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<document id="FFD18A80EFB563D134980B8B8438023A" ID-DOI="10.11646/zootaxa.4103.4.1" ID-GBIF-Dataset="cbd5532b-82f3-4fc1-a7f3-cd1a0e795c49" ID-ISSN="1175-5326" ID-Zenodo-Dep="264759" ID-ZooBank="B6428194-1C7A-4A9F-9038-EE4A78222A06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460614658210" checkinUser="plazi" docAuthor="Liu, Xingyue, Lu, Xiumei &amp; Zhang, Weiwei" docDate="2016" docId="03FBD81AAF20FFADFF58FBDCAC44A096" docLanguage="en" docName="zootaxa.4103.4.1.pdf.imf" docOrigin="Zootaxa 4103 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nanoraphidia Engel 2002" docType="treatment" docVersion="8" lastPageNumber="316" masterDocId="FFC2A062AF2FFFA2FFCFFFC0AF18A724" masterDocTitle="New genera and species of the minute snakeflies (Raphidioptera: Mesoraphidiidae: Nanoraphidiini) from the mid Cretaceous of Myanmar" masterLastPageNumber="324" masterPageNumber="301" pageNumber="316" updateTime="1698329959600" updateUser="plazi">
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<mods:title id="52BB833BA2A251B4B3CEBDFC1C80BF71">New genera and species of the minute snakeflies (Raphidioptera: Mesoraphidiidae: Nanoraphidiini) from the mid Cretaceous of Myanmar</mods:title>
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<mods:namePart id="CAA596903A064A152786A3C725D3AD46">Liu, Xingyue</mods:namePart>
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<mods:namePart id="D74200F28978BAA1D2A2E17CEA66C819">Lu, Xiumei</mods:namePart>
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<mods:namePart id="04F76A6BFBC030A09F51BE11B4A48BD1">Zhang, Weiwei</mods:namePart>
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<treatment id="03FBD81AAF20FFADFF58FBDCAC44A096" ID-DOI="http://doi.org/10.5281/zenodo.6055085" ID-GBIF-Taxon="119568748" ID-Zenodo-Dep="6055085" LSID="urn:lsid:plazi:treatment:03FBD81AAF20FFADFF58FBDCAC44A096" httpUri="http://treatment.plazi.org/id/03FBD81AAF20FFADFF58FBDCAC44A096" lastPageNumber="316" pageId="15" pageNumber="316">
<subSubSection id="C3483A87AF20FFADFF58FBDCAE35A373" pageId="15" pageNumber="316" type="nomenclature">
<paragraph id="8BED690CAF20FFADFF58FBDCAD28A312" blockId="15.[151,560,1052,1111]" box="[151,560,1052,1078]" pageId="15" pageNumber="316">
<heading id="D0A5DE60AF20FFADFF58FBDCAD28A312" bold="true" box="[151,560,1052,1078]" fontSize="11" level="1" pageId="15" pageNumber="316" reason="1">
<emphasis id="B926B51EAF20FFADFF58FBDCAD28A312" bold="true" box="[151,560,1052,1078]" pageId="15" pageNumber="316">
Genus
<taxonomicName id="4C52128FAF20FFADFF21FBDCAD28A312" ID-CoL="5XHB" authority="Engel, 2002" authorityName="Engel" authorityYear="2002" box="[238,560,1052,1078]" class="Insecta" family="Mesoraphidiidae" genus="Nanoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B926B51EAF20FFADFF21FBDCAE80A312" bold="true" box="[238,408,1052,1078]" italics="true" pageId="15" pageNumber="316">Nanoraphidia</emphasis>
Engel, 2002
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BED690CAF20FFADFF58FBFEAE35A373" blockId="15.[151,560,1052,1111]" box="[151,301,1086,1111]" pageId="15" pageNumber="316">
(
<figureCitation id="13697589AF20FFADFF50FBFEAE3EA373" box="[159,294,1086,1111]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStartId-0="18.[151,250,1906,1928]" captionStartId-1="19.[151,250,1126,1148]" captionTargetBox-0="[165,1422,1025,1879]" captionTargetBox-1="[167,1436,194,1092]" captionTargetId-0="figure@18.[160,1426,1021,1885]" captionTargetId-1="figure@19.[164,1449,193,1101]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURE 12. Nanoraphidia electroburmica Engel, EMTG BA- 001202. A. Habitus in dorsal view; B. Drawing of fore- and hindwing; C. Drawing of female genitalia, lateral view; D. Drawing of female genitalia, dorsal view. T: tergum; S: sternum; gst: gonostylus; op: ovipositor; e: ectoproct. Scale bar = 1.0 mm." captionText-1="FIGURE 13. Nanoraphidia electroburmica Engel. A. EMTG BA- 002164, habitus (left: male; right: female); B. PCXJ BA- 0 0 0 4, habitus in dorsal view; C. Same specimen, male genitalia in dorsal view; D. Same specimen, drawing of male genitalia in dorsal view; E. Same specimen, male genitalia in ventral view; F. Same specimen, drawing of male genitalia in ventral view. Arrows indicating putative gonocoxites 9 or gonocoxites + gonostyli 9. T: tergum; e: ectoproct. Scale bar = 1.0 mm (A B) and 0.5 mm (C F)." httpUri-0="https://zenodo.org/record/264771/files/figure.png" httpUri-1="https://zenodo.org/record/264772/files/figure.png" pageId="15" pageNumber="316">Figs. 1213</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF20FFADFF58FB47ADEDA3B8" box="[151,757,1158,1180]" pageId="15" pageNumber="316" type="materials_examined">
<paragraph id="8BED690CAF20FFADFF58FB47ADEDA3B8" blockId="15.[151,757,1158,1180]" box="[151,757,1158,1180]" pageId="15" pageNumber="316">
<typeStatus id="54E9D7AEAF20FFADFF58FB47AFD1A3B8" box="[151,201,1159,1180]" pageId="15" pageNumber="316">Type</typeStatus>
species:
<taxonomicName id="4C52128FAF20FFADFEE9FB46ADE9A3B8" authority="Engel, 2002: 3" authorityName="Engel" authorityPageNumber="3" authorityYear="2002" box="[294,753,1158,1180]" class="Insecta" family="Mesoraphidiidae" genus="Nanoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="electroburmica">
<emphasis id="B926B51EAF20FFADFEE9FB46AD4CA3B8" box="[294,596,1158,1180]" italics="true" pageId="15" pageNumber="316">Nanoraphidia electroburmica</emphasis>
Engel, 2002: 3
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF20FFADFF58FB09ABB4A102" pageId="15" pageNumber="316" type="diagnosis">
<paragraph id="8BED690CAF20FFADFF58FB09ABB4A102" blockId="15.[151,1436,1225,1970]" pageId="15" pageNumber="316">
<emphasis id="B926B51EAF20FFADFF58FB09AE09A3C6" bold="true" box="[151,273,1225,1250]" pageId="15" pageNumber="316">Diagnosis.</emphasis>
Minute size. Head ovoid, with portion posterior to compound eyes nearly as long as eye length and tapering caudad, and with clypeus slightly shorter than compound eye length; three small ocelli present on posterior half of head; antenna nearly as long as head plus prothorax, with 2030 flagellomeres. Pro- and mesotibiae distinctly swollen, metatibia feebly swollen, all tibiae lacking tibial spur. Forewing: Costal space distinctly broadened; pterostigma elongate, ~1/3× wing length, without pterostigmal crossvein, proximally closed by a crossvein and distally closed by a veinlet of RA; Scp terminating into C slightly distad wing midpoint; posterior veinlet of RA zig-zagged with stem of RA;
<collectingCountry id="F345299CAF20FFADFCCBFA62AC3EA29D" box="[772,806,1442,1465]" name="Poland" pageId="15" pageNumber="316">RP</collectingCountry>
distally bifurcated; MA distally trifurcated; two radial cells, one discal cell, and two discoidal cells present. Hindwing: Configuration of pterostigmal area same to that of male; stem of MA proximally fused with MP; a slightly oblique proximal mp-cua present; two radial cells, one discal cell, and one discoidal cell present; A1 proximally fused with CuP for a short distance.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF20FFADFF08F9F1AC44A096" pageId="15" pageNumber="316" type="discussion">
<paragraph id="8BED690CAF20FFADFF08F9F1AC30A1DA" blockId="15.[151,1436,1225,1970]" pageId="15" pageNumber="316">
<emphasis id="B926B51EAF20FFADFF08F9F1AE23A16E" bold="true" box="[199,315,1585,1610]" pageId="15" pageNumber="316">Remarks.</emphasis>
This genus appears to be closely related to
<taxonomicName id="4C52128FAF20FFADFCE0F9F1ACE2A16D" box="[815,1018,1585,1609]" class="Insecta" family="Mesoraphidiidae" genus="Cantabroraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B926B51EAF20FFADFCE0F9F1ACE2A16D" box="[815,1018,1585,1609]" italics="true" pageId="15" pageNumber="316">Cantabroraphidia</emphasis>
</taxonomicName>
Pérez-de la Fuente, Nel, Peñalver &amp; Delclòs, 2010 and
<emphasis id="B926B51EAF20FFADFEB1F995AD57A149" box="[382,591,1621,1645]" italics="true" pageId="15" pageNumber="316">Amarantoraphidia</emphasis>
Pérez-de la Fuente, Peñalver, Delclòs &amp; Engel, 2012 from the Early Cretaceous amber of
<collectingCountry id="F345299CAF20FFADFE66F9B9AEF1A1B6" box="[425,489,1657,1682]" name="Spain" pageId="15" pageNumber="316">Spain</collectingCountry>
in having generally similar cephalic feature and wing venation. However,
<taxonomicName id="4C52128FAF20FFADFF58F95DAE2CA191" box="[151,308,1693,1717]" class="Insecta" family="Mesoraphidiidae" genus="Nanoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B926B51EAF20FFADFF58F95DAE2CA191" box="[151,308,1693,1717]" italics="true" pageId="15" pageNumber="316">Nanoraphidia</emphasis>
</taxonomicName>
can be distinguished from the latter two genera by the distally trifurcate MA (MA distally bifurcated in
<taxonomicName id="4C52128FAF20FFADFF7AF901AE98A1FD" box="[181,384,1729,1753]" class="Insecta" family="Mesoraphidiidae" genus="Cantabroraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B926B51EAF20FFADFF7AF901AE98A1FD" box="[181,384,1729,1753]" italics="true" pageId="15" pageNumber="316">Cantabroraphidia</emphasis>
</taxonomicName>
and
<emphasis id="B926B51EAF20FFADFE75F901AD93A1FD" box="[442,651,1729,1753]" italics="true" pageId="15" pageNumber="316">Amarantoraphidia</emphasis>
) and the presence of only one forewing discoidal cell (two discoidal cells present in
<taxonomicName id="4C52128FAF20FFADFE89F925AD09A1D9" box="[326,529,1765,1789]" class="Insecta" family="Mesoraphidiidae" genus="Cantabroraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B926B51EAF20FFADFE89F925AD09A1D9" box="[326,529,1765,1789]" italics="true" pageId="15" pageNumber="316">Cantabroraphidia</emphasis>
</taxonomicName>
and
<emphasis id="B926B51EAF20FFADFD88F925AC0FA1D9" box="[583,791,1765,1789]" italics="true" pageId="15" pageNumber="316">Amarantoraphidia</emphasis>
).
</paragraph>
<paragraph id="8BED690CAF20FFADFF08F8C9AC44A096" blockId="15.[151,1436,1225,1970]" pageId="15" pageNumber="316">
<taxonomicName id="4C52128FAF20FFADFF08F8C9ACA5A006" authority="Jepson, Ansorge &amp; Jarzemborski, 2011" authorityName="Jepson, Ansorge &amp; Jarzemborski" authorityYear="2011" box="[199,957,1801,1826]" class="Insecta" family="Mesoraphidiidae" genus="Nanoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="lithographica">
<emphasis id="B926B51EAF20FFADFF08F8C9AD1BA005" box="[199,515,1801,1825]" italics="true" pageId="15" pageNumber="316">Nanoraphidia lithographica</emphasis>
Jepson, Ansorge &amp; Jarzemborski, 2011
</taxonomicName>
from the Early Cretaceous of
<collectingCountry id="F345299CAF20FFADFAC2F8C9AA55A006" box="[1293,1357,1801,1826]" name="Spain" pageId="15" pageNumber="316">Spain</collectingCountry>
differs from
<taxonomicName id="4C52128FAF20FFADFF15F8EEAEB2A061" box="[218,426,1837,1861]" class="Insecta" family="Mesoraphidiidae" genus="Nanoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="electroburmica">
<emphasis id="B926B51EAF20FFADFF15F8EEAEB2A061" box="[218,426,1837,1861]" italics="true" pageId="15" pageNumber="316">N. electroburmica</emphasis>
</taxonomicName>
by the forewing RA with posterior branch continuous with its stem and the distally bifurcated MA, and it more resembles the species of
<emphasis id="B926B51EAF20FFADFCC3F891ACC4A04D" box="[780,988,1873,1897]" italics="true" pageId="15" pageNumber="316">Amarantoraphidia</emphasis>
and
<taxonomicName id="4C52128FAF20FFADFBD4F891ABFEA04D" box="[1051,1254,1873,1897]" class="Insecta" family="Mesoraphidiidae" genus="Cantabroraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B926B51EAF20FFADFBD4F891ABFEA04D" box="[1051,1254,1873,1897]" italics="true" pageId="15" pageNumber="316">Cantabroraphidia</emphasis>
</taxonomicName>
from the Early Cretaceous amber of
<collectingCountry id="F345299CAF20FFADFE45F8B5AED5A0AA" box="[394,461,1909,1934]" name="Spain" pageId="15" pageNumber="316">Spain</collectingCountry>
. However, the latter two genera possess two forewing discoidal cells. At any rate, the generic placement of
<taxonomicName id="4C52128FAF20FFADFE46F85AAD59A095" box="[393,577,1945,1969]" class="Insecta" family="Mesoraphidiidae" genus="Nanoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="lithographica">
<emphasis id="B926B51EAF20FFADFE46F85AAD59A095" box="[393,577,1945,1969]" italics="true" pageId="15" pageNumber="316">N. lithographica</emphasis>
</taxonomicName>
should be re-considered.
</paragraph>
</subSubSection>
</treatment>
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