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<document id="4BB1F71F823F41A1708924EBA682C0A1" ID-CLB-Dataset="29418" ID-DOI="10.11646/zootaxa.4482.3.2" ID-GBIF-Dataset="fd24ef18-76ac-46f1-aec2-2faf5b0ff325" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440707" ID-ZooBank="3FBE11E2-9F97-4A29-92A9-67780C782E0D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538375301724" checkinUser="plazi" docAuthor="Li, Jheng-Jhang, Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2018" docId="376A1B4DFFADFF8BD1CDFD444536FC87" docLanguage="en" docName="zootaxa.4482.3.2.pdf" docOrigin="Zootaxa 4482 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parasesarma gecko Li &amp; Rahayu &amp; Ng 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="464" masterDocId="CB536335FFA4FF86D15AFFF74432FFDA" masterDocTitle="Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific" masterLastPageNumber="490" masterPageNumber="451" pageNumber="460" updateTime="1698559507815" updateUser="ExternalLinkService">
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<mods:title id="5551B44B7D5A1D1D517BC4113BE192DF">Identity of the tree-spider crab, Parasesarma leptosoma (Hilgendorf, 1869) (Decapoda: Brachyura: Sesarmidae), with descriptions of seven new species from the Western Pacific</mods:title>
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(
<figureCitation id="27F8B6DEFFADFF8FD1C5FD214539FD35" box="[159,267,726,751]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="10.[151,250,1920,1942]" captionStartId-1="11.[151,250,1376,1398]" captionStartId-2="12.[151,250,815,837]" captionTargetBox-0="[192,1407,193,1911]" captionTargetBox-1="[186,1355,232,1354]" captionTargetBox-2="[175,1397,233,748]" captionTargetId-0="figure@10.[192,1407,193,1911]" captionTargetId-1="figure@11.[152,1435,193,1354]" captionTargetId-2="figure@12.[151,1435,193,794]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 4. Parasesarma gecko n. sp.. A–D, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; E, F, paratype female (15.3×13.0 mm) (ASIZ 73381), Iriomote Island, Japan; G, H, female (20.2×17.5 mm) (NMNS-7779-002), Okinawa Island, Japan, showing abnormal right second ambulatory leg; I, J, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan, showing asymmetrical chelae. A, E, G, I, dorsal views; B, F, H, J, ventral views; C, D, frontal views." captionText-1="FIGURE 5. Parasesarma gecko n. sp., holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan. A, dorsal view of carapace; B, lateral view of pectinated cristae on palm of left chela; C, dorsal view of tip of left cheliped; D–G, left G1. D, F, dorsal (sternal) views; E, G, ventral (pleonal) views; D, E, distal part of G1. Scale bars: 1.0 mm." captionText-2="FIGURE 6. Parasesarma gecko n. sp., left chela. A, D, paratype male (14.4×12.6 mm) (ZRC 2008.1252), Miyako Island, Japan; B, E, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan; C, F, paratype male (14.5× 12.3 mm) (ZRC 2008.0501), Guam. A–C, dorsal views of pectinated cristae on palm, arrow points to direction of dactylar finger; D–F, lateral views of pectinated cristae on palm. Scale bars: 1.0 mm." httpUri-0="https://zenodo.org/record/1440717/files/figure.png" httpUri-1="https://zenodo.org/record/1440719/files/figure.png" httpUri-2="https://zenodo.org/record/1440721/files/figure.png" pageId="9" pageNumber="460">Figs. 4–6</figureCitation>
, 
<figureCitation id="27F8B6DEFFADFF8FD04CFD20457AFD35" box="[278,328,727,751]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="9" pageNumber="460">19B</figureCitation>
, 
<figureCitation id="27F8B6DEFFADFF8FD009FD2045B4FD35" box="[339,390,727,751]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="9" pageNumber="460">20A</figureCitation>
, 
<figureCitation id="27F8B6DEFFADFF8FD0CBFD2045F0FD35" box="[401,450,727,751]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="9" pageNumber="460">21B</figureCitation>
, 
<figureCitation id="27F8B6DEFFADFF8FD094FD204633FD35" box="[462,513,727,751]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="9" pageNumber="460">22A</figureCitation>
, 
<figureCitation id="27F8B6DEFFADFF8FD356FD20460DFD35" box="[524,575,727,751]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="9" pageNumber="460">23A</figureCitation>
, 
<figureCitation id="27F8B6DEFFADFF8FD310FD20464EFD35" box="[586,636,727,751]" captionStart="FIGURE 24" captionStartId="34.[151,250,1531,1553]" captionTargetBox="[151,1436,218,1490]" captionTargetId="figure@34.[151,1436,218,1490]" captionTargetPageId="34" captionText="FIGURE 24. In situ photographs of Parasesarma leptosoma species-complex, specimens not collected. A, P. leptosoma (Hilgendorf, 1869), Kenya (Photograph: S. Cannicci); B, P. gecko n. sp., Okinawa, Japan; C, P. macaco n. sp., Pingtung, Taiwan; D, P. kui n. sp., Pingtung, Taiwan; E, P. purpureum n. sp., Borneo, Malaysia; F, P. tarantula n. sp., Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440757/files/figure.png" pageId="9" pageNumber="460">24B</figureCitation>
, 
<figureCitation id="27F8B6DEFFADFF8FD3DDFD214687FD35" box="[647,693,726,751]" captionStart="FIGURE 25" captionStartId="35.[151,250,1877,1899]" captionTargetBox="[170,1415,193,1866]" captionTargetId="figure@35.[170,1415,193,1866]" captionTargetPageId="35" captionText="FIGURE 25. Habitats associated with Parasesarma macaco n. sp., P. gecko n. sp. and P. kui n. sp., specimens not collected. A–D, P. macaco n. sp., Pingtung (Paoli River), Taiwan; E, P. gecko n. sp., group of crabs on cement wall near mangrove, Okinawa, Japan; F, G, P. kui n. sp., Pingtung (Kangkou River), Taiwan. A, habitat constituted by Talipariti tiliaceum and Clerodendrum inerme; B, male climbing on T. tiliaceum; C, female climbing on C. inerme; D, ovigerous female during breeding migration; F, habitat constituted by T. tiliaceum; G, male climbing on shrubs." httpUri="https://zenodo.org/record/1440759/files/figure.png" pageId="9" pageNumber="460">25E</figureCitation>
)
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<treatmentCitationGroup id="9FD38D75FFADFF8FD1CDFCE841AFFCEE" box="[151,1437,798,821]" pageId="9" pageNumber="460">
<taxonomicName id="78C3D1D8FFADFF8FD1CDFCE8461BFCEE" ID-CoL="7Z29P" authority=", De Man 1889: 436" authorityName="De Man" authorityPageNumber="436" authorityYear="1889" box="[151,553,798,820]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFADFF8FD1CDFCE8456EFCEE" box="[151,348,798,820]" italics="true" pageId="9" pageNumber="460">Sesarma leptosoma</emphasis>
, 
<treatmentCitation id="3E628C4AFFADFF8FD033FCE8461BFCEE" author="" box="[361,553,798,820]" page="436" pageId="9" pageNumber="460">
<bibRefCitation id="DB52D7AAFFADFF8FD033FCE845C5FCEE" box="[361,503,798,820]" pageId="9" pageNumber="460" refId="ref17312" refString="Man, J. G. De (1889) Uber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 4, 409 - 452." type="journal article">De Man 1889</bibRefCitation>
: 436
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</taxonomicName>
, pl. 10, fig. 11.—Ortmann 1894: 725. Not 
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<emphasis id="8DB77649FFADFF8FD281FCD740E2FCEE" box="[987,1232,798,821]" italics="true" pageId="9" pageNumber="460">Parasesarma leptosoma</emphasis>
(
<bibRefCitation id="DB52D7AAFFADFF8FD587FCE941A3FCEE" box="[1245,1425,798,820]" pageId="9" pageNumber="460" refId="ref16852" refString="Hilgendorf, F. (1869) Crustaceen. In: C. V. D. Decken (Ed), Reisen in Ost-Afrika in den Jahren 1859 - 1865. Heidelberg - Leipzig, pp. 67 - 116. http: // dx. doi. org / 10.5962 / bhl. title. 10640" type="book chapter">Hilgendorf, 1869</bibRefCitation>
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.
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<taxonomicName id="78C3D1D8FFADFF8FD1CDFCC946B4FC89" ID-CoL="7XX38" authority=", Nakasone 1977: 10" authorityName="Nakasone" authorityPageNumber="10" authorityYear="1977" box="[151,646,829,851]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFADFF8FD1CDFCC9452CFC89" box="[151,286,830,851]" italics="true" pageId="9" pageNumber="460">Parasesarma</emphasis>
aff. 
<emphasis id="8DB77649FFADFF8FD016FCCA4586FC89" box="[332,436,829,851]" italics="true" pageId="9" pageNumber="460">leptosoma</emphasis>
, 
<treatmentCitation id="3E628C4AFFADFF8FD09BFCCA46B4FC89" author="" box="[449,646,829,851]" page="10" pageId="9" pageNumber="460">
<bibRefCitation id="DB52D7AAFFADFF8FD09BFCCA4653FC89" box="[449,609,829,851]" pageId="9" pageNumber="460" refId="ref17664" refString="Nakasone, Y. (1977) The ecology and distribution of mangrove animals. In: Investigations of natural monument from Okinawa 10: the emergency of investigations Rhizophoraceae mangrove of Gesashi Bay. Education Section, Okinawa Prefecture, Naha, pp. 9 - 38." type="book chapter">Nakasone 1977</bibRefCitation>
: 10
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, 11.—Nishidaira 1980: 66.
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</paragraph>
<paragraph id="BF7CAA5BFFADFF8FD1CDFCAB416EFCA8" blockId="9.[151,1437,798,974]" box="[151,1372,860,882]" pageId="9" pageNumber="460">
<treatmentCitationGroup id="9FD38D75FFADFF8FD1CDFCAB416EFCA8" box="[151,1372,860,882]" pageId="9" pageNumber="460">
<taxonomicName id="78C3D1D8FFADFF8FD1CDFCAB4720FCA8" ID-CoL="7Z29P" authority=", McLay &amp; Ryan 1990: 115" authorityName=", McLay &amp; Ryan" authorityPageNumber="115" authorityYear="1990" box="[151,786,860,882]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="leptosoma" subGenus="Parasesarma">
<emphasis id="8DB77649FFADFF8FD1CDFCAB44DCFCAB" box="[151,238,860,881]" italics="true" pageId="9" pageNumber="460">Sesarma</emphasis>
(
<emphasis id="8DB77649FFADFF8FD1A7FCAA45B6FCA8" box="[253,388,861,882]" italics="true" pageId="9" pageNumber="460">Parasesarma</emphasis>
) 
<emphasis id="8DB77649FFADFF8FD0C8FCAB45C8FCA8" box="[402,506,860,882]" italics="true" pageId="9" pageNumber="460">leptosoma</emphasis>
, 
<treatmentCitation id="3E628C4AFFADFF8FD35DFCAA4720FCA8" author="" box="[519,786,860,882]" page="115" pageId="9" pageNumber="460">
<bibRefCitation id="DB52D7AAFFADFF8FD35DFCAA46D0FCA8" box="[519,738,860,882]" pageId="9" pageNumber="460" refId="ref17530" refString="McLay, C. L. &amp; Ryan, P. A. (1990) The terrestrial crabs Sesarma (Sesarmops) impressum and Geograpsus crinipes (Brachyura, Grapsidae, Sesarminae) recorded from the Fiji Island. Journal of the Royal Society of New Zealand, 20, 107 - 118. http: // dx. doi. org / 10.1080 / 03036758.1990.10426735" type="journal article">McLay &amp; Ryan 1990</bibRefCitation>
: 115
</treatmentCitation>
</taxonomicName>
, table 2. Not 
<taxonomicName id="78C3D1D8FFADFF8FD2C0FCAA416BFCA8" ID-CoL="7XX38" authority="(Hilgendorf, 1869)" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[922,1369,860,882]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFADFF8FD2C0FCAA40BDFCA8" box="[922,1167,860,882]" italics="true" pageId="9" pageNumber="460">Parasesarma leptosoma</emphasis>
(
<bibRefCitation id="DB52D7AAFFADFF8FD5C7FCAB4163FCA8" box="[1181,1361,860,882]" pageId="9" pageNumber="460" refId="ref16852" refString="Hilgendorf, F. (1869) Crustaceen. In: C. V. D. Decken (Ed), Reisen in Ost-Afrika in den Jahren 1859 - 1865. Heidelberg - Leipzig, pp. 67 - 116. http: // dx. doi. org / 10.5962 / bhl. title. 10640" type="book chapter">Hilgendorf, 1869</bibRefCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BF7CAA5BFFADFF8FD1CDFC8C40A7FC14" blockId="9.[151,1437,798,974]" pageId="9" pageNumber="460">
<treatmentCitationGroup id="9FD38D75FFADFF8FD1CDFC8C40A7FC14" pageId="9" pageNumber="460">
<taxonomicName id="78C3D1D8FFADFF8FD1CDFC8C46E6FC4A" ID-CoL="7XX38" authority=", Takeda &amp; Nunomura 1976: 86" authorityName="Takeda &amp; Nunomura" authorityPageNumber="86" authorityYear="1976" box="[151,724,890,912]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFADFF8FD1CDFC8C45BFFC4A" box="[151,397,890,912]" italics="true" pageId="9" pageNumber="460">Parasesarma leptosoma</emphasis>
, Takeda &amp; Nunomura 1976: 86
</taxonomicName>
.— 
<treatmentCitation id="3E628C4AFFADFF8FD3A8FC8D47ADFC4A" author="" box="[754,927,890,912]" page="223" pageId="9" pageNumber="460">
<bibRefCitation id="DB52D7AAFFADFF8FD3A8FC8D4742FC4A" box="[754,880,890,912]" pageId="9" pageNumber="460" refId="ref16423" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda-Anomura, Brachyura. In: Wells, A. &amp; Houston, W. W. K. (Eds.), Zological Catalogue of Australia, vol. 19.3 B. CSIRO Publishing Australia, Melbourne, pp. 641." type="book chapter">Davie 2002</bibRefCitation>
: 223
</treatmentCitation>
.—Shokita 
<emphasis id="8DB77649FFADFF8FD54EFC8C4070FC4A" box="[1044,1090,890,912]" italics="true" pageId="9" pageNumber="460">et al</emphasis>
. 2002: 78. —Shokita 
<emphasis id="8DB77649FFADFF8FD47DFC8C4167FC4A" box="[1319,1365,890,912]" italics="true" pageId="9" pageNumber="460">et al</emphasis>
. 2003: 102.—Rahayu &amp; Ng 2009: 35.— 
<treatmentCitation id="3E628C4AFFADFF8FD34FFC6E46CFFC75" author="" box="[533,765,921,943]" page="359" pageId="9" pageNumber="460">
<bibRefCitation id="DB52D7AAFFADFF8FD34FFC6E46FDFC75" box="[533,719,921,943]" pageId="9" pageNumber="460" refId="ref17023" refString="Koller, P., Liu, H. C. &amp; Schubart, C. D. (2010) A new semiterrestrial species of Parasesarma De Man, 1895, from Taiwan (Decapoda, Brachyura, Sesarmidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Folume. Crustaceana Monographs, 14, 357 - 368." type="journal article">
Koller 
<emphasis id="8DB77649FFADFF8FD305FC6D46B9FC75" box="[607,651,921,943]" italics="true" pageId="9" pageNumber="460">et al</emphasis>
. 2010
</bibRefCitation>
: 359
</treatmentCitation>
.— 
<treatmentCitation id="3E628C4AFFADFF8FD241FC6E47F0FC75" author="" box="[795,962,921,943]" page="51" pageId="9" pageNumber="460">
<bibRefCitation id="DB52D7AAFFADFF8FD241FC6E4790FC75" box="[795,930,921,943]" pageId="9" pageNumber="460" refId="ref17763" refString="Naruse, T. (2010) Catalogue of materials deposited in the University Museum (Fujukan). University of the Ryukyus No. 3. University of the Ryukyus. The University Museum (Fujukan), University of the Ryukyus, Okinawa. [in Japanese]" type="book">Naruse 2010</bibRefCitation>
: 51
</treatmentCitation>
.— 
<treatmentCitation id="3E628C4AFFADFF8FD2BAFC6E4111FC75" author="" box="[992,1315,921,943]" page="13" pageId="9" pageNumber="460">
<bibRefCitation id="DB52D7AAFFADFF8FD2BAFC6E40CCFC75" box="[992,1278,921,943]" pageId="9" pageNumber="460" refId="ref17144" refString="Maenosono, T. &amp; Naruse, T. (2015) Morphological characteristics and taxonomical problems of the genus Parasesarma De Man, 1895 (Crustacea: Decapoda: Brachyura: Sesarmidae) from the Ryukyu Archipelago, Japan. Fauna Ryukyuana, 23, 1 - 41 [in Japanese]." type="journal article">Maenosono &amp; Naruse 2015</bibRefCitation>
: 13
</treatmentCitation>
, figs. 2E, 6, 11E, F.— 
<treatmentCitation id="3E628C4AFFADFF8FD07CFC4E4678FC14" author="" box="[294,586,952,974]" page="8" pageId="9" pageNumber="460">
<bibRefCitation id="DB52D7AAFFADFF8FD07CFC4E4601FC14" box="[294,563,952,974]" pageId="9" pageNumber="460" refId="ref17198" refString="Maenosono, T. &amp; Saeki, T. (2016) The sesarmid (Crustacea: Decapoda: Brachyura) fauna of Ishigaki-jima Island, Ryukyu Archipelago, Japan, with new distributional records. Fauna Ryukyuana, 33, 1 - 13 [in Japanese with English abstract, Figure captions]." type="journal article">Maenosono &amp; Saeki 2016</bibRefCitation>
: 8
</treatmentCitation>
, table 1. Not 
<taxonomicName id="78C3D1D8FFADFF8FD388FC4E40A3FC14" ID-CoL="7XX38" authority="(Hilgendorf, 1869)" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[722,1169,952,974]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFADFF8FD388FC4E47F5FC14" box="[722,967,952,974]" italics="true" pageId="9" pageNumber="460">Parasesarma leptosoma</emphasis>
(
<bibRefCitation id="DB52D7AAFFADFF8FD28FFC4F40BBFC14" box="[981,1161,952,974]" pageId="9" pageNumber="460" refId="ref16852" refString="Hilgendorf, F. (1869) Crustaceen. In: C. V. D. Decken (Ed), Reisen in Ost-Afrika in den Jahren 1859 - 1865. Heidelberg - Leipzig, pp. 67 - 116. http: // dx. doi. org / 10.5962 / bhl. title. 10640" type="book chapter">Hilgendorf, 1869</bibRefCitation>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFADFF8FD1CDFC0D4639F804" pageId="9" pageNumber="460" type="materials_examined">
<paragraph id="BF7CAA5BFFADFF8FD1CDFC0D4639F804" blockId="9.[151,1437,1018,2015]" pageId="9" pageNumber="460">
<materialsCitation id="0FABA006FFADFF8FD1CDFC0D474AFBED" ID-GBIF-Occurrence="1913889779" collectingDate="2016-07-18" collectorName="J. Li" country="Japan" location="Awase" municipality="Material" pageId="9" pageNumber="460" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="8DB77649FFADFF8FD1CDFC0D45B0FBC9" bold="true" box="[151,386,1018,1043]" pageId="9" pageNumber="460">
<collectingMunicipality id="5F183021FFADFF8FD1CDFC0D44CDFBC9" box="[151,255,1018,1043]" pageId="9" pageNumber="460">Material</collectingMunicipality>
examined.
</emphasis>
<typeStatus id="607814F9FFADFF8FD0D1FC0C4611FBC9" box="[395,547,1019,1043]" pageId="9" pageNumber="460" type="holotype">HOLOTYPE</typeStatus>
: male (16.2× 
<quantity id="783B07BEFFADFF8FD3E2FC0D4712FBC9" box="[696,800,1018,1043]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="9" pageNumber="460" unit="mm" value="14.6">14.6 mm</quantity>
) (RUMF-ZC-4710), 
<collectingCountry id="C7D4EACBFFADFF8FD54BFC0C4067FBC8" box="[1041,1109,1019,1042]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD53BFC0D40F4FBC9" box="[1121,1222,1018,1043]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Island: 
<location id="BA1CFC80FFADFF8FD47EFC0C415CFBC9" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD47EFC0C415CFBC9" box="[1316,1390,1019,1043]" country="Japan" municipality="Material" name="Awase" pageId="9" pageNumber="460" stateProvince="Okinawa">Awase</location>
(
<collectedFrom id="0CDFE9A6FFADFF8FD425FC0C466FFBED" pageId="9" pageNumber="460">
on 
<taxonomicName id="78C3D1D8FFADFF8FD1CDFBE9466FFBED" authority="Sheue, Liu &amp; Yong" authorityName="Sheue, Liu &amp; Yong" box="[151,605,1054,1079]" class="Magnoliopsida" family="Rhizophoraceae" genus="Kandelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="9" pageNumber="460" phylum="Tracheophyta" rank="species" species="obovata">
<emphasis id="8DB77649FFADFF8FD1CDFBE94556FBED" box="[151,356,1054,1079]" italics="true" pageId="9" pageNumber="460">Kandelia obovata</emphasis>
Sheue, Liu &amp; Yong
</taxonomicName>
</collectedFrom>
), 
<collectingDate id="DB397573FFADFF8FD32FFBE84721FBED" box="[629,787,1054,1079]" pageId="9" pageNumber="460" value="2016-07-18">
<date id="CB7D8C9BFFADFF8FD32FFBE84721FBED" box="[629,787,1054,1079]" pageId="9" pageNumber="460" value="2016-07-18">18 July 2016</date>
</collectingDate>
, J.- 
<collectorName id="1236CF8DFFADFF8FD264FBE8474AFBED" box="[830,888,1054,1079]" pageId="9" pageNumber="460">J. Li</collectorName>
</materialsCitation>
. 
<materialsCitation id="0FABA006FFADFF8FD2D2FBE84776FB81" ID-GBIF-Occurrence="1913889769" collectingDate="2001-08" collectionCode="ZRC" collectorName="C. Liu." country="Guam" location="Pago Bay" pageId="9" pageNumber="460" specimenCode="ZRC 2008.0501" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="607814F9FFADFF8FD2D2FBE84015FBED" box="[904,1063,1055,1079]" pageId="9" pageNumber="460" type="paratype">PARATYPES</typeStatus>
: 
<specimenCount id="A9C561D2FFADFF8FD56DFBE840B5FBED" box="[1079,1159,1054,1079]" pageId="9" pageNumber="460" type="male">1 male</specimenCount>
(14.5× 
<quantity id="783B07BEFFADFF8FD587FBE9417BFBED" box="[1245,1353,1054,1079]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="9" pageNumber="460" unit="mm" value="12.3">12.3 mm</quantity>
) (
<specimenCode id="EF650220FFADFF8FD43EFBE84520FB81" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="460">ZRC 2008.0501</specimenCode>
), 
<collectingCountry id="C7D4EACBFFADFF8FD07CFBB4455DFB81" box="[294,367,1091,1115]" name="Guam" pageId="9" pageNumber="460">Guam</collectingCountry>
: near 
<location id="BA1CFC80FFADFF8FD0E9FBB44613FB81" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD0E9FBB44613FB81" box="[435,545,1091,1115]" country="Guam" name="Pago Bay" pageId="9" pageNumber="460">Pago Bay</location>
, 
<collectingDate id="DB397573FFADFF8FD374FBB446F0FB81" box="[558,706,1091,1115]" pageId="9" pageNumber="460" value="2001-08">
<date id="CB7D8C9BFFADFF8FD374FBB446F0FB81" box="[558,706,1091,1115]" pageId="9" pageNumber="460" value="2001-08">August 2001</date>
</collectingDate>
, H.- 
<collectorName id="1236CF8DFFADFF8FD3A9FBB44776FB81" box="[755,836,1090,1115]" pageId="9" pageNumber="460">C. Liu.</collectorName>
</materialsCitation>
—1 male (14.4× 
<quantity id="783B07BEFFADFF8FD2A6FBB54054FB81" box="[1020,1126,1090,1115]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="9" pageNumber="460" unit="mm" value="12.6">12.6 mm</quantity>
), 
<materialsCitation id="0FABA006FFADFF8FD520FBB44075FBA5" ID-GBIF-Occurrence="1913889773" collectingDate="2001-03-20" collectionCode="ZRC" collectorName="P. K. L. Ng." country="Japan" location="Shimajiri" municipality="Miyako Island" pageId="9" pageNumber="460" specimenCode="ZRC 2008.1252" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD520FBB440EFFB81" box="[1146,1245,1090,1115]" pageId="9" pageNumber="460" type="female">1 female</specimenCount>
(12.5×11.0 mm) (
<specimenCode id="EF650220FFADFF8FD1C4FB90456CFBA5" box="[158,350,1126,1151]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="460">ZRC 2008.1252</specimenCode>
), 
<collectingCountry id="C7D4EACBFFADFF8FD02CFB904588FBA4" box="[374,442,1127,1150]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingMunicipality id="5F183021FFADFF8FD090FB914644FBA5" box="[458,630,1126,1151]" pageId="9" pageNumber="460">Miyako Island</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD3DCFB9146C3FBA5" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD3DCFB9146C3FBA5" box="[646,753,1126,1151]" country="Japan" municipality="Miyako Island" name="Shimajiri" pageId="9" pageNumber="460">Shimajiri</location>
, 
<collectingDate id="DB397573FFADFF8FD25BFB904788FBA5" box="[769,954,1126,1151]" pageId="9" pageNumber="460" value="2001-03-20">
<date id="CB7D8C9BFFADFF8FD25BFB904788FBA5" box="[769,954,1126,1151]" pageId="9" pageNumber="460" value="2001-03-20">20 March 2001</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD290FB904075FBA5" box="[970,1095,1127,1151]" pageId="9" pageNumber="460">P.K.L. Ng.</collectorName>
</materialsCitation>
—1 male (15.9× 
<quantity id="783B07BEFFADFF8FD452FB914144FBA5" box="[1288,1398,1126,1151]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="9" pageNumber="460" unit="mm" value="13.7">13.7 mm</quantity>
), 
<materialsCitation id="0FABA006FFADFF8FD4D4FB90414DFB79" ID-GBIF-Occurrence="1913889778" collectingDate="2004-09-17" collectionCode="ASIZ" collectorName="C. Liu." country="Japan" location="Iriomote Island" municipality="Yaeyama Islands" pageId="9" pageNumber="460" specimenCode="ASIZ 73381" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD4D4FB9044D1FB79" pageId="9" pageNumber="460" type="female">1 female</specimenCount>
(15.3×13.0 mm) (
<specimenCode id="EF650220FFADFF8FD0EAFB7C4672FB79" box="[432,576,1163,1187]" collectionCode="ASIZ" pageId="9" pageNumber="460">ASIZ 73381</specimenCode>
) 
<collectingCountry id="C7D4EACBFFADFF8FD314FB7C46A0FB78" box="[590,658,1163,1186]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingMunicipality id="5F183021FFADFF8FD3C7FB7C476DFB79" box="[669,863,1162,1187]" pageId="9" pageNumber="460">Yaeyama Islands</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD230FB7D402AFB79" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD230FB7D402AFB79" box="[874,1048,1162,1187]" country="Japan" municipality="Yaeyama Islands" name="Iriomote Island" pageId="9" pageNumber="460">Iriomote Island</location>
, 
<collectingDate id="DB397573FFADFF8FD57EFB7C4130FB79" box="[1060,1282,1162,1187]" pageId="9" pageNumber="460" value="2004-09-17">
<date id="CB7D8C9BFFADFF8FD57EFB7C4130FB79" box="[1060,1282,1162,1187]" pageId="9" pageNumber="460" value="2004-09-17">17 September 2004</date>
</collectingDate>
, H.- 
<collectorName id="1236CF8DFFADFF8FD46BFB7C414DFB79" box="[1329,1407,1162,1187]" pageId="9" pageNumber="460">C. Liu.</collectorName>
</materialsCitation>
— 3 males (13.0× 
<quantity id="783B07BEFFADFF8FD065FB584594FB1C" box="[319,422,1199,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="9" pageNumber="460" unit="mm" value="11.3">11.3 mm</quantity>
, 12.5×11.0 mm, 10.4×9.0 mm), 
<materialsCitation id="0FABA006FFADFF8FD24FFB5847D8FB31" ID-GBIF-Occurrence="1913889774" collectingDate="2016-07-18" collectorName="J. Li." country="Japan" location="Awase" pageId="9" pageNumber="460" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD24FFB584745FB1D" box="[789,887,1198,1223]" pageId="9" pageNumber="460" type="female">1 female</specimenCount>
(10.1× 
<quantity id="783B07BEFFADFF8FD292FB594016FB1D" box="[968,1060,1198,1223]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="460" unit="mm" value="8.5">8.5 mm</quantity>
), 1 ovigerous female (11.9×10.3) (NMNS-7779-001), 
<collectingCountry id="C7D4EACBFFADFF8FD025FB2445F1FB30" box="[383,451,1235,1258]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD08AFB254607FB31" box="[464,565,1234,1259]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Island: 
<location id="BA1CFC80FFADFF8FD3CFFB2446D0FB31" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD3CFFB2446D0FB31" box="[661,738,1235,1259]" country="Japan" name="Awase" pageId="9" pageNumber="460" stateProvince="Okinawa">Awase</location>
, 
<collectingDate id="DB397573FFADFF8FD3B5FB2447B5FB31" box="[751,903,1234,1259]" pageId="9" pageNumber="460" value="2016-07-18">
<date id="CB7D8C9BFFADFF8FD3B5FB2447B5FB31" box="[751,903,1234,1259]" pageId="9" pageNumber="460" value="2016-07-18">18 July 2016</date>
</collectingDate>
, J.- 
<collectorName id="1236CF8DFFADFF8FD2EAFB2447D8FB31" box="[944,1002,1234,1259]" pageId="9" pageNumber="460">J. Li.</collectorName>
</materialsCitation>
— 
<materialsCitation id="0FABA006FFADFF8FD553FB254688FAE9" ID-GBIF-Occurrence="1913889764" collectingDate="2016-07-19" collectorName="J. Li." country="Japan" location="Awase" pageId="9" pageNumber="460" specimenCount="5" specimenCount-male="5" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD553FB254052FB31" box="[1033,1120,1234,1259]" pageId="9" pageNumber="460" type="male">5 males</specimenCount>
(17.0× 
<quantity id="783B07BEFFADFF8FD5EEFB25412EFB30" box="[1204,1308,1234,1259]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="9" pageNumber="460" unit="mm" value="15.1">15.1 mm</quantity>
, 16.9× 
<quantity id="783B07BEFFADFF8FD436FB2544F7FAD4" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="9" pageNumber="460" unit="mm" value="15.1">15.1 mm</quantity>
, 15.9× 
<quantity id="783B07BEFFADFF8FD04EFB01454FFAD4" box="[276,381,1270,1295]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="9" pageNumber="460" unit="mm" value="14.3">14.3 mm</quantity>
, 15.5× 
<quantity id="783B07BEFFADFF8FD096FB014607FAD4" box="[460,565,1270,1295]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="9" pageNumber="460" unit="mm" value="13.7">13.7 mm</quantity>
, 13.4×12.0 mm), 1 ovigerous female (11.8×9.9) (NMNS-7779-003), 
<collectingCountry id="C7D4EACBFFADFF8FD40FFB0041ABFAD4" box="[1365,1433,1271,1294]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD1CDFAED44CEFAE9" box="[151,252,1306,1331]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Island: 
<location id="BA1CFC80FFADFF8FD004FAEC4599FAE9" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD004FAEC4599FAE9" box="[350,427,1307,1331]" country="Japan" name="Awase" pageId="9" pageNumber="460" stateProvince="Okinawa">Awase</location>
, 
<collectingDate id="DB397573FFADFF8FD0E3FAEC4667FAE9" box="[441,597,1306,1331]" pageId="9" pageNumber="460" value="2016-07-19">
<date id="CB7D8C9BFFADFF8FD0E3FAEC4667FAE9" box="[441,597,1306,1331]" pageId="9" pageNumber="460" value="2016-07-19">19 July 2016</date>
</collectingDate>
, J.- 
<collectorName id="1236CF8DFFADFF8FD325FAEC4688FAE9" box="[639,698,1306,1331]" pageId="9" pageNumber="460">J. Li.</collectorName>
</materialsCitation>
— 
<materialsCitation id="0FABA006FFADFF8FD382FAED4754FA8D" ID-GBIF-Occurrence="1913889760" collectingDate="2016-07-20" collectorName="J. Li." country="Japan" location="Yagaji Island" pageId="9" pageNumber="460" specimenCount="2" specimenCount-male="2" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD382FAED4700FAE9" box="[728,818,1306,1331]" pageId="9" pageNumber="460" type="male">2 males</specimenCount>
(16.6× 
<quantity id="783B07BEFFADFF8FD2DCFAED47C3FAE8" box="[902,1009,1306,1331]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="9" pageNumber="460" unit="mm" value="14.4">14.4 mm</quantity>
, 15.6× 
<quantity id="783B07BEFFADFF8FD518FAED4099FAE9" box="[1090,1195,1306,1331]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="9" pageNumber="460" unit="mm" value="14.1">14.1 mm</quantity>
) (NMNS-7779-004), 
<collectingCountry id="C7D4EACBFFADFF8FD1CDFAC844E9FA8C" box="[151,219,1343,1366]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD1B3FAC9457CFA8D" box="[233,334,1342,1367]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Islands: 
<location id="BA1CFC80FFADFF8FD0E1FAC94667FA8D" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD0E1FAC94667FA8D" box="[443,597,1342,1367]" country="Japan" name="Yagaji Island" pageId="9" pageNumber="460" stateProvince="Okinawa">Yagaji Island</location>
, 
<collectingDate id="DB397573FFADFF8FD33EFAC84732FA8D" box="[612,768,1342,1367]" pageId="9" pageNumber="460" value="2016-07-20">
<date id="CB7D8C9BFFADFF8FD33EFAC84732FA8D" box="[612,768,1342,1367]" pageId="9" pageNumber="460" value="2016-07-20">20 July 2016</date>
</collectingDate>
, J.- 
<collectorName id="1236CF8DFFADFF8FD273FAC84754FA8D" box="[809,870,1342,1367]" pageId="9" pageNumber="460">J. Li.</collectorName>
</materialsCitation>
— 
<materialsCitation id="0FABA006FFADFF8FD2DDFAC94765FAA1" ID-GBIF-Occurrence="1913889775" collectingDate="2010-04-13" collectorName="T. Naruse." country="Japan" location="Oura River" municipality="Nago" pageId="9" pageNumber="460" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD2DDFAC947D5FA8D" box="[903,999,1342,1367]" pageId="9" pageNumber="460" type="female">1 female</specimenCount>
(11.7×9.7) (RUMF-ZC-2898), 
<collectingCountry id="C7D4EACBFFADFF8FD40FFAC841ABFA8C" box="[1365,1433,1343,1366]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD1CDFA9544CEFAA1" box="[151,252,1378,1403]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Island: 
<collectingMunicipality id="5F183021FFADFF8FD003FA9445A8FAA1" box="[345,410,1379,1403]" pageId="9" pageNumber="460">Nago</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD0FCFA944614FAA1" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD0FCFA944614FAA1" box="[422,550,1378,1403]" country="Japan" municipality="Nago" name="Oura River" pageId="9" pageNumber="460" stateProvince="Okinawa">Oura River</location>
, 
<collectingDate id="DB397573FFADFF8FD369FA9446E4FAA1" box="[563,726,1378,1403]" pageId="9" pageNumber="460" value="2010-04-13">
<date id="CB7D8C9BFFADFF8FD369FA9446E4FAA1" box="[563,726,1378,1403]" pageId="9" pageNumber="460" value="2010-04-13">13 April 2010</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD3B8FA944765FAA1" box="[738,855,1379,1403]" pageId="9" pageNumber="460">T. Naruse.</collectorName>
</materialsCitation>
— 
<materialsCitation id="0FABA006FFADFF8FD22EFA94403FFA45" ID-GBIF-Occurrence="1913889765" collectingDate="2010-06-04" collectorName="T. Naruse." country="Japan" location="Oura River" municipality="Nago" pageId="9" pageNumber="460" specimenCount="2" specimenCount-male="2" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD22EFA9447F8FAA1" box="[884,970,1378,1403]" pageId="9" pageNumber="460" type="male">2 males</specimenCount>
(13.4× 
<quantity id="783B07BEFFADFF8FD546FA9440B6FAA0" box="[1052,1156,1379,1403]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="9" pageNumber="460" unit="mm" value="11.4">11.4 mm</quantity>
, 10.6× 
<quantity id="783B07BEFFADFF8FD58BFA954119FAA1" box="[1233,1323,1378,1403]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="9" pageNumber="460" unit="mm" value="8.9">8.9 mm</quantity>
) (RUMF- ZC-2899), 
<collectingCountry id="C7D4EACBFFADFF8FD04EFA70456AFA44" box="[276,344,1415,1438]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD03EFA7145FBFA45" box="[356,457,1414,1439]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Island: 
<collectingMunicipality id="5F183021FFADFF8FD37FFA704654FA45" box="[549,614,1415,1439]" pageId="9" pageNumber="460">Nago</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD32BFA7046C3FA45" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD32BFA7046C3FA45" box="[625,753,1414,1439]" country="Japan" municipality="Nago" name="Oura River" pageId="9" pageNumber="460" stateProvince="Okinawa">Oura River</location>
, 
<collectingDate id="DB397573FFADFF8FD3A7FA7047B8FA45" box="[765,906,1415,1439]" pageId="9" pageNumber="460" value="2010-06-04">
<date id="CB7D8C9BFFADFF8FD3A7FA7047B8FA45" box="[765,906,1415,1439]" pageId="9" pageNumber="460" value="2010-06-04">4 June 2010</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD2CCFA70403FFA45" box="[918,1037,1415,1439]" pageId="9" pageNumber="460">T. Naruse.</collectorName>
</materialsCitation>
— 
<materialsCitation id="0FABA006FFADFF8FD571FA70407CFA19" ID-GBIF-Occurrence="1913889763" collectingDate="2010-06-09" collectorName="T. Naruse." country="Japan" location="Gabu" municipality="Yagaji Island" pageId="9" pageNumber="460" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD571FA704040FA45" box="[1067,1138,1414,1439]" pageId="9" pageNumber="460" type="male">1 male</specimenCount>
(14.8× 
<quantity id="783B07BEFFADFF8FD59EFA714118FA45" box="[1220,1322,1414,1439]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="460" unit="mm" value="12.5">12.5 mm</quantity>
) (RUMF- ZC-2900), 
<collectingCountry id="C7D4EACBFFADFF8FD04DFA5C4569FA18" box="[279,347,1451,1474]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD033FA5D45FCFA19" box="[361,462,1450,1475]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Islands: 
<collectingMunicipality id="5F183021FFADFF8FD361FA5D46E5FA19" box="[571,727,1450,1475]" pageId="9" pageNumber="460">Yagaji Island</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD3BFFA5D4717FA19" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD3BFFA5D4717FA19" box="[741,805,1450,1475]" country="Japan" municipality="Yagaji Island" name="Gabu" pageId="9" pageNumber="460" stateProvince="Okinawa">Gabu</location>
, 
<collectingDate id="DB397573FFADFF8FD269FA5C47F7FA19" box="[819,965,1451,1475]" pageId="9" pageNumber="460" value="2010-06-09">
<date id="CB7D8C9BFFADFF8FD269FA5C47F7FA19" box="[819,965,1451,1475]" pageId="9" pageNumber="460" value="2010-06-09">9 June 2010</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD28EFA5C407CFA19" box="[980,1102,1451,1475]" pageId="9" pageNumber="460">T. Naruse.</collectorName>
</materialsCitation>
—1 male (12.3×10.0 mm), 
<materialsCitation id="0FABA006FFADFF8FD4D4FA5C44C2F9D1" ID-GBIF-Occurrence="1913889771" collectingDate="2014-02-08" collectorName="T. Naruse." country="Japan" location="Okukubi River" municipality="Kin" pageId="9" pageNumber="460" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD4D4FA5C44D1FA3D" pageId="9" pageNumber="460" type="female">1 female</specimenCount>
(10.8× 
<quantity id="783B07BEFFADFF8FD063FA3845AAFA3D" box="[313,408,1487,1511]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="9" pageNumber="460" unit="mm" value="8.3">8.3 mm</quantity>
) (RUMF-ZC-2901), 
<collectingCountry id="C7D4EACBFFADFF8FD3CAFA3846E6FA3C" box="[656,724,1487,1510]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD3BEFA39477BFA3D" box="[740,841,1486,1511]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Island: 
<collectingMunicipality id="5F183021FFADFF8FD2F5FA3947EFFA3D" box="[943,989,1486,1511]" pageId="9" pageNumber="460">Kin</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD2B4FA3940AFFA3D" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD2B4FA3940AFFA3D" box="[1006,1181,1486,1511]" country="Japan" municipality="Kin" name="Okukubi River" pageId="9" pageNumber="460" stateProvince="Okinawa">Okukubi River</location>
, 
<collectingDate id="DB397573FFADFF8FD5F4FA384147FA3D" box="[1198,1397,1486,1511]" pageId="9" pageNumber="460" value="2014-02-08">
<date id="CB7D8C9BFFADFF8FD5F4FA384147FA3D" box="[1198,1397,1486,1511]" pageId="9" pageNumber="460" value="2014-02-08">8 February 2014</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD4DCFA3844C2F9D1" pageId="9" pageNumber="460">T. Naruse.</collectorName>
</materialsCitation>
— 
<materialsCitation id="0FABA006FFADFF8FD054FA0444C2F9F5" ID-GBIF-Occurrence="1913889759" collectingDate="2014-02-18" collectorName="T. Naruse." country="Japan" location="Kawamitsu" municipality="Miyako Island" pageId="9" pageNumber="460" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD054FA04456AF9D1" box="[270,344,1522,1547]" pageId="9" pageNumber="460" type="male">1 male</specimenCount>
(15.9× 
<quantity id="783B07BEFFADFF8FD0F6FA054624F9D1" box="[428,534,1522,1547]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="9" pageNumber="460" unit="mm" value="13.9">13.9 mm</quantity>
) (RUMF-ZC-2902), 
<collectingCountry id="C7D4EACBFFADFF8FD250FA04477CF9D0" box="[778,846,1523,1546]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingMunicipality id="5F183021FFADFF8FD206FA054035F9D1" box="[860,1031,1522,1547]" pageId="9" pageNumber="460">Miyako Island</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD54FFA0540A5F9D1" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD54FFA0540A5F9D1" box="[1045,1175,1522,1547]" country="Japan" municipality="Miyako Island" name="Kawamitsu" pageId="9" pageNumber="460">Kawamitsu</location>
, 
<collectingDate id="DB397573FFADFF8FD5FCFA044145F9D1" box="[1190,1399,1522,1547]" pageId="9" pageNumber="460" value="2014-02-18">
<date id="CB7D8C9BFFADFF8FD5FCFA044145F9D1" box="[1190,1399,1522,1547]" pageId="9" pageNumber="460" value="2014-02-18">18 February 2014</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD4DCFA0444C2F9F5" pageId="9" pageNumber="460">T. Naruse.</collectorName>
</materialsCitation>
— 
<materialsCitation id="0FABA006FFADFF8FD054F9E044DFF989" ID-GBIF-Occurrence="1913889758" collectingDate="2010-03-09" collectorName="Naruse." country="Japan" location="Gabu" municipality="Yagaji Island" pageId="9" pageNumber="460" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD054F9E0456AF9F5" box="[270,344,1558,1583]" pageId="9" pageNumber="460" type="male">1 male</specimenCount>
(16.9×14.5) (RUMF-ZC-2903), 
<collectingCountry id="C7D4EACBFFADFF8FD38FF9E0472BF9F4" box="[725,793,1559,1582]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFADFF8FD27DF9E147BEF9F5" box="[807,908,1558,1583]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Islands: 
<collectingMunicipality id="5F183021FFADFF8FD2A3F9E140A6F9F5" box="[1017,1172,1558,1583]" pageId="9" pageNumber="460">Yagaji Island</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD5F8F9E140D3F9F5" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD5F8F9E140D3F9F5" box="[1186,1249,1558,1583]" country="Japan" municipality="Yagaji Island" name="Gabu" pageId="9" pageNumber="460" stateProvince="Okinawa">Gabu</location>
, 
<collectingDate id="DB397573FFADFF8FD5ABF9E041AAF9F5" box="[1265,1432,1558,1583]" pageId="9" pageNumber="460" value="2010-03-09">
<date id="CB7D8C9BFFADFF8FD5ABF9E041AAF9F5" box="[1265,1432,1558,1583]" pageId="9" pageNumber="460" value="2010-03-09">9 March 2010</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD1CDF9CC44DFF989" box="[151,237,1595,1619]" pageId="9" pageNumber="460">Naruse.</collectorName>
</materialsCitation>
—2 males (17.4×14.7, 15.5×13.0 mm), 
<materialsCitation id="0FABA006FFADFF8FD39EF9CC4778F9AD" ID-GBIF-Occurrence="1913889767" collectingDate="2014-05-04" collectorName="Naruse." country="Japan" county="Yaeyama Islands" location="Utara River" municipality="Iriomote Island" pageId="9" pageNumber="460" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD39EF9CC4718F989" box="[708,810,1594,1619]" pageId="9" pageNumber="460" type="female">1 female</specimenCount>
(16.5× 
<quantity id="783B07BEFFADFF8FD2DAF9CD47D9F989" box="[896,1003,1594,1619]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="9" pageNumber="460" unit="mm" value="14.3">14.3 mm</quantity>
) (RUMF-ZC-2930), 
<collectingCountry id="C7D4EACBFFADFF8FD5B8F9CC4114F988" box="[1250,1318,1595,1618]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
: 
<collectingCounty id="561DD2D7FFADFF8FD46FF9CC44D9F9AD" pageId="9" pageNumber="460">Yaeyama Islands</collectingCounty>
: 
<collectingMunicipality id="5F183021FFADFF8FD1A3F9A9459FF9AD" box="[249,429,1630,1655]" pageId="9" pageNumber="460">Iriomote Island</collectingMunicipality>
: 
<location id="BA1CFC80FFADFF8FD0E1F9A84676F9AD" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD0E1F9A84676F9AD" box="[443,580,1630,1655]" country="Japan" county="Yaeyama Islands" municipality="Iriomote Island" name="Utara River" pageId="9" pageNumber="460">Utara River</location>
, 
<collectingDate id="DB397573FFADFF8FD308F9A846D6F9AD" box="[594,740,1631,1655]" pageId="9" pageNumber="460" value="2014-05-04">
<date id="CB7D8C9BFFADFF8FD308F9A846D6F9AD" box="[594,740,1631,1655]" pageId="9" pageNumber="460" value="2014-05-04">4 May 2014</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD3A9F9A84778F9AD" box="[755,842,1631,1655]" pageId="9" pageNumber="460">Naruse.</collectorName>
</materialsCitation>
— 
<materialsCitation id="0FABA006FFADFF8FD23DF9A84791F940" ID-GBIF-Occurrence="1913889777" country="Japan" location="Japan" pageId="9" pageNumber="460" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD23DF9A8478DF9AD" box="[871,959,1630,1655]" pageId="9" pageNumber="460" type="male">3 males</specimenCount>
(1.7× 
<quantity id="783B07BEFFADFF8FD55EF9A8405DF9AC" box="[1028,1135,1631,1655]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="9" pageNumber="460" unit="mm" value="12.4">12.4 mm</quantity>
, 13.4×12.0 mm, 10.9× 
<quantity id="783B07BEFFADFF8FD423F9A944F5F941" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="9" pageNumber="460" unit="mm" value="9.5">9.5 mm</quantity>
), 1 ovigerous female (15.6× 
<quantity id="783B07BEFFADFF8FD35DF975465DF941" box="[519,623,1666,1691]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="9" pageNumber="460" unit="mm" value="13.8">13.8 mm</quantity>
) (RUMF-ZC-4711), 
<location id="BA1CFC80FFADFF8FD205F9744791F940" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD205F9744791F940" box="[863,931,1667,1690]" country="Japan" name="Japan" pageId="9" pageNumber="460">
<collectingCountry id="C7D4EACBFFADFF8FD205F9744791F940" box="[863,931,1667,1690]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
</location>
</materialsCitation>
: 
<materialsCitation id="0FABA006FFADFF8FD2F5F975402FF964" ID-GBIF-Occurrence="1913889768" collectingDate="2016-07-17" collectorName="J. Li." country="Japan" location="Awase" pageId="9" pageNumber="460" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
<collectingRegion id="7D0764B9FFADFF8FD2F5F9754026F941" box="[943,1044,1666,1691]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Island: 
<location id="BA1CFC80FFADFF8FD528F974408DF941" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD528F974408DF941" box="[1138,1215,1667,1691]" country="Japan" name="Awase" pageId="9" pageNumber="460" stateProvince="Okinawa">Awase</location>
, 
<collectingDate id="DB397573FFADFF8FD591F9744150F941" box="[1227,1378,1666,1691]" pageId="9" pageNumber="460" value="2016-07-17">
<date id="CB7D8C9BFFADFF8FD591F9744150F941" box="[1227,1378,1666,1691]" pageId="9" pageNumber="460" value="2016-07-17">17 July 2016</date>
</collectingDate>
, J.- 
<collectorName id="1236CF8DFFADFF8FD4D0F9744485F965" pageId="9" pageNumber="460">J. Li.</collectorName>
—1 ovigerous female (20.2×17.5) (NMNS-7779-002) (leg variation), 
<collectingCountry id="C7D4EACBFFADFF8FD283F950402FF964" box="[985,1053,1703,1726]" name="Japan" pageId="9" pageNumber="460">Japan</collectingCountry>
</materialsCitation>
: 
<materialsCitation id="0FABA006FFADFF8FD570F95145D6F939" ID-GBIF-Occurrence="1913889780" collectingDate="2016-07-18" collectorName="J. Li. Other" country="Japan" location="Awase" pageId="9" pageNumber="460" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
<collectingRegion id="7D0764B9FFADFF8FD570F95140BDF965" box="[1066,1167,1702,1727]" country="Japan" name="Okinawa" pageId="9" pageNumber="460">Okinawa</collectingRegion>
Island: 
<location id="BA1CFC80FFADFF8FD5B4F9504109F965" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD5B4F9504109F965" box="[1262,1339,1703,1727]" country="Japan" name="Awase" pageId="9" pageNumber="460" stateProvince="Okinawa">Awase</location>
, 
<collectingDate id="DB397573FFADFF8FD412F95044E0F939" pageId="9" pageNumber="460" value="2016-07-18">
<date id="CB7D8C9BFFADFF8FD412F95044E0F939" pageId="9" pageNumber="460" value="2016-07-18">18 July 2016</date>
</collectingDate>
, J.- 
<collectorName id="1236CF8DFFADFF8FD1A3F93C454EF939" box="[249,380,1738,1763]" pageId="9" pageNumber="460">J. Li. Other</collectorName>
material
</materialsCitation>
: 
<materialsCitation id="0FABA006FFADFF8FD0B5F93D4606F8F1" ID-GBIF-Occurrence="1913889782" collectingDate="2008-09" collectorName="Liu." country="Guam" location="Guam" pageId="9" pageNumber="460" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="A9C561D2FFADFF8FD0B5F93D4675F939" box="[495,583,1738,1763]" pageId="9" pageNumber="460" type="male">5 males</specimenCount>
(19.1× 
<quantity id="783B07BEFFADFF8FD3C2F93D4733F938" box="[664,769,1738,1763]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="9" pageNumber="460" unit="mm" value="16.6">16.6 mm</quantity>
, 14.9× 
<quantity id="783B07BEFFADFF8FD215F93C4784F938" box="[847,950,1739,1763]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="9" pageNumber="460" unit="mm" value="13.3">13.3 mm</quantity>
, 13.7× 
<quantity id="783B07BEFFADFF8FD559F93D4058F938" box="[1027,1130,1738,1763]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="9" pageNumber="460" unit="mm" value="11.5">11.5 mm</quantity>
, 12.9×11.0 mm, 12.4× 
<quantity id="783B07BEFFADFF8FD431F93C44F5F8DD" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="9" pageNumber="460" unit="mm" value="10.4">10.4 mm</quantity>
), 4 ovigerous females (18.1× 
<quantity id="783B07BEFFADFF8FD34DF91946B0F8DC" box="[535,642,1774,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="9" pageNumber="460" unit="mm" value="15.4">15.4 mm</quantity>
, 16.8× 
<quantity id="783B07BEFFADFF8FD38BF9194709F8DC" box="[721,827,1774,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="9" pageNumber="460" unit="mm" value="14.8">14.8 mm</quantity>
, 16.5× 
<quantity id="783B07BEFFADFF8FD2D3F91947C1F8DC" box="[905,1011,1774,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="460" unit="mm" value="14.2">14.2 mm</quantity>
, 16.2× 
<quantity id="783B07BEFFADFF8FD51EF9194099F8DD" box="[1092,1195,1774,1799]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="9" pageNumber="460" unit="mm" value="14.1">14.1 mm</quantity>
) (NMNS-7779-006), 
<collectingCountry id="C7D4EACBFFADFF8FD1CDF8E444ECF8F1" box="[151,222,1811,1835]" name="Guam" pageId="9" pageNumber="460">Guam</collectingCountry>
, 
<collectingDate id="DB397573FFADFF8FD1B7F8E54599F8F1" box="[237,427,1810,1835]" pageId="9" pageNumber="460" value="2008-09">
<date id="CB7D8C9BFFADFF8FD1B7F8E54599F8F1" box="[237,427,1810,1835]" pageId="9" pageNumber="460" value="2008-09">September 2008</date>
</collectingDate>
, H.-C. 
<collectorName id="1236CF8DFFADFF8FD35EF8E54606F8F1" box="[516,564,1810,1835]" pageId="9" pageNumber="460">Liu.</collectorName>
</materialsCitation>
—1 male (13.3× 
<quantity id="783B07BEFFADFF8FD3A8F8E5476EF8F1" box="[754,860,1810,1835]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="9" pageNumber="460" unit="mm" value="11.5">11.5 mm</quantity>
) (ZRC 2017.196), Vanuatu: 
<materialsCitation id="0FABA006FFADFF8FD5F7F8E4414FF8A9" ID-GBIF-Occurrence="1913889784" collectingDate="2006-09-11" collectionCode="SANTO" collectorName="P. K. L. Ng" country="Guam" county="West Baldwin Cove" latitude="-15.581667" location="Expedition" longLatPrecision="128" longitude="167.03" municipality="Nasouli River" pageId="9" pageNumber="460" specimenCode="SANTO 2006" specimenCount="1" specimenCount-adult="1" typeStatus="paratype">
<collectingCounty id="561DD2D7FFADFF8FD5F7F8E441AAF8F1" box="[1197,1432,1810,1835]" pageId="9" pageNumber="460">West Baldwin Cove</collectingCounty>
, 
<collectingMunicipality id="5F183021FFADFF8FD1CDF8C1450EF895" box="[151,316,1846,1871]" pageId="9" pageNumber="460">Nasouli River</collectingMunicipality>
mouth intertidal, hard bottom, 
<location id="BA1CFC80FFADFF8FD39DF8C14770F895" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD39DF8C14770F895" box="[711,834,1846,1871]" country="Guam" county="West Baldwin Cove" latitude="-15.581667" longLatPrecision="128" longitude="167.03" municipality="Nasouli River" name="Expedition" pageId="9" pageNumber="460">Expédition</location>
<specimenCode id="EF650220FFADFF8FD20BF8C047CBF895" box="[849,1017,1846,1871]" collectionCode="SANTO" pageId="9" pageNumber="460">SANTO 2006</specimenCode>
, station VM4, intertidal area near mangroves, 
<geoCoordinate id="DAF7CC9CFFADFF8FD07FF8AD45A0F8A9" box="[293,402,1882,1907]" direction="south" orientation="latitude" pageId="9" pageNumber="460" precision="92" value="-15.581667">15°34.9'S</geoCoordinate>
<geoCoordinate id="DAF7CC9CFFADFF8FD0C4F8AD462CF8A9" box="[414,542,1882,1907]" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="92" value="167.03">167°01.8'E</geoCoordinate>
, 
<collectingDate id="DB397573FFADFF8FD374F8AC4727F8A9" box="[558,789,1882,1907]" pageId="9" pageNumber="460" value="2006-09-11">
<date id="CB7D8C9BFFADFF8FD374F8AC4727F8A9" box="[558,789,1882,1907]" pageId="9" pageNumber="460" value="2006-09-11">11 September 2006</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD27FF8AC47A8F8A9" box="[805,922,1883,1907]" pageId="9" pageNumber="460">P.K.L. Ng</collectorName>
<emphasis id="8DB77649FFADFF8FD2FDF8AB47ECF8A9" box="[935,990,1882,1907]" italics="true" pageId="9" pageNumber="460">et al</emphasis>
..— 
<specimenCount id="A9C561D2FFADFF8FD556F8AC4068F8A9" box="[1036,1114,1882,1907]" pageId="9" pageNumber="460" type="adult">1 adult</specimenCount>
female (13.2× 
<quantity id="783B07BEFFADFF8FD452F8AD4144F8A9" box="[1288,1398,1882,1907]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="9" pageNumber="460" unit="mm" value="11.5">11.5 mm</quantity>
)
</materialsCitation>
, 2 juvenile females (8.9× 
<quantity id="783B07BEFFADFF8FD0CEF88945DDF84C" box="[404,495,1918,1943]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="460" unit="mm" value="7.6">7.6 mm</quantity>
, 8.6× 
<quantity id="783B07BEFFADFF8FD374F88946BAF84D" box="[558,648,1918,1943]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="9" pageNumber="460" unit="mm" value="7.2">7.2 mm</quantity>
) (ZRC 2017.197), Vanuatu: 
<materialsCitation id="0FABA006FFADFF8FD295F8894639F804" ID-GBIF-Occurrence="1913889781" collectingDate="2006-09-13" collectionCode="SANTO" collectorName="P. K. L. Ng" country="Guam" latitude="-15.596666" location="Expedition" longLatPrecision="128" longitude="167.10333" municipality="Belmoul" pageId="9" pageNumber="460" specimenCode="SANTO 2006" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="5F183021FFADFF8FD295F889401DF84D" box="[975,1071,1918,1943]" pageId="9" pageNumber="460">Belmoul</collectingMunicipality>
lagoon channel, intertidal, sand and muddy sand, 
<location id="BA1CFC80FFADFF8FD006F85545E5F861" LSID="urn:lsid:plazi:treatment:376A1B4DFFADFF8BD1CDFD444536FC87:BA1CFC80FFADFF8FD006F85545E5F861" box="[348,471,1954,1979]" country="Guam" latitude="-15.596666" longLatPrecision="128" longitude="167.10333" municipality="Belmoul" name="Expedition" pageId="9" pageNumber="460">Expédition</location>
<specimenCode id="EF650220FFADFF8FD084F854464EF861" box="[478,636,1954,1979]" collectionCode="SANTO" pageId="9" pageNumber="460">SANTO 2006</specimenCode>
, station VM9, intertidal area near mangroves, 
<geoCoordinate id="DAF7CC9CFFADFF8FD5D9F85540C2F861" box="[1155,1264,1954,1979]" direction="south" orientation="latitude" pageId="9" pageNumber="460" precision="92" value="-15.596666">15°35.8'S</geoCoordinate>
<geoCoordinate id="DAF7CC9CFFADFF8FD5ACF8554147F861" box="[1270,1397,1954,1979]" direction="east" orientation="longitude" pageId="9" pageNumber="460" precision="92" value="167.10333">167°06.2'E</geoCoordinate>
, 
<collectingDate id="DB397573FFADFF8FD4DAF8544562F805" pageId="9" pageNumber="460" value="2006-09-13">
<date id="CB7D8C9BFFADFF8FD4DAF8544562F805" pageId="9" pageNumber="460" value="2006-09-13">13 September 2006</date>
</collectingDate>
, 
<collectorName id="1236CF8DFFADFF8FD006F83045F9F805" box="[348,459,1991,2015]" pageId="9" pageNumber="460">P.K.L. Ng</collectorName>
<emphasis id="8DB77649FFADFF8FD089F8304636F805" box="[467,516,1990,2015]" italics="true" pageId="9" pageNumber="460">et al</emphasis>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="EBBCFAD3FFAEFF8CD1CDF877414CF828" httpUri="https://zenodo.org/record/1440717/files/figure.png" pageId="10" pageNumber="461" startId="10.[151,250,1920,1942]" targetBox="[192,1407,193,1911]" targetPageId="10">
<paragraph id="BF7CAA5BFFAEFF8CD1CDF877414CF828" blockId="10.[151,1436,1920,2034]" pageId="10" pageNumber="461">
<emphasis id="8DB77649FFAEFF8CD1CDF877453FF84F" bold="true" box="[151,269,1920,1942]" pageId="10" pageNumber="461">FIGURE 4</emphasis>
. 
<taxonomicName id="78C3D1D8FFAEFF8CD043F87645D0F84C" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[281,482,1920,1942]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFAEFF8CD043F87645D0F84C" box="[281,482,1920,1942]" italics="true" pageId="10" pageNumber="461">Parasesarma gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFAEFF8CD0B2F8774610F84F" box="[488,546,1920,1941]" pageId="10" pageNumber="461" rank="species">
<emphasis id="8DB77649FFAEFF8CD0B2F8774610F84F" bold="true" box="[488,546,1920,1941]" pageId="10" pageNumber="461">n. sp.</emphasis>
</taxonomicNameLabel>
. A–D, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; E, F, paratype female (15.3×13.0 mm) (ASIZ 73381), Iriomote Island, Japan; G, H, female (20.2×17.5 mm) (NMNS-7779-002), Okinawa Island, Japan, showing abnormal right second ambulatory leg; I, J, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan, showing asymmetrical chelae. A, E, G, I, dorsal views; B, F, H, J, ventral views; C, D, frontal views.
</paragraph>
</caption>
<caption id="EBBCFAD3FFAFFF8DD1CDFA974051FA69" httpUri="https://zenodo.org/record/1440719/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,250,1376,1398]" targetBox="[186,1355,232,1354]" targetPageId="11">
<paragraph id="BF7CAA5BFFAFFF8DD1CDFA974051FA69" blockId="11.[151,1436,1376,1459]" pageId="11" pageNumber="462">
<emphasis id="8DB77649FFAFFF8DD1CDFA97453FFAAF" bold="true" box="[151,269,1376,1398]" pageId="11" pageNumber="462">FIGURE 5</emphasis>
. 
<taxonomicName id="78C3D1D8FFAFFF8DD040FA9645D0FAAC" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[282,482,1376,1398]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFAFFF8DD040FA9645D0FAAC" box="[282,482,1376,1398]" italics="true" pageId="11" pageNumber="462">Parasesarma gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFAFFF8DD0B3FA974610FAAF" box="[489,546,1376,1397]" pageId="11" pageNumber="462" rank="species">
<emphasis id="8DB77649FFAFFF8DD0B3FA974610FAAF" bold="true" box="[489,546,1376,1397]" pageId="11" pageNumber="462">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan. A, dorsal view of carapace; B, lateral view of pectinated cristae on palm of left chela; C, dorsal view of tip of left cheliped; D–G, left G1. D, F, dorsal (sternal) views; E, G, ventral (pleonal) views; D, E, distal part of G1. Scale bars: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="F7D9F9D0FFAFFF8AD19DFA2844D8FB89" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462" type="diagnosis">
<paragraph id="BF7CAA5BFFAFFF8DD19DFA2844E4F82A" blockId="11.[151,1437,1503,2032]" pageId="11" pageNumber="462">
<emphasis id="8DB77649FFAFFF8DD19DFA284573FA22" bold="true" box="[199,321,1503,1528]" pageId="11" pageNumber="462">Diagnosis.</emphasis>
Carapace (
<figureCitation id="27F8B6DEFFAFFF8DD09FFA2846B8FA2D" box="[453,650,1503,1528]" captionStart="FIGURE 4" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[192,1407,193,1911]" captionTargetId="figure@10.[192,1407,193,1911]" captionTargetPageId="10" captionText="FIGURE 4. Parasesarma gecko n. sp.. A–D, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; E, F, paratype female (15.3×13.0 mm) (ASIZ 73381), Iriomote Island, Japan; G, H, female (20.2×17.5 mm) (NMNS-7779-002), Okinawa Island, Japan, showing abnormal right second ambulatory leg; I, J, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan, showing asymmetrical chelae. A, E, G, I, dorsal views; B, F, H, J, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440717/files/figure.png" pageId="11" pageNumber="462">Figs. 4A, E, G, I</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD3C0FA28468DFA22" box="[666,703,1503,1528]" captionStart="FIGURE 5" captionStartId="11.[151,250,1376,1398]" captionTargetBox="[186,1355,232,1354]" captionTargetId="figure@11.[152,1435,193,1354]" captionTargetPageId="11" captionText="FIGURE 5. Parasesarma gecko n. sp., holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan. A, dorsal view of carapace; B, lateral view of pectinated cristae on palm of left chela; C, dorsal view of tip of left cheliped; D–G, left G1. D, F, dorsal (sternal) views; E, G, ventral (pleonal) views; D, E, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440719/files/figure.png" pageId="11" pageNumber="462">5A</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD394FA1746CDFA22" box="[718,767,1504,1528]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="11" pageNumber="462">22A</figureCitation>
) squarish in general outline, 1.2 times broader than long; regions well defined, separated by shallow grooves; postfrontal region distinct, separated into 4 lobes by deep grooves; front deflexed downwards (
<figureCitation id="27F8B6DEFFAFFF8DD318F9D0469AF99A" box="[578,680,1575,1600]" captionStart="FIGURE 4" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[192,1407,193,1911]" captionTargetId="figure@10.[192,1407,193,1911]" captionTargetPageId="10" captionText="FIGURE 4. Parasesarma gecko n. sp.. A–D, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; E, F, paratype female (15.3×13.0 mm) (ASIZ 73381), Iriomote Island, Japan; G, H, female (20.2×17.5 mm) (NMNS-7779-002), Okinawa Island, Japan, showing abnormal right second ambulatory leg; I, J, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan, showing asymmetrical chelae. A, E, G, I, dorsal views; B, F, H, J, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440717/files/figure.png" pageId="11" pageNumber="462">Figs. 4C</figureCitation>
), margin slightly concave in dorsal view; lateral margin straight, subparallel along most of length before curving to join almost straight posterior carapace margin; cornea extending or just reaching tip of external orbital tooth (
<figureCitation id="27F8B6DEFFAFFF8DD3D4F998477BF95D" box="[654,841,1647,1672]" captionStart="FIGURE 4" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[192,1407,193,1911]" captionTargetId="figure@10.[192,1407,193,1911]" captionTargetPageId="10" captionText="FIGURE 4. Parasesarma gecko n. sp.. A–D, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; E, F, paratype female (15.3×13.0 mm) (ASIZ 73381), Iriomote Island, Japan; G, H, female (20.2×17.5 mm) (NMNS-7779-002), Okinawa Island, Japan, showing abnormal right second ambulatory leg; I, J, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan, showing asymmetrical chelae. A, E, G, I, dorsal views; B, F, H, J, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440717/files/figure.png" pageId="11" pageNumber="462">Figs. 4A, E, G, I</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD20CF9984748F952" box="[854,890,1647,1672]" captionStart="FIGURE 5" captionStartId="11.[151,250,1376,1398]" captionTargetBox="[186,1355,232,1354]" captionTargetId="figure@11.[152,1435,193,1354]" captionTargetPageId="11" captionText="FIGURE 5. Parasesarma gecko n. sp., holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan. A, dorsal view of carapace; B, lateral view of pectinated cristae on palm of left chela; C, dorsal view of tip of left cheliped; D–G, left G1. D, F, dorsal (sternal) views; E, G, ventral (pleonal) views; D, E, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440719/files/figure.png" pageId="11" pageNumber="462">5A</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD2DDF9874789F952" box="[903,955,1648,1672]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="11" pageNumber="462">22A</figureCitation>
). Ischium of third maxilliped with shallow median sulcus, merus with distinct submedian ridge; exopod slender, tip reaching half-length of outer margin of merus, flagellum long. Male cheliped palm with 2 (sometimes 3) transverse pectinate crests (
<date id="CB7D8C9BFFAFFF8DD5BAF94F41AEF90A" box="[1248,1436,1719,1744]" pageId="11" pageNumber="462" value="2010-11-13" valueMax="2010-11-03" valueMin="2010-11-13">11–13 and 3–10</date>
corneous teeth, respectively) (
<figureCitation id="27F8B6DEFFAFFF8DD0B0F92B465CF92E" box="[490,622,1756,1781]" captionStart="FIGURE 5" captionStartId="11.[151,250,1376,1398]" captionTargetBox="[186,1355,232,1354]" captionTargetId="figure@11.[152,1435,193,1354]" captionTargetPageId="11" captionText="FIGURE 5. Parasesarma gecko n. sp., holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan. A, dorsal view of carapace; B, lateral view of pectinated cristae on palm of left chela; C, dorsal view of tip of left cheliped; D–G, left G1. D, F, dorsal (sternal) views; E, G, ventral (pleonal) views; D, E, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440719/files/figure.png" pageId="11" pageNumber="462">Figs. 5B, C</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD321F92B468DF92F" box="[635,703,1756,1781]" captionStart="FIGURE 6" captionStartId="12.[151,250,815,837]" captionTargetBox="[175,1397,233,748]" captionTargetId="figure@12.[151,1435,193,794]" captionTargetPageId="12" captionText="FIGURE 6. Parasesarma gecko n. sp., left chela. A, D, paratype male (14.4×12.6 mm) (ZRC 2008.1252), Miyako Island, Japan; B, E, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan; C, F, paratype male (14.5× 12.3 mm) (ZRC 2008.0501), Guam. A–C, dorsal views of pectinated cristae on palm, arrow points to direction of dactylar finger; D–F, lateral views of pectinated cristae on palm. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440721/files/figure.png" pageId="11" pageNumber="462">6A–C</figureCitation>
) on upper surface; upper surface of dactylus with 10–15 (mostly 11) symmetrical, obliquely elongate dactylar tubercles, proximal 1–5 tubercles steep, sharp, other tubercles large but becoming lower distally, distalmost tubercle indistinct (
<figureCitation id="27F8B6DEFFAFFF8DD26BF8D347A6F8E7" box="[817,916,1828,1853]" captionStart="FIGURE 5" captionStartId="11.[151,250,1376,1398]" captionTargetBox="[186,1355,232,1354]" captionTargetId="figure@11.[152,1435,193,1354]" captionTargetPageId="11" captionText="FIGURE 5. Parasesarma gecko n. sp., holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan. A, dorsal view of carapace; B, lateral view of pectinated cristae on palm of left chela; C, dorsal view of tip of left cheliped; D–G, left G1. D, F, dorsal (sternal) views; E, G, ventral (pleonal) views; D, E, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440719/files/figure.png" pageId="11" pageNumber="462">Figs. 5C</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD2FAF8D347D2F8E7" box="[928,992,1828,1853]" captionStart="FIGURE 6" captionStartId="12.[151,250,815,837]" captionTargetBox="[175,1397,233,748]" captionTargetId="figure@12.[151,1435,193,794]" captionTargetPageId="12" captionText="FIGURE 6. Parasesarma gecko n. sp., left chela. A, D, paratype male (14.4×12.6 mm) (ZRC 2008.1252), Miyako Island, Japan; B, E, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan; C, F, paratype male (14.5× 12.3 mm) (ZRC 2008.0501), Guam. A–C, dorsal views of pectinated cristae on palm, arrow points to direction of dactylar finger; D–F, lateral views of pectinated cristae on palm. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440721/files/figure.png" pageId="11" pageNumber="462">6D–F</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD2B1F8D3402FF8E6" box="[1003,1053,1828,1852]" captionStart="FIGURE 19" captionStartId="28.[151,250,1811,1833]" captionTargetBox="[152,1435,199,1754]" captionTargetId="figure@28.[152,1435,199,1754]" captionTargetPageId="28" captionText="FIGURE 19. Outer views of chelae, holotype males. A, Parasesarma leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; C, P. macaco n. sp. (13.8×12.3 mm) (NMNS-7779- 005), Pingtung, Taiwan; D, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440747/files/figure.png" pageId="11" pageNumber="462">19B</figureCitation>
). Ambulatory legs relatively stout for this species-complex, P3 and P4 about 1.7 times carapace width; P3 and P4 coxae without dense setae; P3 merus 2.5 times as long as broad; P3 propodus 4.8 times as long as broad; P3 dactylus 0.4 times length of propodus (
<figureCitation id="27F8B6DEFFAFFF8DD1C5F878453AF872" box="[159,264,1935,1960]" captionStart="FIGURE 20" captionStartId="30.[151,250,1268,1290]" captionTargetBox="[152,1435,393,1246]" captionTargetId="figure@30.[152,1435,393,1246]" captionTargetPageId="30" captionText="FIGURE 20. First to fourth ambulatory legs (P2–P5) (from top to bottom), holotype males. A, Parasesarma gecko n. sp. (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; B, P. macaco n. sp. male (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C, P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. gracilipes n. sp. (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, P. purpureum n. sp. (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; F, P. parvulum n. sp. (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; G, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440749/files/figure.png" pageId="11" pageNumber="462">Fig. 20A</figureCitation>
). G1 relatively slender (
<figureCitation id="27F8B6DEFFAFFF8DD349F87846A4F872" box="[531,662,1935,1960]" captionStart="FIGURE 5" captionStartId="11.[151,250,1376,1398]" captionTargetBox="[186,1355,232,1354]" captionTargetId="figure@11.[152,1435,193,1354]" captionTargetPageId="11" captionText="FIGURE 5. Parasesarma gecko n. sp., holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan. A, dorsal view of carapace; B, lateral view of pectinated cristae on palm of left chela; C, dorsal view of tip of left cheliped; D–G, left G1. D, F, dorsal (sternal) views; E, G, ventral (pleonal) views; D, E, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440719/files/figure.png" pageId="11" pageNumber="462">Figs. 5D–G</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD3FBF86746E1F872" box="[673,723,1936,1960]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="11" pageNumber="462">21B</figureCitation>
), slightly bent; apical process corneous, elongated, slightly bent at angle of 45°, long, stout, ending in rounded tip (
<figureCitation id="27F8B6DEFFAFFF8DD394F8434763F817" box="[718,849,1972,1997]" captionStart="FIGURE 5" captionStartId="11.[151,250,1376,1398]" captionTargetBox="[186,1355,232,1354]" captionTargetId="figure@11.[152,1435,193,1354]" captionTargetPageId="11" captionText="FIGURE 5. Parasesarma gecko n. sp., holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan. A, dorsal view of carapace; B, lateral view of pectinated cristae on palm of left chela; C, dorsal view of tip of left cheliped; D–G, left G1. D, F, dorsal (sternal) views; E, G, ventral (pleonal) views; D, E, distal part of G1. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440719/files/figure.png" pageId="11" pageNumber="462">Figs. 5D–G</figureCitation>
, 
<figureCitation id="27F8B6DEFFAFFF8DD207F84347BDF816" box="[861,911,1972,1996]" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="11" pageNumber="462">21B</figureCitation>
). G2 short, less than quarter length of G1 (
<figureCitation id="27F8B6DEFFAFFF8DD435F84344FBF82A" captionStart="FIGURE 21" captionStartId="31.[151,250,1815,1837]" captionTargetBox="[152,1435,260,1731]" captionTargetId="figure@31.[152,1435,260,1731]" captionTargetPageId="31" captionText="FIGURE 21. Gonopods of Parasesarma leptosoma species-complex. A, P. leptosoma (17.0×15.2 mm) (ZRC 2000.1718), Kenya; B, P. gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; C, P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; D, P. kui n. sp., holotype (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; E, P. gracilipes n. sp., holotype (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; F, P. purpureum n. sp., holotype (15.9×13.7 mm) (ZRC 2012.0755a), Tioman Island, Peninsular Malaysia; G, P. parvulum n. sp., holotype (9.6×8.4 mm) (NMCR 50702), Mindoro Island, Philippines; H, P. tarantula n. sp., holotype (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440751/files/figure.png" pageId="11" pageNumber="462">Fig. 21B</figureCitation>
).
</paragraph>
<caption id="EBBCFAD3FFA8FF8AD1CDFCD846E0FC7B" httpUri="https://zenodo.org/record/1440721/files/figure.png" pageId="12" pageNumber="463" startId="12.[151,250,815,837]" targetBox="[175,1397,233,748]" targetPageId="12">
<paragraph id="BF7CAA5BFFA8FF8AD1CDFCD846E0FC7B" blockId="12.[151,1436,815,929]" pageId="12" pageNumber="463">
<emphasis id="8DB77649FFA8FF8AD1CDFCD8453DFC9E" bold="true" box="[151,271,815,837]" pageId="12" pageNumber="463">FIGURE 6</emphasis>
. 
<taxonomicName id="78C3D1D8FFA8FF8AD044FCC745DBFC9F" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[286,489,815,837]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD044FCC745DBFC9F" box="[286,489,815,837]" italics="true" pageId="12" pageNumber="463">Parasesarma gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD0A8FCD8461CFC9E" box="[498,558,815,836]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD0A8FCD8461CFC9E" bold="true" box="[498,558,815,836]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
, left chela. A, D, paratype male (14.4×12.6 mm) (ZRC 2008.1252), Miyako Island, Japan; B, E, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan; C, F, paratype male (14.5× 12.3 mm) (ZRC 2008.0501), Guam. A–C, dorsal views of pectinated cristae on palm, arrow points to direction of dactylar finger; D–F, lateral views of pectinated cristae on palm. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="BF7CAA5BFFA8FF8AD19DFC3944D8FB89" blockId="12.[151,1437,974,2007]" pageId="12" pageNumber="463">
<emphasis id="8DB77649FFA8FF8AD19DFC3945C0FC3D" bold="true" box="[199,498,974,999]" pageId="12" pageNumber="463">Morphological variation.</emphasis>
Several male specimens have asymmetrical chelipeds (
<figureCitation id="27F8B6DEFFA8FF8AD539FC3940E4FC3D" box="[1123,1238,974,999]" captionStart="FIGURE 4" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[192,1407,193,1911]" captionTargetId="figure@10.[192,1407,193,1911]" captionTargetPageId="10" captionText="FIGURE 4. Parasesarma gecko n. sp.. A–D, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; E, F, paratype female (15.3×13.0 mm) (ASIZ 73381), Iriomote Island, Japan; G, H, female (20.2×17.5 mm) (NMNS-7779-002), Okinawa Island, Japan, showing abnormal right second ambulatory leg; I, J, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan, showing asymmetrical chelae. A, E, G, I, dorsal views; B, F, H, J, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440717/files/figure.png" pageId="12" pageNumber="463">
Figs. 
<date id="CB7D8C9BFFA8FF8AD5F9FC38408EFC3D" box="[1187,1212,975,999]" pageId="12" pageNumber="463">4I</date>
, J
</figureCitation>
), probably due to regeneration of lost limbs. The dactylar tubercles of the male chela ranges from 10 to 15 (average 11) (
<figureCitation id="27F8B6DEFFA8FF8AD457FC0541A2FBD1" box="[1293,1424,1010,1035]" captionStart="FIGURE 6" captionStartId="12.[151,250,815,837]" captionTargetBox="[175,1397,233,748]" captionTargetId="figure@12.[151,1435,193,794]" captionTargetPageId="12" captionText="FIGURE 6. Parasesarma gecko n. sp., left chela. A, D, paratype male (14.4×12.6 mm) (ZRC 2008.1252), Miyako Island, Japan; B, E, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan; C, F, paratype male (14.5× 12.3 mm) (ZRC 2008.0501), Guam. A–C, dorsal views of pectinated cristae on palm, arrow points to direction of dactylar finger; D–F, lateral views of pectinated cristae on palm. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440721/files/figure.png" pageId="12" pageNumber="463">Figs. 6D–F</figureCitation>
). The corneous teeth of the proximal transverse pectinate crest on the male cheliped palm varies from 3 to 10 (
<figureCitation id="27F8B6DEFFA8FF8AD43EFBE144ECFB89" captionStart="FIGURE 6" captionStartId="12.[151,250,815,837]" captionTargetBox="[175,1397,233,748]" captionTargetId="figure@12.[151,1435,193,794]" captionTargetPageId="12" captionText="FIGURE 6. Parasesarma gecko n. sp., left chela. A, D, paratype male (14.4×12.6 mm) (ZRC 2008.1252), Miyako Island, Japan; B, E, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan; C, F, paratype male (14.5× 12.3 mm) (ZRC 2008.0501), Guam. A–C, dorsal views of pectinated cristae on palm, arrow points to direction of dactylar finger; D–F, lateral views of pectinated cristae on palm. Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1440721/files/figure.png" pageId="12" pageNumber="463">Figs. 6A–C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFA8FF8AD19DFBA9400CFB41" pageId="12" pageNumber="463" type="description">
<paragraph id="BF7CAA5BFFA8FF8AD19DFBA9400CFB41" blockId="12.[151,1437,974,2007]" pageId="12" pageNumber="463">
<emphasis id="8DB77649FFA8FF8AD19DFBA9455EFBAD" bold="true" box="[199,364,1118,1143]" pageId="12" pageNumber="463">Colour in life.</emphasis>
Carapace dark brown, mottled with darker and lighter blotches; chelipeds brownish orange, but more yellowish in large males; ambulatory legs brown (
<figureCitation id="27F8B6DEFFA8FF8AD252FB754748FB40" box="[776,890,1154,1179]" captionStart="FIGURE 22" captionStartId="32.[151,250,1490,1512]" captionTargetBox="[152,1435,193,1469]" captionTargetId="figure@32.[152,1435,193,1469]" captionTargetPageId="32" captionText="FIGURE 22. Colour in life, males, dorsal views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440753/files/figure.png" pageId="12" pageNumber="463">Figs. 22A</figureCitation>
, 
<figureCitation id="27F8B6DEFFA8FF8AD2DCFB75478BFB40" box="[902,953,1154,1178]" captionStart="FIGURE 23" captionStartId="33.[151,250,1605,1627]" captionTargetBox="[152,1435,245,1548]" captionTargetId="figure@33.[152,1435,245,1548]" captionTargetPageId="33" captionText="FIGURE 23. Colour in life, males, ventral views. A. Parasesarma gecko n. sp., holotype (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa, Japan; B. P. macaco n. sp., holotype (13.8×12.3 mm) (NMNS-7779-005), Pingtung, Taiwan; C. P. kui n. sp. (14.7×13.7 mm) (NMNS-7779-015), Pingtung, Taiwan; D, P. purpureum n. sp. (13.4×11.8 mm) (ZRC 2018.0799), Borneo, Malaysia; E, P. tarantula n. sp. (13.4×11.5 mm) (MZB Cru 4801), Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440755/files/figure.png" pageId="12" pageNumber="463">23A</figureCitation>
, 
<figureCitation id="27F8B6DEFFA8FF8AD29EFB7547C7FB40" box="[964,1013,1154,1178]" captionStart="FIGURE 24" captionStartId="34.[151,250,1531,1553]" captionTargetBox="[151,1436,218,1490]" captionTargetId="figure@34.[151,1436,218,1490]" captionTargetPageId="34" captionText="FIGURE 24. In situ photographs of Parasesarma leptosoma species-complex, specimens not collected. A, P. leptosoma (Hilgendorf, 1869), Kenya (Photograph: S. Cannicci); B, P. gecko n. sp., Okinawa, Japan; C, P. macaco n. sp., Pingtung, Taiwan; D, P. kui n. sp., Pingtung, Taiwan; E, P. purpureum n. sp., Borneo, Malaysia; F, P. tarantula n. sp., Sulawesi, Indonesia." httpUri="https://zenodo.org/record/1440757/files/figure.png" pageId="12" pageNumber="463">24B</figureCitation>
, 
<figureCitation id="27F8B6DEFFA8FF8AD55BFB754000FB41" box="[1025,1074,1154,1179]" captionStart="FIGURE 25" captionStartId="35.[151,250,1877,1899]" captionTargetBox="[170,1415,193,1866]" captionTargetId="figure@35.[170,1415,193,1866]" captionTargetPageId="35" captionText="FIGURE 25. Habitats associated with Parasesarma macaco n. sp., P. gecko n. sp. and P. kui n. sp., specimens not collected. A–D, P. macaco n. sp., Pingtung (Paoli River), Taiwan; E, P. gecko n. sp., group of crabs on cement wall near mangrove, Okinawa, Japan; F, G, P. kui n. sp., Pingtung (Kangkou River), Taiwan. A, habitat constituted by Talipariti tiliaceum and Clerodendrum inerme; B, male climbing on T. tiliaceum; C, female climbing on C. inerme; D, ovigerous female during breeding migration; F, habitat constituted by T. tiliaceum; G, male climbing on shrubs." httpUri="https://zenodo.org/record/1440759/files/figure.png" pageId="12" pageNumber="463">25E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFA8FF8BD19DFB5145FDFE9A" lastPageId="13" lastPageNumber="464" pageId="12" pageNumber="463" type="discussion">
<paragraph id="BF7CAA5BFFA8FF8AD19DFB5145AFFAF0" blockId="12.[151,1437,974,2007]" pageId="12" pageNumber="463">
<emphasis id="8DB77649FFA8FF8AD19DFB514509FB65" bold="true" box="[199,315,1190,1215]" pageId="12" pageNumber="463">Remarks.</emphasis>
<taxonomicName id="78C3D1D8FFA8FF8AD01BFB50462CFB65" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[321,542,1190,1215]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD01BFB50462CFB65" box="[321,542,1190,1215]" italics="true" pageId="12" pageNumber="463">Parasesarma gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD37CFB514656FB65" box="[550,612,1190,1215]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD37CFB514656FB65" bold="true" box="[550,612,1190,1215]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
most closely resembles 
<taxonomicName id="78C3D1D8FFA8FF8AD22CFB5047DAFB64" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[886,1000,1191,1214]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD22CFB5047DAFB64" box="[886,1000,1191,1214]" italics="true" pageId="12" pageNumber="463">P. macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD2B5FB51401FFB65" box="[1007,1069,1190,1215]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD2B5FB51401FFB65" bold="true" box="[1007,1069,1190,1215]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
in having yellowish chela in life and the relatively slender G1, but it can be distinguished from the latter by the number of dactylar tubercles on the male chela (
<quantity id="783B07BEFFA8FF8AD07CFB1945BCFADD" box="[294,398,1262,1287]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="12" pageNumber="463" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10–15 in</quantity>
<taxonomicName id="78C3D1D8FFA8FF8AD0C3FB1845CBFADD" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[409,505,1262,1287]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD0C3FB1845CBFADD" box="[409,505,1262,1287]" italics="true" pageId="12" pageNumber="463">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD35EFB194675FADD" box="[516,583,1262,1287]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD35EFB194675FADD" bold="true" box="[516,583,1262,1287]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
versus 
<quantity id="783B07BEFFA8FF8AD3FEFB1946DDFADD" box="[676,751,1262,1287]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="12" pageNumber="463" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">6–8 in</quantity>
<taxonomicName id="78C3D1D8FFA8FF8AD3A3FB18475FFADC" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[761,877,1263,1286]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD3A3FB18475FFADC" box="[761,877,1263,1286]" italics="true" pageId="12" pageNumber="463">P. macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD220FB19478FFADD" box="[890,957,1262,1287]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD220FB19478FFADD" bold="true" box="[890,957,1262,1287]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
). The other differentiating characters are summarized in 
<tableCitation id="F2419FE0FFA8FF8AD01EFAE545A5FAF0" box="[324,407,1298,1323]" captionStart="TABLE 1" captionStartId="7.[147,224,586,606]" captionText="TABLE 1. Comparison of major differences between members of the Parasesarma leptosoma species-complex." pageId="12" pageNumber="463">Table 1</tableCitation>
.
</paragraph>
<paragraph id="BF7CAA5BFFA8FF8AD19DFAC144DDF8B1" blockId="12.[151,1437,974,2007]" pageId="12" pageNumber="463">
The sizes of ovigerous females of 
<taxonomicName id="78C3D1D8FFA8FF8AD30FFAC04687FA95" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[597,693,1334,1359]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD30FFAC04687FA95" box="[597,693,1334,1359]" italics="true" pageId="12" pageNumber="463">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD3E4FAC146CDFA95" box="[702,767,1334,1359]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD3E4FAC146CDFA95" bold="true" box="[702,767,1334,1359]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
show substantial disparity (the smallest carapace width is 
<quantity id="783B07BEFFA8FF8AD1CDFAAD44CEFAA8" box="[151,252,1370,1394]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="12" pageNumber="463" unit="mm" value="11.8">11.8 mm</quantity>
and the largest is 
<quantity id="783B07BEFFA8FF8AD08BFAAD4609FAA8" box="[465,571,1370,1394]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="12" pageNumber="463" unit="mm" value="20.2">20.2 mm</quantity>
), but no other obvious morphological variation detected between small and large specimens. The cheliped asymmetry in 
<taxonomicName id="78C3D1D8FFA8FF8AD3F2FA88473AFA4D" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[680,776,1406,1431]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD3F2FA88473AFA4D" box="[680,776,1406,1431]" italics="true" pageId="12" pageNumber="463">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD24EFA894765FA4D" box="[788,855,1406,1431]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD24EFA894765FA4D" bold="true" box="[788,855,1406,1431]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
is common in males (
<figureCitation id="27F8B6DEFFA8FF8AD53BFA8940E9FA4D" box="[1121,1243,1406,1431]" captionStart="FIGURE 4" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[192,1407,193,1911]" captionTargetId="figure@10.[192,1407,193,1911]" captionTargetPageId="10" captionText="FIGURE 4. Parasesarma gecko n. sp.. A–D, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; E, F, paratype female (15.3×13.0 mm) (ASIZ 73381), Iriomote Island, Japan; G, H, female (20.2×17.5 mm) (NMNS-7779-002), Okinawa Island, Japan, showing abnormal right second ambulatory leg; I, J, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan, showing asymmetrical chelae. A, E, G, I, dorsal views; B, F, H, J, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440717/files/figure.png" pageId="12" pageNumber="463">
Figs. 
<date id="CB7D8C9BFFA8FF8AD5FFFA88408CFA4D" box="[1189,1214,1407,1431]" pageId="12" pageNumber="463">4I</date>
, J
</figureCitation>
). Other than the asymmetry, the dactylar tubercles and the proximal pectinate crest on the palm vary substantially from 
<date id="CB7D8C9BFFA8FF8AD47CFA5544E3FA05" pageId="12" pageNumber="463" value="2010-10-15" valueMax="2010-10-03" valueMin="2010-10-15">10–15 and 3–10</date>
, respectively. Such wide range of variation has not been observed in other species of this complex. We also found a female with malformed ambulatory legs (
<figureCitation id="27F8B6DEFFA8FF8AD39CFA1D4778F9D8" box="[710,842,1514,1539]" captionStart="FIGURE 4" captionStartId="10.[151,250,1920,1942]" captionTargetBox="[192,1407,193,1911]" captionTargetId="figure@10.[192,1407,193,1911]" captionTargetPageId="10" captionText="FIGURE 4. Parasesarma gecko n. sp.. A–D, holotype male (16.2×14.6 mm) (RUMF-ZC-4710), Okinawa Island, Japan; E, F, paratype female (15.3×13.0 mm) (ASIZ 73381), Iriomote Island, Japan; G, H, female (20.2×17.5 mm) (NMNS-7779-002), Okinawa Island, Japan, showing abnormal right second ambulatory leg; I, J, paratype male (15.9×13.7 mm) (ASIZ 73381), Iriomote Island, Japan, showing asymmetrical chelae. A, E, G, I, dorsal views; B, F, H, J, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440717/files/figure.png" pageId="12" pageNumber="463">Figs. 4G, H</figureCitation>
). The variations might result from the tendency of 
<taxonomicName id="78C3D1D8FFA8FF8AD4DDFA1C44EAF9FD" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD4DDFA1C44EAF9FD" italics="true" pageId="12" pageNumber="463">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD1BEF9F9451AF9FD" box="[228,296,1550,1575]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD1BEF9F9451AF9FD" bold="true" box="[228,296,1550,1575]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
to automise its chelipeds and ambulatory legs more readily compared to other species we have observed (see “Ecology”). There are several records from 
<collectingCountry id="C7D4EACBFFA8FF8AD26DF9C44745F990" box="[823,887,1587,1610]" name="Japan" pageId="12" pageNumber="463">Japan</collectingCountry>
referred to 
<taxonomicName id="78C3D1D8FFA8FF8AD55BF9C440A0F991" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[1025,1170,1586,1611]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD55BF9C440A0F991" box="[1025,1170,1586,1611]" italics="true" pageId="12" pageNumber="463">P. leptosoma</emphasis>
</taxonomicName>
. 
<bibRefCitation id="DB52D7AAFFA8FF8AD5F9F9C54157F990" box="[1187,1381,1586,1611]" pageId="12" pageNumber="463" refId="ref17664" refString="Nakasone, Y. (1977) The ecology and distribution of mangrove animals. In: Investigations of natural monument from Okinawa 10: the emergency of investigations Rhizophoraceae mangrove of Gesashi Bay. Education Section, Okinawa Prefecture, Naha, pp. 9 - 38." type="book chapter">Nakasone (1977)</bibRefCitation>
first recorded “ 
<taxonomicName id="78C3D1D8FFA8FF8AD04BF9A045E7F9B5" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[273,469,1622,1647]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD04BF9A04511F9B4" box="[273,291,1623,1646]" italics="true" pageId="12" pageNumber="463">P.</emphasis>
aff. 
<emphasis id="8DB77649FFA8FF8AD038F9A145E7F9B5" box="[354,469,1622,1647]" italics="true" pageId="12" pageNumber="463">leptosoma</emphasis>
</taxonomicName>
” from Gesaji River, 
<collectingRegion id="7D0764B9FFA8FF8AD388F9A14705F9B5" box="[722,823,1622,1647]" country="Japan" name="Okinawa" pageId="12" pageNumber="463">Okinawa</collectingRegion>
Island, and noted that the crabs often climb on the mangrove aerial roots and leaves. Nishidaira (1980: 66) also recorded 
<taxonomicName id="78C3D1D8FFA8FF8AD2F1F98C4054F949" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[939,1126,1658,1683]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD2F1F98C478FF948" box="[939,957,1659,1682]" italics="true" pageId="12" pageNumber="463">P.</emphasis>
aff. 
<emphasis id="8DB77649FFA8FF8AD2A9F98D4054F949" box="[1011,1126,1658,1683]" italics="true" pageId="12" pageNumber="463">leptosoma</emphasis>
</taxonomicName>
from Ishigaki and Iriomote Islands. Shokita (2002) first formally recorded 
<taxonomicName id="78C3D1D8FFA8FF8AD3E4F9684763F96D" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[702,849,1694,1719]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD3E4F9684763F96D" box="[702,849,1694,1719]" italics="true" pageId="12" pageNumber="463">P. leptosoma</emphasis>
</taxonomicName>
from Oura River, 
<collectingRegion id="7D0764B9FFA8FF8AD575F96940A6F96D" box="[1071,1172,1694,1719]" country="Japan" name="Okinawa" pageId="12" pageNumber="463">Okinawa</collectingRegion>
Island, and later from Sakagawa River, 
<collectingRegion id="7D0764B9FFA8FF8AD007F93545F0F901" box="[349,450,1730,1755]" country="Japan" name="Okinawa" pageId="12" pageNumber="463">Okinawa</collectingRegion>
Island (Shokita 2003). Recently, 
<bibRefCitation id="DB52D7AAFFA8FF8AD212F93440ABF901" box="[840,1177,1730,1755]" pageId="12" pageNumber="463" refId="ref17144" refString="Maenosono, T. &amp; Naruse, T. (2015) Morphological characteristics and taxonomical problems of the genus Parasesarma De Man, 1895 (Crustacea: Decapoda: Brachyura: Sesarmidae) from the Ryukyu Archipelago, Japan. Fauna Ryukyuana, 23, 1 - 41 [in Japanese]." type="journal article">Maenosono &amp; Naruse (2015)</bibRefCitation>
reported 
<taxonomicName id="78C3D1D8FFA8FF8AD450F93441AEF901" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[1290,1436,1730,1755]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD450F93441AEF901" box="[1290,1436,1730,1755]" italics="true" pageId="12" pageNumber="463">P. leptosoma</emphasis>
</taxonomicName>
from the Ryukyu Islands, and concluded that the “ 
<taxonomicName id="78C3D1D8FFA8FF8AD387F91047ACF925" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[733,926,1766,1791]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD387F91046D9F924" box="[733,747,1767,1790]" italics="true" pageId="12" pageNumber="463">P</emphasis>
. aff. 
<emphasis id="8DB77649FFA8FF8AD271F91147ACF925" box="[811,926,1766,1791]" italics="true" pageId="12" pageNumber="463">leptosoma</emphasis>
</taxonomicName>
” (listed by 
<bibRefCitation id="DB52D7AAFFA8FF8AD573F91140DEF925" box="[1065,1260,1766,1791]" pageId="12" pageNumber="463" refId="ref17664" refString="Nakasone, Y. (1977) The ecology and distribution of mangrove animals. In: Investigations of natural monument from Okinawa 10: the emergency of investigations Rhizophoraceae mangrove of Gesashi Bay. Education Section, Okinawa Prefecture, Naha, pp. 9 - 38." type="book chapter">Nakasone (1977)</bibRefCitation>
and Nishidaira (1980) and 
<taxonomicName id="78C3D1D8FFA8FF8AD041F8FC459EF8F9" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[283,428,1802,1827]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD041F8FC459EF8F9" box="[283,428,1802,1827]" italics="true" pageId="12" pageNumber="463">P. leptosoma</emphasis>
</taxonomicName>
represent the same species. Our study has shown that only 
<taxonomicName id="78C3D1D8FFA8FF8AD502F8FC4081F8F9" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[1112,1203,1802,1827]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA8FF8AD502F8FC4081F8F9" box="[1112,1203,1802,1827]" italics="true" pageId="12" pageNumber="463">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA8FF8AD59AF8FD4133F8F9" box="[1216,1281,1802,1827]" pageId="12" pageNumber="463" rank="species">
<emphasis id="8DB77649FFA8FF8AD59AF8FD4133F8F9" bold="true" box="[1216,1281,1802,1827]" pageId="12" pageNumber="463">n. sp.</emphasis>
</taxonomicNameLabel>
occurs in the Ryukyu Islands, and consequently, the previous records of 
<taxonomicName id="78C3D1D8FFA8FF8AD26FF8D847F7F89D" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[821,965,1838,1863]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD26FF8D847F7F89D" box="[821,965,1838,1863]" italics="true" pageId="12" pageNumber="463">P. leptosoma</emphasis>
</taxonomicName>
from the area are all referred to this new species.
</paragraph>
<paragraph id="BF7CAA5BFFA8FF8BD19DF88045FDFE9A" blockId="12.[151,1437,974,2007]" lastBlockId="13.[151,1437,151,861]" lastPageId="13" lastPageNumber="464" pageId="12" pageNumber="463">
<bibRefCitation id="DB52D7AAFFA8FF8AD19DF8804544F855" box="[199,374,1911,1935]" pageId="12" pageNumber="463" refId="ref17312" refString="Man, J. G. De (1889) Uber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 4, 409 - 452." type="journal article">De Man (1889)</bibRefCitation>
, Ortmann (1894) and 
<bibRefCitation id="DB52D7AAFFA8FF8AD334F880475DF855" box="[622,879,1911,1935]" pageId="12" pageNumber="463" refId="ref17530" refString="McLay, C. L. &amp; Ryan, P. A. (1990) The terrestrial crabs Sesarma (Sesarmops) impressum and Geograpsus crinipes (Brachyura, Grapsidae, Sesarminae) recorded from the Fiji Island. Journal of the Royal Society of New Zealand, 20, 107 - 118. http: // dx. doi. org / 10.1080 / 03036758.1990.10426735" type="journal article">McLay &amp; Ryan (1990)</bibRefCitation>
reported 
<taxonomicName id="78C3D1D8FFA8FF8AD281F8804084F855" authorityName=", McLay &amp; Ryan" authorityYear="1990" box="[987,1206,1910,1935]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD281F8804084F855" box="[987,1206,1910,1935]" italics="true" pageId="12" pageNumber="463">Sesarma leptosoma</emphasis>
</taxonomicName>
from 
<collectingCountry id="C7D4EACBFFA8FF8AD5A1F8814110F855" box="[1275,1314,1910,1935]" name="Fiji" pageId="12" pageNumber="463">Fiji</collectingCountry>
Island. 
<bibRefCitation id="DB52D7AAFFA8FF8AD421F8804510F869" pageId="12" pageNumber="463" refId="ref17312" refString="Man, J. G. De (1889) Uber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 4, 409 - 452." type="journal article">De Man (1889)</bibRefCitation>
’s illustrations (pl. 10 
<figureCitation id="27F8B6DEFFA8FF8AD341F86D4659F869" box="[539,619,1946,1971]" captionStart="FIGURE 11" captionStartId="19.[151,250,1654,1676]" captionTargetBox="[152,1435,481,1633]" captionTargetId="figure@19.[152,1435,481,1633]" captionTargetPageId="19" captionText="FIGURE 11. Parasesarma gracilipes n. sp.. A–D, holotype male (13.6×11.9 mm) (MZB Cru 4803), Papua, Indonesia; E, F, paratype female (13.4×11.7 mm) (ZRC 2018.0793), same locality. A, E, dorsal views; B, F, ventral views; C, D, frontal views." httpUri="https://zenodo.org/record/1440731/files/figure.png" pageId="12" pageNumber="463">Fig. 11</figureCitation>
) show the shape of frontal border of the carapace in the dorsal view is distinctly concave, but this feature is not observed in the true 
<taxonomicName id="78C3D1D8FFA8FF8AD207F84847DFF80D" authorityName=", McLay &amp; Ryan" authorityYear="1990" box="[861,1005,1982,2007]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA8FF8AD207F84847DFF80D" box="[861,1005,1982,2007]" italics="true" pageId="12" pageNumber="463">P. leptosoma</emphasis>
</taxonomicName>
from Africa. The morphology of this Fijian material instead matches that of 
<taxonomicName id="78C3D1D8FFA9FF8BD314FF6E4699FF6A" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[590,683,151,176]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD314FF6E4699FF6A" box="[590,683,151,176]" italics="true" pageId="13" pageNumber="464">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA9FF8BD3E9FF6046C0FF6A" box="[691,754,151,176]" pageId="13" pageNumber="464" rank="species">
<emphasis id="8DB77649FFA9FF8BD3E9FF6046C0FF6A" bold="true" box="[691,754,151,176]" pageId="13" pageNumber="464">n. sp.</emphasis>
</taxonomicNameLabel>
which is known for certain from 
<collectingCountry id="C7D4EACBFFA9FF8BD535FF6F4080FF75" box="[1135,1202,152,175]" name="Japan" pageId="13" pageNumber="464">Japan</collectingCountry>
, 
<collectingCountry id="C7D4EACBFFA9FF8BD5E7FF6F4130FF6A" box="[1213,1282,152,176]" name="Guam" pageId="13" pageNumber="464">Guam</collectingCountry>
and 
<collectingCountry id="C7D4EACBFFA9FF8BD463FF6F41ABFF75" box="[1337,1433,152,175]" name="Vanuatu" pageId="13" pageNumber="464">Vanuatu</collectingCountry>
. 
<bibRefCitation id="DB52D7AAFFA9FF8BD1CDFF4B451EFF0E" box="[151,300,188,213]" pageId="13" pageNumber="464" refId="ref16423" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda-Anomura, Brachyura. In: Wells, A. &amp; Houston, W. W. K. (Eds.), Zological Catalogue of Australia, vol. 19.3 B. CSIRO Publishing Australia, Melbourne, pp. 641." type="book chapter">Davie (2002)</bibRefCitation>
recorded material from 
<collectingRegion id="7D0764B9FFA9FF8BD365FF4B46F4FF0F" box="[575,710,188,213]" country="Australia" name="Queensland" pageId="13" pageNumber="464">Queensland</collectingRegion>
, 
<collectingCountry id="C7D4EACBFFA9FF8BD388FF4B470EFF0F" box="[722,828,188,213]" name="Australia" pageId="13" pageNumber="464">Australia</collectingCountry>
, and on the basis of geography, his record is probably 
<taxonomicName id="78C3D1D8FFA9FF8BD1CDFF1644C4FF22" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[151,246,223,248]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD1CDFF1644C4FF22" box="[151,246,223,248]" italics="true" pageId="13" pageNumber="464">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA9FF8BD1A4FF284572FF22" box="[254,320,223,248]" pageId="13" pageNumber="464" rank="species">
<emphasis id="8DB77649FFA9FF8BD1A4FF284572FF22" bold="true" box="[254,320,223,248]" pageId="13" pageNumber="464">n. sp.</emphasis>
</taxonomicNameLabel>
Takeda &amp; Nunomura (1976) listed 
<taxonomicName id="78C3D1D8FFA9FF8BD381FF164759FF22" authorityName=", McLay &amp; Ryan" authorityYear="1990" box="[731,875,223,248]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="8DB77649FFA9FF8BD381FF164759FF22" box="[731,875,223,248]" italics="true" pageId="13" pageNumber="464">P. leptosoma</emphasis>
</taxonomicName>
from 
<collectingCountry id="C7D4EACBFFA9FF8BD2EEFF174051FF22" box="[948,1123,223,248]" name="New Caledonia" pageId="13" pageNumber="464">New Caledonia</collectingCountry>
based on two females, and although no figure was provided, this record falls inside the range of 
<taxonomicName id="78C3D1D8FFA9FF8BD2EBFEF2403DFEC7" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[945,1039,260,285]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD2EBFEF2403DFEC7" box="[945,1039,260,285]" italics="true" pageId="13" pageNumber="464">P. gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA9FF8BD542FEF3406BFEC7" box="[1048,1113,260,285]" pageId="13" pageNumber="464" rank="species">
<emphasis id="8DB77649FFA9FF8BD542FEF3406BFEC7" bold="true" box="[1048,1113,260,285]" pageId="13" pageNumber="464">n. sp.</emphasis>
</taxonomicNameLabel>
, and as such, their record is also referred to this species.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFA9FF8BD19DFEBB45EAFE77" pageId="13" pageNumber="464" type="etymology">
<paragraph id="BF7CAA5BFFA9FF8BD19DFEBB45EAFE77" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="464">
<emphasis id="8DB77649FFA9FF8BD19DFEBB457CFEBF" bold="true" box="[199,334,332,357]" pageId="13" pageNumber="464">Etymology.</emphasis>
The specific epithet alludes to the new species’ quick movements on vertical surfaces and its tendency to autotomise its appendages when handled, as also observed in the eponymous lizard. The name is used here as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFA9FF8BD19DFE4040E4FDE7" pageId="13" pageNumber="464" type="distribution">
<paragraph id="BF7CAA5BFFA9FF8BD19DFE4040E4FDE7" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="464">
<emphasis id="8DB77649FFA9FF8BD19DFE404552FE0A" bold="true" box="[199,352,439,464]" pageId="13" pageNumber="464">Distribution.</emphasis>
<collectingCountry id="C7D4EACBFFA9FF8BD035FE4F4581FE15" box="[367,435,440,463]" name="Japan" pageId="13" pageNumber="464">Japan</collectingCountry>
: 
<collectingRegion id="7D0764B9FFA9FF8BD09CFE404619FE0A" box="[454,555,439,464]" country="Japan" name="Okinawa" pageId="13" pageNumber="464">Okinawa</collectingRegion>
Island, Miyako Island, Ishigaki Island, Iriomote Island (
<bibRefCitation id="DB52D7AAFFA9FF8BD586FE4041A5FE0A" box="[1244,1431,439,464]" pageId="13" pageNumber="464" refId="ref17664" refString="Nakasone, Y. (1977) The ecology and distribution of mangrove animals. In: Investigations of natural monument from Okinawa 10: the emergency of investigations Rhizophoraceae mangrove of Gesashi Bay. Education Section, Okinawa Prefecture, Naha, pp. 9 - 38." type="book chapter">Nakasone 1977</bibRefCitation>
; Nishidaira 1980; Shokita 2002, 2003; 
<bibRefCitation id="DB52D7AAFFA9FF8BD31EFE2B473CFE2E" box="[580,782,476,501]" pageId="13" pageNumber="464" refId="ref17023" refString="Koller, P., Liu, H. C. &amp; Schubart, C. D. (2010) A new semiterrestrial species of Parasesarma De Man, 1895, from Taiwan (Decapoda, Brachyura, Sesarmidae). Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Folume. Crustaceana Monographs, 14, 357 - 368." type="journal article">
Koller 
<emphasis id="8DB77649FFA9FF8BD3C9FE2A46F6FE2F" box="[659,708,476,501]" italics="true" pageId="13" pageNumber="464">et al</emphasis>
. 2010
</bibRefCitation>
; 
<bibRefCitation id="DB52D7AAFFA9FF8BD240FE2A4799FE2E" box="[794,939,476,500]" pageId="13" pageNumber="464" refId="ref17763" refString="Naruse, T. (2010) Catalogue of materials deposited in the University Museum (Fujukan). University of the Ryukyus No. 3. University of the Ryukyus. The University Museum (Fujukan), University of the Ryukyus, Okinawa. [in Japanese]" type="book">Naruse 2010</bibRefCitation>
; 
<bibRefCitation id="DB52D7AAFFA9FF8BD2EDFE2A40C2FE2F" box="[951,1264,476,501]" pageId="13" pageNumber="464" refId="ref17144" refString="Maenosono, T. &amp; Naruse, T. (2015) Morphological characteristics and taxonomical problems of the genus Parasesarma De Man, 1895 (Crustacea: Decapoda: Brachyura: Sesarmidae) from the Ryukyu Archipelago, Japan. Fauna Ryukyuana, 23, 1 - 41 [in Japanese]." type="journal article">Maenosono &amp; Naruse 2015</bibRefCitation>
; 
<bibRefCitation id="DB52D7AAFFA9FF8BD5A6FE2A452BFDC2" pageId="13" pageNumber="464" refId="ref17198" refString="Maenosono, T. &amp; Saeki, T. (2016) The sesarmid (Crustacea: Decapoda: Brachyura) fauna of Ishigaki-jima Island, Ryukyu Archipelago, Japan, with new distributional records. Fauna Ryukyuana, 33, 1 - 13 [in Japanese with English abstract, Figure captions]." type="journal article">Maenosono &amp; Saeki 2016</bibRefCitation>
); 
<collectingCountry id="C7D4EACBFFA9FF8BD076FDF74543FDC2" box="[300,369,512,536]" name="Guam" pageId="13" pageNumber="464">Guam</collectingCountry>
(Rahayu &amp; Ng 2009), 
<collectingRegion id="7D0764B9FFA9FF8BD32BFE0846CAFDC2" box="[625,760,511,536]" country="Australia" name="Queensland" pageId="13" pageNumber="464">Queensland</collectingRegion>
, 
<collectingCountry id="C7D4EACBFFA9FF8BD259FE08475BFDC2" box="[771,873,511,536]" name="Australia" pageId="13" pageNumber="464">Australia</collectingCountry>
(
<bibRefCitation id="DB52D7AAFFA9FF8BD222FE084032FDC2" box="[888,1024,511,536]" pageId="13" pageNumber="464" refId="ref16423" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca: Eucarida (Part 2): Decapoda-Anomura, Brachyura. In: Wells, A. &amp; Houston, W. W. K. (Eds.), Zological Catalogue of Australia, vol. 19.3 B. CSIRO Publishing Australia, Melbourne, pp. 641." type="book chapter">Davie 2002</bibRefCitation>
); 
<collectingCountry id="C7D4EACBFFA9FF8BD549FE084008FDC2" box="[1043,1082,511,536]" name="Fiji" pageId="13" pageNumber="464">Fiji</collectingCountry>
(
<bibRefCitation id="DB52D7AAFFA9FF8BD513FDF740DAFDC2" box="[1097,1256,512,536]" pageId="13" pageNumber="464" refId="ref17312" refString="Man, J. G. De (1889) Uber einige neue oder seltene indopacifische Brachyuren. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 4, 409 - 452." type="journal article">De Man 1889</bibRefCitation>
; Ortmann 1894; 
<bibRefCitation id="DB52D7AAFFA9FF8BD1CDFDD245BBFDE6" box="[151,393,548,573]" pageId="13" pageNumber="464" refId="ref17530" refString="McLay, C. L. &amp; Ryan, P. A. (1990) The terrestrial crabs Sesarma (Sesarmops) impressum and Geograpsus crinipes (Brachyura, Grapsidae, Sesarminae) recorded from the Fiji Island. Journal of the Royal Society of New Zealand, 20, 107 - 118. http: // dx. doi. org / 10.1080 / 03036758.1990.10426735" type="journal article">McLay &amp; Ryan 1990</bibRefCitation>
), 
<collectingCountry id="C7D4EACBFFA9FF8BD0C6FDD2467BFDE7" box="[412,585,548,573]" name="New Caledonia" pageId="13" pageNumber="464">New Caledonia</collectingCountry>
(Takeda &amp; Nunomura 1976) and 
<collectingCountry id="C7D4EACBFFA9FF8BD29EFDD24012FDE6" box="[964,1056,549,572]" name="Vanuatu" pageId="13" pageNumber="464">Vanuatu</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="F7D9F9D0FFA9FF8BD19DFDB04536FC87" pageId="13" pageNumber="464" type="biology_ecology">
<paragraph id="BF7CAA5BFFA9FF8BD19DFDB04536FC87" blockId="13.[151,1437,151,861]" pageId="13" pageNumber="464">
<emphasis id="8DB77649FFA9FF8BD19DFDB04519FDBA" bold="true" box="[199,299,583,608]" pageId="13" pageNumber="464">Ecology.</emphasis>
In 
<collectingRegion id="7D0764B9FFA9FF8BD00EFDB0458BFDBA" box="[340,441,583,608]" country="Japan" name="Okinawa" pageId="13" pageNumber="464">Okinawa</collectingRegion>
Island, 
<taxonomicName id="78C3D1D8FFA9FF8BD34FFDBE46C7FDBA" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[533,757,583,608]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD34FFDBE46C7FDBA" box="[533,757,583,608]" italics="true" pageId="13" pageNumber="464">Parasesarma gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA9FF8BD3A7FDB0470CFDBA" box="[765,830,583,608]" pageId="13" pageNumber="464" rank="species">
<emphasis id="8DB77649FFA9FF8BD3A7FDB0470CFDBA" bold="true" box="[765,830,583,608]" pageId="13" pageNumber="464">n. sp.</emphasis>
</taxonomicNameLabel>
is common on the mangrove trees 
<taxonomicName id="78C3D1D8FFA9FF8BD588FDB04549FD5F" authority="Sheue, Liu &amp; Yong" authorityName="Sheue, Liu &amp; Yong" class="Magnoliopsida" family="Rhizophoraceae" genus="Kandelia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="13" pageNumber="464" phylum="Tracheophyta" rank="species" species="obovata">
<emphasis id="8DB77649FFA9FF8BD588FDB041AEFDBA" box="[1234,1436,583,608]" italics="true" pageId="13" pageNumber="464">Kandelia obovata</emphasis>
Sheue, Liu &amp; Yong
</taxonomicName>
, 
<taxonomicName id="78C3D1D8FFA9FF8BD0D0FD9B46D8FD5F" authority="Linn." authorityName="Linn." box="[394,746,620,645]" class="Magnoliopsida" family="Pandanaceae" genus="Pandanus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pandanales" pageId="13" pageNumber="464" phylum="Tracheophyta" rank="species" species="odoratissimus">
<emphasis id="8DB77649FFA9FF8BD0D0FD9B4693FD5F" box="[394,673,620,645]" italics="true" pageId="13" pageNumber="464">Pandanus odoratissimus</emphasis>
Linn.
</taxonomicName>
, as well as on rocks and man-made vertical structures (e.g. cement and pillars) (
<figureCitation id="27F8B6DEFFA9FF8BD0D9FD7845DAFD72" box="[387,488,655,680]" captionStart="FIGURE 25" captionStartId="35.[151,250,1877,1899]" captionTargetBox="[170,1415,193,1866]" captionTargetId="figure@35.[170,1415,193,1866]" captionTargetPageId="35" captionText="FIGURE 25. Habitats associated with Parasesarma macaco n. sp., P. gecko n. sp. and P. kui n. sp., specimens not collected. A–D, P. macaco n. sp., Pingtung (Paoli River), Taiwan; E, P. gecko n. sp., group of crabs on cement wall near mangrove, Okinawa, Japan; F, G, P. kui n. sp., Pingtung (Kangkou River), Taiwan. A, habitat constituted by Talipariti tiliaceum and Clerodendrum inerme; B, male climbing on T. tiliaceum; C, female climbing on C. inerme; D, ovigerous female during breeding migration; F, habitat constituted by T. tiliaceum; G, male climbing on shrubs." httpUri="https://zenodo.org/record/1440759/files/figure.png" pageId="13" pageNumber="464">Fig. 25E</figureCitation>
) near estuarine waters (
<bibRefCitation id="DB52D7AAFFA9FF8BD3A0FD67400BFD72" box="[762,1081,655,680]" pageId="13" pageNumber="464" refId="ref17144" refString="Maenosono, T. &amp; Naruse, T. (2015) Morphological characteristics and taxonomical problems of the genus Parasesarma De Man, 1895 (Crustacea: Decapoda: Brachyura: Sesarmidae) from the Ryukyu Archipelago, Japan. Fauna Ryukyuana, 23, 1 - 41 [in Japanese]." type="journal article">Maenosono &amp; Naruse 2015</bibRefCitation>
; 
<bibRefCitation id="DB52D7AAFFA9FF8BD51DFD674144FD72" box="[1095,1398,655,680]" pageId="13" pageNumber="464" refId="ref17198" refString="Maenosono, T. &amp; Saeki, T. (2016) The sesarmid (Crustacea: Decapoda: Brachyura) fauna of Ishigaki-jima Island, Ryukyu Archipelago, Japan, with new distributional records. Fauna Ryukyuana, 33, 1 - 13 [in Japanese with English abstract, Figure captions]." type="journal article">Maenosono &amp; Saeki 2016</bibRefCitation>
). It was found together with 
<taxonomicName id="78C3D1D8FFA9FF8BD0EAFD4247C4FD17" authority="(White, 1847)" baseAuthorityName="White" baseAuthorityYear="1847" box="[432,1014,692,717]" class="Malacostraca" family="Grapsidae" genus="Metopograpsus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="8DB77649FFA9FF8BD0EAFD4246F6FD17" box="[432,708,692,717]" italics="true" pageId="13" pageNumber="464">Metopograpsus latifrons</emphasis>
(White, 1847) (Grapsidae)
</taxonomicName>
in the same habitat (
<collectingRegion id="7D0764B9FFA9FF8BD5B8FD43417BFD17" box="[1250,1353,692,717]" country="Japan" name="Okinawa" pageId="13" pageNumber="464">Okinawa</collectingRegion>
Island, 
<collectingCountry id="C7D4EACBFFA9FF8BD1CDFD2F44EAFD2A" box="[151,216,728,752]" name="Japan" pageId="13" pageNumber="464">Japan</collectingCountry>
) (present paper). The new species is active during day and night time, moves fast between mangrove roots, branches or other vertical surfaces. 
<taxonomicName id="78C3D1D8FFA9FF8BD360FD0A472EFCCF" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[570,796,764,789]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="gecko" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD360FD0A472EFCCF" box="[570,796,764,789]" italics="true" pageId="13" pageNumber="464">Parasesarma gecko</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA9FF8BD27DFD0B4758FCCF" box="[807,874,764,789]" pageId="13" pageNumber="464" rank="species">
<emphasis id="8DB77649FFA9FF8BD27DFD0B4758FCCF" bold="true" box="[807,874,764,789]" pageId="13" pageNumber="464">n. sp.</emphasis>
</taxonomicNameLabel>
tends to lose its appendages very readily when captured, apparently more so than 
<taxonomicName id="78C3D1D8FFA9FF8BD369FCD64640FCE2" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[563,626,799,824]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="kui" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD369FCD64640FCE2" box="[563,626,799,824]" italics="true" pageId="13" pageNumber="464">P. kui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA9FF8BD3D8FCE846F7FCE2" box="[642,709,799,824]" pageId="13" pageNumber="464" rank="species">
<emphasis id="8DB77649FFA9FF8BD3D8FCE846F7FCE2" bold="true" box="[642,709,799,824]" pageId="13" pageNumber="464">n. sp.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="78C3D1D8FFA9FF8BD382FCD6477FFCE2" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[728,845,801,824]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="macaco" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD382FCD6477FFCE2" box="[728,845,801,824]" italics="true" pageId="13" pageNumber="464">P. macaco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA9FF8BD206FCE847ADFCE2" box="[860,927,799,824]" pageId="13" pageNumber="464" rank="species">
<emphasis id="8DB77649FFA9FF8BD206FCE847ADFCE2" bold="true" box="[860,927,799,824]" pageId="13" pageNumber="464">n. sp.</emphasis>
</taxonomicNameLabel>
from 
<collectingCountry id="C7D4EACBFFA9FF8BD2B4FCE84072FCE2" box="[1006,1088,799,824]" name="Taiwan" pageId="13" pageNumber="464">Taiwan</collectingCountry>
and 
<taxonomicName id="78C3D1D8FFA9FF8BD5DBFCD6413BFCE2" authority="Li &amp; Rahayu &amp; Ng, 2018" authorityName="Li &amp; Rahayu &amp; Ng" authorityYear="2018" box="[1153,1289,799,824]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="tarantula" status="sp. nov.">
<emphasis id="8DB77649FFA9FF8BD5DBFCD6413BFCE2" box="[1153,1289,799,824]" italics="true" pageId="13" pageNumber="464">P. tarantula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9684CB32FFA9FF8BD44FFCE8416BFCE2" box="[1301,1369,799,824]" pageId="13" pageNumber="464" rank="species">
<emphasis id="8DB77649FFA9FF8BD44FFCE8416BFCE2" bold="true" box="[1301,1369,799,824]" pageId="13" pageNumber="464">n. sp.</emphasis>
</taxonomicNameLabel>
from Sulawesi.
</paragraph>
</subSubSection>
</treatment>
</document>