144,1733,1758]" collectionCode="ZRC" collectorName="Cru" pageId="25" pageNumber="476" specimenCode="ZRC 2018.0802" specimenCount="1" typeStatus="paratype">
1 ovigerous female (11.2×
<quantity id="783B07BEFFBDFF9FD31BF93246A8F904" box="[577,666,1733,1758]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="25" pageNumber="476" unit="mm" value="9.6">9.6 mm</quantity>
) (
<specimenCode id="EF650220FFBDFF9FD3EAF9314759F904" box="[688,875,1734,1758]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="25" pageNumber="476">ZRC 2018.0802</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFBDFF9CD19DF91E4051FEA1" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476" type="diagnosis">
<paragraph id="BF7CAA5BFFBDFF9CD19DF91E4745FEEC" blockId="25.[151,1436,1625,2027]" lastBlockId="26.[151,1437,151,379]" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476">
<emphasis id="8DB77649FFBDFF9FD19DF91E4573F8D8" bold="true" box="[199,321,1769,1794]" pageId="25" pageNumber="476">Diagnosis.</emphasis>
Carapace (
<figureCitation id="27F8B6DEFFBDFF9FD09AF91E467FF8DB" box="[448,589,1769,1794]" captionStart="FIGURE 17" captionStartId="26.[151,250,1781,1803]" captionTargetBox="[152,1435,449,1769]" captionTargetId="figure@26.[152,1435,449,1769]" captionTargetPageId="26" captionText="FIGURE 17. Parasesarma tarantula n. sp.. AE, paratype male (12.6×10.9 mm) (ZRC 2018.0801), Sulawesi, Indonesia; F, G, paratype female (11.2×9.6 mm) (ZRC 2018.0802), same locality. A, F, dorsal views; B, E, G, ventral views; C, D, frontal views; E, close-up of gonopods." httpUri="https://zenodo.org/record/1440743/files/figure.png" pageId="25" pageNumber="476">Figs. 17A, F</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD303F91D46B8F8D8" box="[601,650,1770,1794]" captionStart="FIGURE 18" captionStartId="27.[151,250,1907,1929]" captionTargetBox="[192,1347,669,1874]" captionTargetId="figure@27.[160,1426,637,1891]" captionTargetPageId="27" captionText="FIGURE 18. Parasesarma tarantula n. sp., holotype male (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia. A, dorsal view of carapace; BD, tip of left cheliped; EH, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440745/files/figure.png" pageId="25" pageNumber="476">18A</figureCitation>
) squarish in general outline, 1.2 times broader than long; regions well defined, separated by shallow grooves; postfrontal region distinct, separated into 4 lobes by deep grooves; frontal margin slightly concaves in dorsal view; lateral margin straight, subparallel along most of length before curving to join almost straight posterior carapace margin; cornea extending or just reaching tip of external orbital tooth (
<figureCitation id="27F8B6DEFFBDFF9FD43EF8BA44D8F85C" captionStart="FIGURE 17" captionStartId="26.[151,250,1781,1803]" captionTargetBox="[152,1435,449,1769]" captionTargetId="figure@26.[152,1435,449,1769]" captionTargetPageId="26" captionText="FIGURE 17. Parasesarma tarantula n. sp.. AE, paratype male (12.6×10.9 mm) (ZRC 2018.0801), Sulawesi, Indonesia; F, G, paratype female (11.2×9.6 mm) (ZRC 2018.0802), same locality. A, F, dorsal views; B, E, G, ventral views; C, D, frontal views; E, close-up of gonopods." httpUri="https://zenodo.org/record/1440743/files/figure.png" pageId="25" pageNumber="476">Figs. 17A, F</figureCitation>
,
<figureCitation id="27F8B6DEFFBDFF9FD1A0F898451CF85D" box="[250,302,1903,1927]" captionStart="FIGURE 18" captionStartId="27.[151,250,1907,1929]" captionTargetBox="[192,1347,669,1874]" captionTargetId="figure@27.[160,1426,637,1891]" captionTargetPageId="27" captionText="FIGURE 18. Parasesarma tarantula n. sp., holotype male (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia. A, dorsal view of carapace; BD, tip of left cheliped; EH, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440745/files/figure.png" pageId="25" pageNumber="476">18A</figureCitation>
). Ischium of third maxilliped with shallow median sulcus, merus with distinct submedian ridge; exopod slender, tip reaching half-length of outer margin of merus, flagellum long. Male cheliped palm with 2 transverse pectinate crests (11 and 10 corneous teeth, distal and proximal crests, respectively) on upper surface; upper surface of dactylus with 1012 symmetrical, obliquely elongate dactylar tubercles, proximal tubercles steep, sharp, other tubercles large, distalmost tubercle indistinct (
<figureCitation id="27F8B6DEFFBEFF9CD27CFF60478AFF75" box="[806,952,151,176]" captionStart="FIGURE 18" captionStartId="27.[151,250,1907,1929]" captionTargetBox="[192,1347,669,1874]" captionTargetId="figure@27.[160,1426,637,1891]" captionTargetPageId="27" captionText="FIGURE 18. Parasesarma tarantula n. sp., holotype male (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia. A, dorsal view of carapace; BD, tip of left cheliped; EH, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440745/files/figure.png" pageId="26" pageNumber="477">Figs. 18C, D</figureCitation>
,
<figureCitation id="27F8B6DEFFBEFF9CD29EFF6F47CAFF6A" box="[964,1016,152,176]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="26" pageNumber="477">19H</figureCitation>
). Ambulatory legs relatively stout for this species-complex, P3 about 1.5 times carapace width; P3 and P4 coxae without dense setae; P3 merus 2.3 times as long as broad; P3 propodus 3.9 times as long as broad; P3 dactylus 0.5 times length of propodus (
<figureCitation id="27F8B6DEFFBEFF9CD5A7FF2D4155FF29" box="[1277,1383,218,243]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="26" pageNumber="477">Fig. 20G</figureCitation>
). G1 relatively slender (
<figureCitation id="27F8B6DEFFBEFF9CD031FF0B45CCFECE" box="[363,510,252,277]" captionStart="FIGURE 18" captionStartId="27.[151,250,1907,1929]" captionTargetBox="[192,1347,669,1874]" captionTargetId="figure@27.[160,1426,637,1891]" captionTargetPageId="27" captionText="FIGURE 18. Parasesarma tarantula n. sp., holotype male (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia. A, dorsal view of carapace; BD, tip of left cheliped; EH, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440745/files/figure.png" pageId="26" pageNumber="477">Figs. 18EH</figureCitation>
,
<figureCitation id="27F8B6DEFFBEFF9CD351FF0B460DFECE" box="[523,575,252,276]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="26" pageNumber="477">21H</figureCitation>
); apical process corneous, short, bent at angle of 45°, long, stout, ending in rounded tip. G2 longer than quarter length of G1 (
<figureCitation id="27F8B6DEFFBEFF9CD39DFEEA471EFEEC" box="[711,812,285,310]" captionStart="FIGURE 17" captionStartId="26.[151,250,1781,1803]" captionTargetBox="[152,1435,449,1769]" captionTargetId="figure@26.[152,1435,449,1769]" captionTargetPageId="26" captionText="FIGURE 17. Parasesarma tarantula n. sp.. AE, paratype male (12.6×10.9 mm) (ZRC 2018.0801), Sulawesi, Indonesia; F, G, paratype female (11.2×9.6 mm) (ZRC 2018.0802), same locality. A, F, dorsal views; B, E, G, ventral views; C, D, frontal views; E, close-up of gonopods." httpUri="https://zenodo.org/record/1440743/files/figure.png" pageId="26" pageNumber="477">Fig. 17E</figureCitation>
,
<figureCitation id="27F8B6DEFFBEFF9CD26DFEE94759FEEC" box="[823,875,286,310]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="26" pageNumber="477">21H</figureCitation>
).
</paragraph>
<paragraph id="BF7CAA5BFFBEFF9CD19DFEC94051FEA1" blockId="26.[151,1437,151,379]" pageId="26" pageNumber="477">
<emphasis id="8DB77649FFBEFF9CD19DFEC94543FE8D" bold="true" box="[199,369,318,343]" pageId="26" pageNumber="477">Colour in life.</emphasis>
Similar to
<taxonomicName id="78C3D1D8FFBEFF9CD0ACFEC84667FE8D" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[502,597,318,343]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFBEFF9CD0ACFEC84667FE8D" box="[502,597,318,343]" italics="true" pageId="26" pageNumber="477">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFBEFF9CD304FEC946ADFE8D" box="[606,671,318,343]" pageId="26" pageNumber="477" rank="species">
<emphasis id="8DB77649FFBEFF9CD304FEC946ADFE8D" bold="true" box="[606,671,318,343]" pageId="26" pageNumber="477">n. sp.</emphasis>
</taxonomicNameLabel>
, colour in general is dark brown. Carapace mottled dark and light brown, chelipeds brownish yellow, ambulatory legs light brown (
<figureCitation id="27F8B6DEFFBEFF9CD22BFE9547EDFEA1" box="[881,991,354,379]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="26" pageNumber="477">Figs. 22E</figureCitation>
,
<figureCitation id="27F8B6DEFFBEFF9CD2B1FE944029FEA1" box="[1003,1051,355,379]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="26" pageNumber="477">23E</figureCitation>
,
<figureCitation id="27F8B6DEFFBEFF9CD57CFE944064FEA1" box="[1062,1110,355,379]" captionStart="FIGURE 24" captionStartId="34.[151,250,1531,1553]" captionTargetBox="[151,1436,218,1490]" captionTargetId="figure@34.[151,1436,218,1490]" captionTargetPageId="34" captionText="FIGURE 24. In situ photographs of Parasesarma leptosoma species-complex, specimens not collected. A, P. leptosoma (Hilgendorf, 1869), Kenya (Photograph: S. Cannicci); B, P. gecko n. sp., Okinawa, Japan; C, P. macaco n. sp., Pingtung, Taiwan; D, P. kui n. sp., Pingtung, Taiwan; E, P. purpureum n. sp., Borneo, Malaysia; F, P. tarantula n. sp., Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440757/files/figure.png" pageId="26" pageNumber="477">24F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="EBBCFAD3FFBEFF9CD1CDF90245A6F892" httpUri="https://zenodo.org/record/1440743/files/figure.png" pageId="26" pageNumber="477" startId="26.[151,250,1781,1803]" targetBox="[152,1435,449,1769]" targetPageId="26">
<paragraph id="BF7CAA5BFFBEFF9CD1CDF90245A6F892" blockId="26.[151,1436,1781,1864]" pageId="26" pageNumber="477">
<emphasis id="8DB77649FFBEFF9CD1CDF902452AF8D0" bold="true" box="[151,280,1781,1803]" pageId="26" pageNumber="477">FIGURE 17</emphasis>
.
<taxonomicName id="78C3D1D8FFBEFF9CD07FF9014622F8D1" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[293,528,1781,1803]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="tarantula" status="sp. nov.">
<emphasis id="8DB77649FFBEFF9CD07FF9014622F8D1" box="[293,528,1781,1803]" italics="true" pageId="26" pageNumber="477">Parasesarma tarantula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFBEFF9CD34CF902467DF8D0" box="[534,591,1781,1802]" pageId="26" pageNumber="477" rank="species">
<emphasis id="8DB77649FFBEFF9CD34CF902467DF8D0" bold="true" box="[534,591,1781,1802]" pageId="26" pageNumber="477">n. sp.</emphasis>
</taxonomicNameLabel>
. AE, paratype male (12.6×10.9 mm) (ZRC 2018.0801), Sulawesi, Indonesia; F, G, paratype female (11.2×9.6 mm) (ZRC 2018.0802), same locality. A, F, dorsal views; B, E, G, ventral views; C, D, frontal views; E, close-up of gonopods.
</paragraph>
</caption>
<subSubSection id="F7D9F9D0FFBEFF9DD19DF8794696FE86" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477" type="discussion">
<paragraph id="BF7CAA5BFFBEFF9DD19DF8794140FF0F" blockId="26.[151,1436,1934,2031]" lastBlockId="27.[151,1437,151,592]" lastPageId="27" lastPageNumber="478" pageId="26" pageNumber="477">
<emphasis id="8DB77649FFBEFF9CD19DF8794509F87D" bold="true" box="[199,315,1934,1959]" pageId="26" pageNumber="477">Remarks.</emphasis>
The discovery of
<taxonomicName id="78C3D1D8FFBEFF9CD342F8784712F87D" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[536,800,1934,1959]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="tarantula" status="sp. nov.">
<emphasis id="8DB77649FFBEFF9CD342F8784712F87D" box="[536,800,1934,1959]" italics="true" pageId="26" pageNumber="477">Parasesarma tarantula</emphasis>
</taxonomicName>
<emphasis id="8DB77649FFBEFF9CD276F879475DF87D" bold="true" box="[812,879,1934,1959]" pageId="26" pageNumber="477">
<taxonomicNameLabel id="9684CB32FFBEFF9CD276F879475DF87D" box="[812,879,1934,1959]" pageId="26" pageNumber="477" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Sulawesi is unexpected. The species was found in Bitung, a site very close to Lembeh Island, the
<typeStatus id="607814F9FFBEFF9CD245F844477DF811" box="[799,847,1971,1995]" pageId="26" pageNumber="477">type</typeStatus>
locality of
<emphasis id="8DB77649FFBEFF9CD28FF844415AF811" box="[981,1384,1970,1995]" italics="true" pageId="26" pageNumber="477">
<taxonomicName id="78C3D1D8FFBEFF9CD28FF8444068F811" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[981,1114,1970,1995]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="limbense">P. limbense</taxonomicName>
.
<taxonomicName id="78C3D1D8FFBEFF9CD53CF844415AF811" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[1126,1384,1970,1995]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="limbense">Parasesarma limbense</taxonomicName>
</emphasis>
was found originally in Lembeh Island, but we were not found in Bitung, suggesting that the
<typeStatus id="607814F9FFBEFF9CD5F3F82040D6F835" box="[1193,1252,2007,2031]" pageId="26" pageNumber="477">types</typeStatus>
may have been collected in a different microhabitat Having two species of a species-complex occurring nearby or even together is not without precedence: in
<collectingCountry id="C7D4EACBFFBFFF9DD092FF4B462FFF0F" box="[456,541,188,213]" name="Taiwan" pageId="27" pageNumber="478">Taiwan</collectingCountry>
,
<taxonomicName id="78C3D1D8FFBFFF9DD372FF4A4657FF0F" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[552,613,188,213]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="kui" status="sp. nov.">
<emphasis id="8DB77649FFBFFF9DD372FF4A4657FF0F" box="[552,613,188,213]" italics="true" pageId="27" pageNumber="478">P. kui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFBFFF9DD336FF4B4699FF0F" box="[620,683,188,213]" pageId="27" pageNumber="478" rank="species">
<emphasis id="8DB77649FFBFFF9DD336FF4B4699FF0F" bold="true" box="[620,683,188,213]" pageId="27" pageNumber="478">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="78C3D1D8FFBFFF9DD3BBFF4A4766FF0E" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[737,852,189,212]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFBFFF9DD3BBFF4A4766FF0E" box="[737,852,189,212]" italics="true" pageId="27" pageNumber="478">P. macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFBFFF9DD201FF4B47A8FF0F" box="[859,922,188,213]" pageId="27" pageNumber="478" rank="species">
<emphasis id="8DB77649FFBFFF9DD201FF4B47A8FF0F" bold="true" box="[859,922,188,213]" pageId="27" pageNumber="478">n. sp.</emphasis>
</taxonomicNameLabel>
also live sympatrically in some locations.
</paragraph>
<paragraph id="BF7CAA5BFFBFFF9DD1CDFF164696FE86" blockId="27.[151,1437,151,592]" pageId="27" pageNumber="478">
<taxonomicName id="78C3D1D8FFBFFF9DD1CDFF1645AEFF22" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[151,412,223,248]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="tarantula" status="sp. nov.">
<emphasis id="8DB77649FFBFFF9DD1CDFF1645AEFF22" box="[151,412,223,248]" italics="true" pageId="27" pageNumber="478">Parasesarma tarantula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFBFFF9DD0FEFF2845D1FF22" box="[420,483,223,248]" pageId="27" pageNumber="478" rank="species">
<emphasis id="8DB77649FFBFFF9DD0FEFF2845D1FF22" bold="true" box="[420,483,223,248]" pageId="27" pageNumber="478">n. sp.</emphasis>
</taxonomicNameLabel>
most closely resembles
<taxonomicName id="78C3D1D8FFBFFF9DD3ADFF164766FF22" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[759,852,223,248]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFBFFF9DD3ADFF164766FF22" box="[759,852,223,248]" italics="true" pageId="27" pageNumber="478">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFBFFF9DD206FF2847A9FF22" box="[860,923,223,248]" pageId="27" pageNumber="478" rank="species">
<emphasis id="8DB77649FFBFFF9DD206FF2847A9FF22" bold="true" box="[860,923,223,248]" pageId="27" pageNumber="478">n. sp.</emphasis>
</taxonomicNameLabel>
in the carapace shape and live colouration. It differs from all other members of the species-complex in having the P3 propodus distinctly shorter (
<figureCitation id="27F8B6DEFFBFFF9DD45DFEF64143FEC3" box="[1287,1393,257,282]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="27" pageNumber="478">Fig. 20G</figureCitation>
). In addition,
<taxonomicName id="78C3D1D8FFBFFF9DD05BFED345B7FEE1" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[257,389,290,315]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="tarantula">
<emphasis id="8DB77649FFBFFF9DD05BFED345B7FEE1" box="[257,389,290,315]" italics="true" pageId="27" pageNumber="478">P. tarantula</emphasis>
</taxonomicName>
is easily separated from
<taxonomicName id="78C3D1D8FFBFFF9DD3C4FED3472CFEE1" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[670,798,290,315]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="limbense">
<emphasis id="8DB77649FFBFFF9DD3C4FED3472CFEE1" box="[670,798,290,315]" italics="true" pageId="27" pageNumber="478">P. limbense</emphasis>
</taxonomicName>
by its prominently concave frontal margin, as well as its straight and elongated G1 (
<figureCitation id="27F8B6DEFFBFFF9DD09DFEB3466AFE86" box="[455,600,324,349]" captionStart="FIGURE 18" captionStartId="27.[151,250,1907,1929]" captionTargetBox="[192,1347,669,1874]" captionTargetId="figure@27.[160,1426,637,1891]" captionTargetPageId="27" captionText="FIGURE 18. Parasesarma tarantula n. sp., holotype male (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia. A, dorsal view of carapace; BD, tip of left cheliped; EH, left G1. B, lateral view of pectinated cristae on palm; C, dorsal view of palm and dactylar finger; D, lateral view of dactylar finger; E, G, dorsal (sternal) views; F, H, ventral (pleonal) views; E, F, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440745/files/figure.png" pageId="27" pageNumber="478">Figs. 18EH</figureCitation>
,
<figureCitation id="27F8B6DEFFBFFF9DD339FEB346A5FE86" box="[611,663,324,348]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="27" pageNumber="478">21H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFBFFF9DD19DFE9246DCFE1C" pageId="27" pageNumber="478" type="etymology">
<paragraph id="BF7CAA5BFFBFFF9DD19DFE9246DCFE1C" blockId="27.[151,1437,151,592]" pageId="27" pageNumber="478">
<emphasis id="8DB77649FFBFFF9DD19DFE92457CFEA4" bold="true" box="[199,334,357,382]" pageId="27" pageNumber="478">Etymology.</emphasis>
The name is from the old Italian common name for large spiders,
<taxonomicName id="78C3D1D8FFBFFF9DD519FE92409BFEA4" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[1091,1193,357,382]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="tarantula">tarantula</taxonomicName>
. The use of the name here alludes to the tree-climbing habits of the new species, similar to many species of
<taxonomicName id="78C3D1D8FFBFFF9DD507FE7E40F0FE78" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[1117,1218,393,418]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="tarantula">tarantula</taxonomicName>
, some of which are also found in Sulawesi. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFBFFF9DD19DFE26477FFE30" box="[199,845,465,490]" pageId="27" pageNumber="478" type="distribution">
<paragraph id="BF7CAA5BFFBFFF9DD19DFE26477FFE30" blockId="27.[151,1437,151,592]" box="[199,845,465,490]" pageId="27" pageNumber="478">
<emphasis id="8DB77649FFBFFF9DD19DFE264552FE30" bold="true" box="[199,352,465,490]" pageId="27" pageNumber="478">Distribution.</emphasis>
Only known from North Sulawesi, thus far.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFBFFF9DD1CDFE0244CBFD8A" pageId="27" pageNumber="478" type="biology_ecology">
<paragraph id="BF7CAA5BFFBFFF9DD1CDFE0244CBFD8A" blockId="27.[151,1437,151,592]" pageId="27" pageNumber="478">
<emphasis id="8DB77649FFBFFF9DD1CDFE0244C9FDD4" bold="true" box="[151,251,501,526]" pageId="27" pageNumber="478">Ecology.</emphasis>
In Sulawesi,
<taxonomicName id="78C3D1D8FFBFFF9DD0CFFE0146A8FDD4" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[405,666,501,526]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="tarantula" status="sp. nov.">
<emphasis id="8DB77649FFBFFF9DD0CFFE0146A8FDD4" box="[405,666,501,526]" italics="true" pageId="27" pageNumber="478">Parasesarma tarantula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFBFFF9DD3FBFE0246D2FDD4" box="[673,736,501,526]" pageId="27" pageNumber="478" rank="species">
<emphasis id="8DB77649FFBFFF9DD3FBFE0246D2FDD4" bold="true" box="[673,736,501,526]" pageId="27" pageNumber="478">n. sp.</emphasis>
</taxonomicNameLabel>
is often observed climbing on mangrove trees and dead wood above the water. They can also found in tree holes near the water level. They are generally active during the daytime.
</paragraph>
</subSubSection>
</treatment>
</document>