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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.252.3588" ID-GBIF-Dataset="a4366e50-a271-44c0-bdef-b52c8cad0b85" ID-PMC="PMC3560839" ID-Pensoft-Pub="1313-2970-252-1" ID-PubMed="23378811" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-252-1" ModsDocOrigin="ZooKeys 252" ModsDocTitle="Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)" checkinTime="1451247847790" checkinUser="pensoft" docAuthor="Bond, Jason E." docDate="2012" docId="14F0986B5102C4E8318B35FAB426BC23" docLanguage="en" docName="ZooKeys 252: 1-209" docOrigin="ZooKeys 252" docSource="http://dx.doi.org/10.3897/zookeys.252.3588" docTitle="Aptostichus sinnombre Bond, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="132" masterDocId="FFAD871BFFD4FF8CFFED1A5CFFE7FFA0" masterDocTitle="Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)" masterLastPageNumber="209" masterPageNumber="1" pageNumber="131" updateTime="1668155039594" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Phylogenetic treatment and taxonomic revision of the trapdoor spider genus Aptostichus Simon (Araneae, Mygalomorphae, Euctenizidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bond, Jason E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>252</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>209</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.252.3588</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.252.3588</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-252-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152039303" LSID="urn:lsid:zoobank.org:act:2ADC46DE-8C5E-4D67-B8A7-C60522405025" httpUri="http://treatment.plazi.org/id/14F0986B5102C4E8318B35FAB426BC23" lastPageId="131" lastPageNumber="132" pageId="130" pageNumber="131">
<subSubSection pageId="130" pageNumber="131" type="nomenclature">
<paragraph pageId="130" pageNumber="131">
<taxonomicName LSID="The No-name Trapdoor Spider" class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sinnombre" order="Araneae" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="sinnombre">Aptostichus sinnombre</taxonomicName>
<taxonomicNameLabel pageId="130" pageNumber="131">sp. n.</taxonomicNameLabel>
Figures 266-273Map 30
</paragraph>
</subSubSection>
<subSubSection pageId="130" pageNumber="131" type="types">
<paragraph pageId="130" pageNumber="131">Types.</paragraph>
<paragraph pageId="130" pageNumber="131">Male holotype (MY3761) from California, San Diego County, Anza-Borrego Desert State Park, Indian Gorge, 0.4km W mouth of Torote Canyon, 32.86923, -116.23740 1, 350m, coll. M. Hedin 20.ii.2009; male paratype (MY3823) from near type locality, mouth of canyon Sin Nombre, 32.8584, -116.1473 1, 200m, coll. M. Hedin 13.iii.2010; deposited in AUMNH.</paragraph>
</subSubSection>
<subSubSection pageId="130" pageNumber="131" type="etymology">
<paragraph pageId="130" pageNumber="131">Etymology.</paragraph>
<paragraph pageId="130" pageNumber="131">The specific epithet is a noun in apposition taken from name of canyon near the type locality, Sin Nombre.</paragraph>
</subSubSection>
<subSubSection pageId="130" pageNumber="131" type="diagnosis">
<paragraph pageId="130" pageNumber="131">Diagnosis.</paragraph>
<paragraph pageId="130" pageNumber="131">
Males can be differentiated from all other
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus" order="Araneae" pageId="130" pageNumber="131" phylum="Arthropoda" rank="genus">Aptostichus</taxonomicName>
species on the basis of a unique conformation of the retrolateral distal-most spination pattern of tibia I, which comprises 3-6 spines arranged in a distinct cluster, a low rounded metatarsal mating apophysis that bears a single strong spine, and a convex shaped (curved) tarsus (Figs 266, 269, 272); all other known species lack this combination.
</paragraph>
</subSubSection>
<subSubSection lastPageId="131" lastPageNumber="132" pageId="130" pageNumber="131" type="description">
<paragraph pageId="130" pageNumber="131">Description of male holotype.</paragraph>
<paragraph pageId="130" pageNumber="131">
Specimen preparation and condition. Specimen collected live, wandering, preserved in 80% EtOH. Coloration and specimen in relatively good condition. Pedipalp, leg I left side removed, stored in vial with specimen; legs right side removed for tissue storage. General coloration. Carapace, chelicerae, yellowish brown 10YR 5/4. Abdomen light yellowish brown 10YR 6/4, light dorsal chevron markings (Fig. 273). Cephalothorax. Carapace 5.75 long, 5.05 wide, very hirsute with intermingled thin white, black setae; stout black bristles along fringe; surface smooth, pars cephalica elevated. Fringe, posterior margin lacks black bristles. Foveal groove deep, straight. Eyes elevated on low mound. AER slightly procurved, PER strongly recurved.
<normalizedToken originalValue="PMEs">PME's</normalizedToken>
smaller in diameter than
<normalizedToken originalValue="AMEs">AME's</normalizedToken>
. Sternum moderately setose, STRl 3.10, STRw 2.67. Posterior sternal sigilla moderate in size, positioned towards center, not contiguous, anterior sigilla pairs small, oval, marginal. Chelicerae with distinct anterior tooth row comprising 7 teeth, posterior margin with patch of small denticles. Palpal endites with patch of small cuspules on proximal, inner margin, labium lacks cuspules, LBw 0.97, LBl 0.65. Rastellum consists of 3 spines not on prominent mound. Abdomen. Setose, heavy black setae intermingled with fine black setae. Legs. Legs long; Leg I: 6.88, 5.19, 4.35, 2.88, 1.90; leg IV: 6.19, 3.05. Tarsus I, swollen distally, tarsus IV straight. Heavy short scopulae on tarsi I, II; lighter on legs III, IV. Tarsus I with single, slightly staggered row of 13 trichobothria. Leg I spination pattern illustrated in Figures 266, 267, 269, 270, 272; TSp 4, TSr 4, TSrd 3. Knob-like metatarsal apophysis bearing a single large spine. Pedipalp. Articles slender (Fig. 268), distal prolateral spine on tibia (Fig. 271). PTw 0.85, PTl 2.57, Bl 1.45. Embolus slender, tapering sharply toward tip, lacking serrations (Fig. 271).
</paragraph>
<paragraph lastPageId="131" lastPageNumber="132" pageId="130" pageNumber="131">
Variation (2). Cl 5.25-5.75, Cw 4.70-5.05, STRl 3.10-3.13, STRw 2.57-2.67, LBw 0.90-0.97, LBl 0.56-0.65, leg I: 6.38-6.88, 4.70-5.19, 4.15-4.35, 2.67-2.88,
<pageBreakToken pageId="131" pageNumber="132" start="start">1.75</pageBreakToken>
-1.90; leg IV: 5.88-6.19, 2.92-3.05; PTl 2.38-2.57, PTw 0.85-0.85, Bl 1.45-1.53, TSp 2-4, TSr 4-4, TSrd 3-6.
</paragraph>
</subSubSection>
<subSubSection pageId="131" pageNumber="132" type="description">
<paragraph pageId="131" pageNumber="132">Description of female.</paragraph>
<paragraph pageId="131" pageNumber="132">Known only from male specimens.</paragraph>
<caption pageId="131" pageNumber="132">
<paragraph pageId="131" pageNumber="132">
Figures 266-272.
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sinnombre" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="sinnombre">Aptostichus sinnombre</taxonomicName>
sp. n., male specimens from San Diego County; scale bars = 1.0mm. 266-268 male holotype (MY3761) 266 retrolateral aspect, leg I [806546] 267 prolateral aspect, leg I [806550] 268 retrolateral aspect, pedipalp [806552] 269-271 male paratype (MY3823) 269 retrolateral aspect, leg I [806554] 270 prolateral aspect, leg I [806558] 271 retrolateral aspect, pedipalp [806560] 272 line drawing, spination pattern, leg I tibia and metatarsus of holotype.
</paragraph>
</caption>
<caption pageId="131" pageNumber="132">
<paragraph pageId="131" pageNumber="132">
Figure 273. Live photograph,
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sinnombre" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="sinnombre">Aptostichus sinnombre</taxonomicName>
sp. n. male holotype (MY3761); note leg I, right hand side missing.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="131" pageNumber="132" type="material examined">
<paragraph pageId="131" pageNumber="132">Material examined.</paragraph>
<paragraph pageId="131" pageNumber="132">Known only from the type material.</paragraph>
</subSubSection>
<subSubSection pageId="131" pageNumber="132" type="distribution">
<paragraph pageId="131" pageNumber="132">Distribution and natural history.</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sinnombre" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="sinnombre">Aptostichus sinnombre</taxonomicName>
is known from only two specimens from the Anza-Borrego Desert State Park, Colorado Desert habitat, in San Diego County and is sympatric with
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus anzaborrego" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="anzaborrego">Aptostichus anzaborrego</taxonomicName>
. Males appear to disperse during late winter months (February, March), based on the limited data available.
</paragraph>
</subSubSection>
<subSubSection pageId="131" pageNumber="132" type="conservation status">
<paragraph pageId="131" pageNumber="132">Conservation status.</paragraph>
<paragraph pageId="131" pageNumber="132">
The conservation status of
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus sinnombre" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="sinnombre">Aptostichus sinnombre</taxonomicName>
is likely imperiled given its restricted distribution and rarity in collections.
</paragraph>
</subSubSection>
<subSubSection pageId="131" pageNumber="132" type="species concept applied">
<paragraph pageId="131" pageNumber="132">Species concept applied.</paragraph>
<paragraph pageId="131" pageNumber="132">Morphological/Phylogenetic.</paragraph>
<subSection pageId="131" pageNumber="132" type="the simus species group">
<paragraph pageId="131" pageNumber="132">
The
<taxonomicName class="Insecta" family="Staphylinidae" genus="Simus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Simus" order="Coleoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="genus">Simus</taxonomicName>
species group
</paragraph>
<paragraph pageId="131" pageNumber="132">Included Species.</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus simus" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="simus">Aptostichus simus</taxonomicName>
Chamberlin, 1917
</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus satleri" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="satleri">Aptostichus satleri</taxonomicName>
sp. n.
</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus elisabethae" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="elisabethae">Aptostichus elisabethae</taxonomicName>
sp. n.
</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus fornax" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="fornax">Aptostichus fornax</taxonomicName>
sp. n.
</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus lucerne" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="lucerne">Aptostichus lucerne</taxonomicName>
sp. n.
</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus bonoi" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="bonoi">Aptostichus bonoi</taxonomicName>
sp. n.
</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus fisheri" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="fisheri">Aptostichus fisheri</taxonomicName>
sp. n.
</paragraph>
<paragraph pageId="131" pageNumber="132">
<taxonomicName class="Arachnida" family="Euctenizidae" genus="Aptostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aptostichus cajalco" order="Araneae" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="cajalco">Aptostichus cajalco</taxonomicName>
sp. n.
</paragraph>
</subSection>
</subSubSection>
</treatment>
</document>