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<document id="AE76378EE9098CBAAC70AA5D19612F28" ID-DOI="10.5281/zenodo.5404276" ID-ISSN="0374-1036" ID-Zenodo-Dep="5404276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132078781" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="142187F5FFD6FF9EF5CFFF66FBA2FEB3" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.161-190.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus gabonensis , Levasseur 1966" docType="treatment" docVersion="4" lastPageNumber="178" masterDocId="E818FF8DFFC6FF8FF464FFBCFFA1FFC7" masterDocTitle="Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="190" masterPageNumber="161" pageNumber="177" updateTime="1699162834734" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CB8672C4005B48E2D88FA46F45D45F61">Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
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<mods:namePart id="09BA4EA349BAC8382ADF43154F2F2641">Hromádka, Lubomír</mods:namePart>
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<mods:typeOfResource id="59B18ABB10C85534C3F02CDCF7CDB4EE">text</mods:typeOfResource>
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<mods:title id="30D20266CFC88FFC568834918DD80E36">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="6C761B55BA9B679F7DCF1ACA22AC3A05">
<mods:date id="7018553DDEC8FF0E88B9133A942AE86C">2009</mods:date>
<mods:detail id="AA32CABA9CF1848E4F1806A560A28D8A" type="pubDate">
<mods:number id="99AFAC7AD30EFB72AFF05688E6118191">2009-06-30</mods:number>
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<mods:number id="F1E3E952E660A4A27C4094967E02FB0F">49</mods:number>
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<mods:number id="43AE2E5E602536F97FFEA33ECF5E469B">1</mods:number>
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<mods:classification id="F3B826397B85E3E39DEF123541AE108C">journal article</mods:classification>
<mods:identifier id="CE3C7205FAEC5535BBCC23567F4A1443" type="DOI">10.5281/zenodo.5404276</mods:identifier>
<mods:identifier id="B779EA8B1BB65762CC4A4446F339A66D" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="C457667417E636A93207F84AE643721E" type="Zenodo-Dep">5404276</mods:identifier>
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<treatment id="142187F5FFD6FF9EF5CFFF66FBA2FEB3" ID-DOI="http://doi.org/10.5281/zenodo.5337806" ID-Zenodo-Dep="5337806" LSID="urn:lsid:plazi:treatment:142187F5FFD6FF9EF5CFFF66FBA2FEB3" httpUri="http://treatment.plazi.org/id/142187F5FFD6FF9EF5CFFF66FBA2FEB3" lastPageId="17" lastPageNumber="178" pageId="16" pageNumber="177">
<subSubSection id="D4926568FFD6FF9FF5CFFF66FCC1FF34" box="[427,864,218,243]" pageId="16" pageNumber="177" type="nomenclature">
<paragraph id="9C3736E3FFD6FF9FF5CFFF66FCC1FF34" blockId="16.[427,864,218,243]" box="[427,864,218,243]" pageId="16" pageNumber="177">
<heading id="C77F818FFFD6FF9FF5CFFF66FCC1FF34" bold="true" box="[427,864,218,243]" centered="true" fontSize="8" level="2" pageId="16" pageNumber="177" reason="3">
<taxonomicName id="5B884D60FFD6FF9FF5CFFF66FCC1FF34" ID-CoL="76ZCM" authority="Levasseur, 1966" authorityName=", Levasseur" authorityYear="1966" box="[427,864,218,243]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="species" species="gabonensis">
<emphasis id="AEFCEAF1FFD6FF9FF5CFFF66FCC1FF34" bold="true" box="[427,864,218,243]" pageId="16" pageNumber="177">
<emphasis id="AEFCEAF1FFD6FF9FF5CFFF66FD07FF34" bold="true" box="[427,678,218,243]" italics="true" pageId="16" pageNumber="177">Philonthus gabonensis</emphasis>
<bibRefCitation id="F8194B12FFD6FF9FF6C9FF67FCC1FF34" author="LEVASSEUR K." box="[685,864,219,243]" pageId="16" pageNumber="177" pagination="1210 - 1219" refId="ref16620" refString="LEVASSEUR K. 1966: Contribution a la faune du Congo (Brazzaville). Mission A. Villiers et A. Descarpentries, XXXII. Coleopteres Staphylinoidea (pars). Bulletin de l'Institut Fondamental d'Afrique Noire, Series A 28 (3): 1210 - 1219." type="journal article" year="1966">Levasseur, 1966</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD6FF9FF62CFF45FD17FECB" box="[584,694,249,268]" pageId="16" pageNumber="177" type="description">
<paragraph id="9C3736E3FFD6FF9FF62CFF45FD17FECB" blockId="16.[151,694,249,303]" box="[584,694,249,268]" pageId="16" pageNumber="177">
(
<figureCitation id="04B32A66FFD6FF9FF62AFF45FD0EFECB" box="[590,687,249,268]" captionStart="Figs" captionStartId="14.[151,190,1467,1486]" captionTargetBox="[222,1059,214,1418]" captionTargetId="figure-9@14.[212,1068,195,1435]" captionTargetPageId="14" captionText="Figs. 25–37. 25–29 – Philonthus combustus Fauvel, 1907; 30–34 – P. falco sp. nov.; 35–37 – P.gabonensis Levasseur, 1966. 25, 30, 35 – aedeagus, ventral view; 26, 31, 36 – aedeagus, lateral view; 27, 33 – apex of paramere with sensory peg setae, ventral view; 28 – female tergite X, ventral view; 29 – gonocoxite of female genital segment; 32, 37 – aedeagus without paramere, ventral view; 34 – elytra." figureDoi="http://doi.org/10.5281/zenodo.5404282" httpUri="https://zenodo.org/record/5404282/files/figure.png" pageId="16" pageNumber="177">Figs 35–37</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD6FF9FF4F3FEA1FD84FEE8" box="[151,549,284,303]" pageId="16" pageNumber="177" type="reference_group">
<paragraph id="9C3736E3FFD6FF9FF4F3FEA1FD84FEE8" blockId="16.[151,694,249,303]" box="[151,549,284,303]" pageId="16" pageNumber="177">
<treatmentCitationGroup id="BC9811CDFFD6FF9FF4F3FEA1FD84FEE8" box="[151,549,284,303]" pageId="16" pageNumber="177">
<treatmentCitation id="1D2910F2FFD6FF9FF4F3FEA1FD80FEE8" author="LEVASSEUR K." box="[151,545,284,303]" page="1211" pageId="16" pageNumber="177" year="1966">
<taxonomicName id="5B884D60FFD6FF9FF4F3FEA1FD80FEE8" ID-CoL="76ZCM" authority="Levasseur, 1966: 1211" authorityName="Levasseur" authorityPageNumber="1211" authorityYear="1966" box="[151,545,284,303]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="species" species="gabonensis">
<emphasis id="AEFCEAF1FFD6FF9FF4F3FEA1FEF8FEE8" box="[151,345,285,303]" italics="true" pageId="16" pageNumber="177">Philonthus gabonensis</emphasis>
<bibRefCitation id="F8194B12FFD6FF9FF53AFEA0FD80FEE8" author="LEVASSEUR K." box="[350,545,284,303]" pageId="16" pageNumber="177" pagination="1210 - 1219" refId="ref16620" refString="LEVASSEUR K. 1966: Contribution a la faune du Congo (Brazzaville). Mission A. Villiers et A. Descarpentries, XXXII. Coleopteres Staphylinoidea (pars). Bulletin de l'Institut Fondamental d'Afrique Noire, Series A 28 (3): 1210 - 1219." type="journal article" year="1966">Levasseur, 1966: 1211</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD6FF9FF4F3FEEDFE9DFE73" pageId="16" pageNumber="177" type="materials_examined">
<paragraph id="9C3736E3FFD6FF9FF4F3FEEDFE03FEA3" blockId="16.[151,1127,336,436]" box="[151,418,336,356]" pageId="16" pageNumber="177">
<emphasis id="AEFCEAF1FFD6FF9FF4F3FEEDFEB1FEA3" bold="true" box="[151,272,336,356]" pageId="16" pageNumber="177">
<typeStatus id="43338841FFD6FF9FF4F3FEEDFF65FEA3" box="[151,196,337,356]" pageId="16" pageNumber="177">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E49F7673FFD6FF9FF571FEEDFEF1FEA3" box="[277,336,337,356]" name="Gabon" pageId="16" pageNumber="177">Gabon</collectingCountry>
, Belinga.
</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4F3FED0FE9DFE73" blockId="16.[151,1127,336,436]" pageId="16" pageNumber="177">
<emphasis id="AEFCEAF1FFD6FF9FF4F3FED0FED8FEB8" bold="true" box="[151,377,363,383]" pageId="16" pageNumber="177">Type material examined.</emphasis>
<typeStatus id="43338841FFD6FF9FF51AFED0FE77FEBA" box="[382,470,364,383]" pageId="16" pageNumber="177" type="paratype">PARATYPES</typeStatus>
: 2 JJ, ‘
<collectingCountry id="E49F7673FFD6FF9FF67AFED0FDFCFEB8" box="[542,605,364,383]" name="Gabon" pageId="16" pageNumber="177">Gabon</collectingCountry>
, Belinga,
<date id="E8361023FFD6FF9FF6D6FED0FCB1FEB8" box="[690,784,364,383]" pageId="16" pageNumber="177" value="1963-03-12">12.iii.1963</date>
, (199 L), Mission biologique au
<collectingCountry id="E49F7673FFD6FF9FF04CFED0FBC2FEB8" box="[1064,1123,364,383]" name="Gabon" pageId="16" pageNumber="177">Gabon</collectingCountry>
, H. Coiffait leg., Coll. I. R. Sc. N. B. //
<typeStatus id="43338841FFD6FF9FF5B9FE3BFD8BFE5D" box="[477,554,391,410]" pageId="16" pageNumber="177" type="paratype">Paratype</typeStatus>
,
<taxonomicName id="5B884D60FFD6FF9FF656FE3BFCF5FE5D" authority=", Levasseur" authorityName=", Levasseur" authorityYear="1966" box="[562,852,391,410]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="177" phylum="Arthropoda" rank="species" species="gabonensis">
<emphasis id="AEFCEAF1FFD6FF9FF656FE3BFD55FE5E" box="[562,756,391,409]" italics="true" pageId="16" pageNumber="177">Philonthus gabonensis</emphasis>
, Levasseur
</taxonomicName>
det. [ochre oblong labels, handwritten]’ (
<collectionCode id="FA99AE26FFD6FF9FF488FE1DFE93FE73" box="[236,306,417,436]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="177">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD6FF9FF4F3FE70FE23F9E3" pageId="16" pageNumber="177" type="description">
<paragraph id="9C3736E3FFD6FF9FF4F3FE70FBC2FE23" blockId="16.[151,1127,460,1572]" box="[151,1123,460,485]" pageId="16" pageNumber="177">
<emphasis id="AEFCEAF1FFD6FF9FF4F3FE70FE9CFE23" bold="true" box="[151,317,460,484]" pageId="16" pageNumber="177">Redescription.</emphasis>
Body length
<quantity id="5B709B06FFD6FF9FF5A9FE71FDEEFE23" box="[461,591,461,485]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="8.1" metricValueMin="7.1" pageId="16" pageNumber="177" unit="mm" value="7.6" valueMax="8.1" valueMin="7.1">7.1–8.1 mm</quantity>
, length of forebody (to end of elytra)
<quantity id="5B709B06FFD6FF9FF786FE70FBFEFE23" box="[994,1119,460,484]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="4.1" metricValueMin="3.6" pageId="16" pageNumber="177" unit="mm" value="3.8499999999999996" valueMax="4.1" valueMin="3.6">3.6–4.1mm</quantity>
.
</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FE50FE49FD63" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">Colouration. Head and pronotum black with blue-violet metallic reflex, scutellum black, elytra red, elytral shoulders smoky black, abdomen black with blue-greenish metallic reflex, clypeus along anterior margin, antennal sockets and maxillary and labial palpi narrowly yellow-brown, mandibles somewhat darker, medial face black, ventral side of antennomere 1 brown-yellow, dorsal side and remaining antennomeres black, femora and tarsi yellowbrown, tibiae somewhat darker.</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FD10FCEAFC83" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">Head of rounded shape, as long as wide, slightly narrowed posteriad. Eyes slightly projecting, longer than temples (ratio 12.5: 9). Posterior angles of head each with one long and several short bristles. Four coarse punctures between eyes, distance between medial interocular punctures 3 times as large as distance between medial and lateral interocular punctures. Temporal area densely punctate; surface without microsculpture.</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FCF1FEB1FC63" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">Antennae long, reaching posterior margin of pronotum when reclined, all antennomeres longer than wide. Relative length of antennomeres: 1 = 10; 2 = 5.5; 3 = 6; 4–7 = 4; 8–10 = 2.5; 11 = 5.</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FC11FBB4FBC3" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">Pronotum somewhat longer than wide (ratio 30: 28), slightly narrowed posteriad in straight line. Each dorsal row with 5 coarse punctures. Each sublateral row with 2 punctures, puncture 1 situated behind level of puncture 3 of dorsal row. Surface without microsculpture.</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FBB0FCFFFB83" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">Scutellum densely and coarsely punctate, punctures as large as eye-facets, separated in transverse direction by distance smaller than a puncture diameters.</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FBF0FC78FB03" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">Elytra combined vaguely wider than long (ratio 40: 38), slightly widened posteriad, densely and finely punctate, punctures somewhat larger than eye-facets, separated in transverse direction by about a puncture diameter. Anterior angles and lateral margins each with several dark bristles of variable length. Setation dark; surface without microsculpture.</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FB71FE72FAC3" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">Legs. Metatibia shorter than metatarsus (ratio 21: 26), relative length of metatarsomeres: 1 = 9; 2 = 3.5; 3–4 = 3; 5 = 7.</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FAB0FEBBFA63" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">Abdomen parallel-sided, slightly narrowed from tergite V towards apex. Elevated area between two basal lines on first three visible tergites densely punctate, punctation of visible tergites finer and denser than that on elytra, becoming slightly sparser towards posterior margin of each tergite. Surface between punctures without microsculpture; setation similar as on elytra.</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1FA10FE8FF9C3" blockId="16.[151,1127,460,1572]" pageId="16" pageNumber="177">
Male. Protarsomeres 1–3 strongly dilated and sub-bilobed, covered with modified pale setae ventrally, protarsomere 4 triangular, distinctly narrower than preceding ones. Aedeagus (
<figureCitation id="04B32A66FFD6FF9FF4FBFA50FE83F9C3" box="[159,290,1516,1540]" captionStart="Figs" captionStartId="14.[151,190,1467,1486]" captionTargetBox="[222,1059,214,1418]" captionTargetId="figure-9@14.[212,1068,195,1435]" captionTargetPageId="14" captionText="Figs. 25–37. 25–29 – Philonthus combustus Fauvel, 1907; 30–34 – P. falco sp. nov.; 35–37 – P.gabonensis Levasseur, 1966. 25, 30, 35 – aedeagus, ventral view; 26, 31, 36 – aedeagus, lateral view; 27, 33 – apex of paramere with sensory peg setae, ventral view; 28 – female tergite X, ventral view; 29 – gonocoxite of female genital segment; 32, 37 – aedeagus without paramere, ventral view; 34 – elytra." figureDoi="http://doi.org/10.5281/zenodo.5404282" httpUri="https://zenodo.org/record/5404282/files/figure.png" pageId="16" pageNumber="177">Figs. 35–37</figureCitation>
).
</paragraph>
<paragraph id="9C3736E3FFD6FF9FF4D1F9B0FE23F9E3" blockId="16.[151,1127,460,1572]" box="[181,386,1548,1572]" pageId="16" pageNumber="177">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD7FF9EF415FF67FEC7FE93" pageId="17" pageNumber="178" type="diagnosis">
<paragraph id="9C3736E3FFD7FF9EF415FF67FD4FFEF3" blockId="17.[113,1089,219,372]" pageId="17" pageNumber="178">
<emphasis id="AEFCEAF1FFD7FF9EF415FF67FECAFF34" bold="true" box="[113,363,219,243]" pageId="17" pageNumber="178">Differential diagnosis.</emphasis>
<taxonomicName id="5B884D60FFD7FF9EF515FF61FDC2FF33" authorityName=", Levasseur" authorityYear="1966" box="[369,611,221,244]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="178" phylum="Arthropoda" rank="species" species="gabonensis">
<emphasis id="AEFCEAF1FFD7FF9EF515FF61FDC2FF33" box="[369,611,221,244]" italics="true" pageId="17" pageNumber="178">Philonthus gabonensis</emphasis>
</taxonomicName>
may be distinguished from the similar
<taxonomicName id="5B884D60FFD7FF9EF067FF61FF37FED3" authority="Hromádka, 2009" authorityName="Hromádka" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="178" phylum="Arthropoda" rank="species" species="aonyx" status="sp. nov.">
<emphasis id="AEFCEAF1FFD7FF9EF067FF61FF37FED3" italics="true" pageId="17" pageNumber="178">P. aonyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5CF578AFFD7FF9EF4F9FF40FF4EFED4" box="[157,239,252,275]" pageId="17" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
by unicoloured elytra, narrower head, differently coloured antennomere 1, finer punctation of abdomen and different shape of the aedeagus.
</paragraph>
<paragraph id="9C3736E3FFD7FF9EF415FE87FEC7FE93" blockId="17.[113,1089,219,372]" box="[113,358,315,340]" pageId="17" pageNumber="178">
<emphasis id="AEFCEAF1FFD7FF9EF415FE87FF4EFE94" bold="true" box="[113,239,315,339]" pageId="17" pageNumber="178">Bionomics.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD7FF9EF415FEE7FBA2FEB3" box="[113,1027,347,372]" pageId="17" pageNumber="178" type="distribution">
<paragraph id="9C3736E3FFD7FF9EF415FEE7FBA2FEB3" blockId="17.[113,1089,219,372]" box="[113,1027,347,372]" pageId="17" pageNumber="178">
<emphasis id="AEFCEAF1FFD7FF9EF415FEE7FEA5FEB4" bold="true" box="[113,260,347,371]" pageId="17" pageNumber="178">Distribution.</emphasis>
<collectingCountry id="E49F7673FFD7FF9EF56EFEE0FEF0FEB3" box="[266,337,348,372]" name="Gabon" pageId="17" pageNumber="178">Gabon</collectingCountry>
(this paper);
<collectingCountry id="E49F7673FFD7FF9EF584FEE0FD4EFEB3" box="[480,751,348,372]" name="Central African Republic" pageId="17" pageNumber="178">Central African Republic</collectingCountry>
, ‘Congo’ (HERMAN 2001).
</paragraph>
</subSubSection>
</treatment>
</document>
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