288 lines
36 KiB
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288 lines
36 KiB
XML
<document id="8B583003896F33D2C4587AF5F2AA9743" ID-DOI="10.1163/18759866-20191416" ID-GBIF-Dataset="6388d839-a72d-4598-a141-053897bfc900" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. & Köhler, Frank" docDate="2019" docId="03F32A40FFE3B13D51A59A03F053FF12" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.3:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="3" docTitle="Fastosarion helenkingae Scott 1995" docType="treatment" docVersion="10" lastPageNumber="412" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="409" updateTime="1731578815876" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1C5B71BA73ECD86869BE069C79968873">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:namePart id="28D6140494D8BBB70B48C155EDDD435A">Hyman, Isabel T.</mods:namePart>
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<mods:affiliation id="902AB6BD2F38396EB0F759166ED96A43">Australian Museum, 1 William St, Sydney NSW 2010, Australia Isabel. Hyman @ austmus. gov. au</mods:affiliation>
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<mods:namePart id="35CA3BEF8B8B292A474E8DFB7D6D91E7">Köhler, Frank</mods:namePart>
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<mods:affiliation id="E4A805F2FCB40F83D957738A540B9875">Australian Museum, 1 William St, Sydney NSW 2010, Australia</mods:affiliation>
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<mods:title id="FD14A647AA7B965A18D21D7B4AFA9479">Contributions to Zoology</mods:title>
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<paragraph id="8BE59B56FFE3B13A51A59A03F04FF9D6" blockId="58.[671,1080,1554,1581]" box="[671,1080,1554,1581]" pageId="58" pageNumber="409">
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<taxonomicName id="4C5AE0D5FFE3B13A51A59A03F04FF9D6" ID-CoL="3DVWC" ID-ENA="2658019" authority="Scott, 1995" authorityName="Scott" authorityYear="1995" box="[671,1080,1554,1581]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="409" phylum="Mollusca" rank="species" species="helenkingae">
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<emphasis id="B92E4744FFE3B13A51A59A03F04FF9D6" bold="true" box="[671,1080,1554,1581]" pageId="58" pageNumber="409">
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<emphasis id="B92E4744FFE3B13A51A59A03F7CDF9D6" bold="true" box="[671,954,1554,1581]" italics="true" pageId="58" pageNumber="409">Fastosarion helenkingae</emphasis>
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<bibRefCitation id="EFCBE6A7FFE3B13A50859A02F04FF9D6" author="Scott, B." box="[959,1080,1554,1581]" pageId="58" pageNumber="409" pagination="69 - 80" refId="ref42653" refString="Scott, B. (1995) Redescription of Fastosarion superba (Cox, 1871) with descriptions of two new species of Fastosarion from northern Queensland (Pulmonata: Helicarionidae). Molluscan Res., 16, 69 - 80." type="journal article" year="1995">Scott, 1995</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C340C8DDFFE3B13A519F9A27F794F9AA" box="[677,995,1591,1617]" pageId="58" pageNumber="409" type="description">
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<paragraph id="8BE59B56FFE3B13A519F9A27F794F9AA" blockId="58.[676,1187,1591,1761]" box="[677,995,1591,1617]" pageId="58" pageNumber="409">
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<figureCitation id="136187D3FFE3B13A519F9A27F69BF9AA" box="[677,748,1591,1617]" captionStart="FIGURE 6" captionStartId="49.[98,171,1309,1330]" captionTargetBox="[98,1151,188,1277]" captionTargetId="figure-7@49.[98,1157,188,1277]" captionTargetPageId="49" captionText="FIGURE 6 Distribution of Clade A species. ▲ = F.superbus, ◻ = F.aquavitae, ♢ = F.brazieri, ◆ = F.comerfordae, ●= F.griseolus, ▽ = F.helenkingae, █ = F.paluma." figureDoi="http://doi.org/10.5281/zenodo.8343073" httpUri="https://zenodo.org/record/8343073/files/figure.png" pageId="58" pageNumber="409">Figs. 6</figureCitation>
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,
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<figureCitation id="136187D3FFE3B13A51CC9A27F74EF9AA" box="[758,825,1591,1617]" captionStart="FIGURE 7" captionStartId="50.[129,202,1279,1300]" captionTargetBox="[134,1183,193,1242]" captionTargetId="figure-9@50.[129,1188,188,1247]" captionTargetPageId="50" captionText="FIGURE 7 Shells of Clade A species. A. Fastosarion superbus, MO77446.B. Fastosarion aquavitae, MO50851.C. Fastosarion aquavitae, MO20228 (type locality of F.ameyi). D. Fastosarion brazieri, AM C.553533.E. Fastosarion comerfordae, MO63985.F.Fastosarion griseolus, MO55568.G.Fastosarion helenkingae, NMV F219324.H. Fastosarion helenkingae, AM C.559842 (type locality of F.schelli). I. Fastosarion paluma, MO63983.J.Fastosarion paluma, MO48024 (Hinchinbrook Is).Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343075" httpUri="https://zenodo.org/record/8343075/files/figure.png" pageId="58" pageNumber="409">7G–H</figureCitation>
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,
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<figureCitation id="136187D3FFE3B13A50799A27F706F9AA" box="[835,881,1591,1617]" captionStart="FIGURE 8" captionStartId="51.[98,171,1353,1374]" captionTargetBox="[98,1157,188,1321]" captionTargetId="figure-6@51.[98,1157,188,1321]" captionTargetPageId="51" captionText="FIGURE 8 Fastosarion live images (Clade A).A.Fastosarion superbus, MO46340, Mt Dryander (QM image). B.F.aquavitae, NMV F230751, Brandy Creek (A. Moussalli, NMV).C. F.brazieri, AM C.553531, Cape Tribulation (A.Moussalli, NMV).D. F.brazieri, AM C.525003, Mt Lewis.E.F.comerfordae, Broken River NP (Kackland). F.F.paluma, Paluma (QM image). G. F.griseolus, AM C.512514,Byfield SF.H. F.griseolus, AM C.512615,Burleigh Head NP. I. F.helenkingae, NMV F219324,Bowling Green NP (A. Moussalli, NMV).J. F.helenkingae, AM C.559842, Magnetic Island (type locality of F.schelli)." figureDoi="http://doi.org/10.5281/zenodo.8343077" httpUri="https://zenodo.org/record/8343077/files/figure.png" pageId="58" pageNumber="409">8I–J</figureCitation>
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,
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<figureCitation id="136187D3FFE3B13A50419A27F7B4F9AA" box="[891,963,1591,1617]" captionStart="FIGURE 12" captionStartId="56.[129,202,1411,1432]" captionTargetBox="[129,1188,188,1379]" captionTargetId="figure-8@56.[129,1188,188,1379]" captionTargetPageId="56" captionText="FIGURE 12 Spermatophores of Clade A species. A–B. Fastosarion brazieri, MO66680.C–D.Fastosarion comerfordae, MO63985. E-F.F.helenkingae, MO47971." figureDoi="http://doi.org/10.5281/zenodo.8343085" httpUri="https://zenodo.org/record/8343085/files/figure.png" pageId="58" pageNumber="409">12C–D</figureCitation>
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,
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<figureCitation id="136187D3FFE3B13A50F79A27F794F9AA" box="[973,995,1591,1617]" captionStart="FIGURE 16" captionStartId="62.[129,202,1714,1735]" captionTargetBox="[161,1154,170,1682]" captionTargetId="figure-52@62.[161,1154,170,1682]" captionTargetPageId="62" captionText="FIGURE 16 Genitalia of F.helenkingae. A, C, E. MO47971, Mt Elliot. B, D. MO16161, Magnetic Is. A–B, Reproductive system.Arrows show the junction of the penis and epiphallus.C–D, Penis interior.E.Spermatophore. Scale bars 1 mm (C–E), 2 mm (A–B)." figureDoi="http://doi.org/10.5281/zenodo.8343093" httpUri="https://zenodo.org/record/8343093/files/figure.png" pageId="58" pageNumber="409">16</figureCitation>
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</paragraph>
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<subSubSection id="C340C8DDFFE3B13A519F9A4BF7C0F91B" pageId="58" pageNumber="409" type="reference_group">
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<paragraph id="8BE59B56FFE3B13A519F9A4BF01BF962" blockId="58.[676,1187,1591,1761]" pageId="58" pageNumber="409">
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<treatmentCitationGroup id="AB4ABC78FFE3B13A519F9A4BF01BF962" pageId="58" pageNumber="409">
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<treatmentCitation id="0AFBBD47FFE3B13A519F9A4BF03AF98E" author="Scott, B." box="[677,1101,1626,1653]" page="75" pageId="58" pageNumber="409" year="1995">
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<taxonomicName id="4C5AE0D5FFE3B13A519F9A4BF03AF98E" ID-CoL="3DVWC" authority="Scott, 1995: 75" authorityName="Scott" authorityPageNumber="75" authorityYear="1995" box="[677,1101,1626,1653]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="409" phylum="Mollusca" rank="species" species="helenkingae">
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<emphasis id="B92E4744FFE3B13A519F9A4BF7DAF98E" box="[677,941,1627,1653]" italics="true" pageId="58" pageNumber="409">Fastosarion helenkingae</emphasis>
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<bibRefCitation id="EFCBE6A7FFE3B13A50899A4BF03AF98E" author="Scott, B." box="[947,1101,1626,1653]" pageId="58" pageNumber="409" pagination="69 - 80" refId="ref42653" refString="Scott, B. (1995) Redescription of Fastosarion superba (Cox, 1871) with descriptions of two new species of Fastosarion from northern Queensland (Pulmonata: Helicarionidae). Molluscan Res., 16, 69 - 80." type="journal article" year="1995">Scott, 1995: 75</bibRefCitation>
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</taxonomicName>
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, figs. 1c,
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<figureCitation id="136187D3FFE3B13A519E9A6FF6C5F962" box="[676,690,1663,1689]" captionStart="FIGURE 2" captionStartId="31.[98,171,1692,1713]" captionTargetBox="[98,1125,170,1660]" captionTargetId="figure-6@31.[98,1125,169,1660]" captionText="FIGURE 2 Majority-rule consensus tree based on Bayesian analysis of the concatenated data set of fragments of the mitochondrial genes 16S and COI. Ambiguous alignment sites in 16S removed using MAFFT. Numbers on branches indicate posterior probabilities. Scale bar indicating 40% of modelled sequence divergence. Shells shown to scale." figureDoi="http://doi.org/10.5281/zenodo.8343065" httpUri="https://zenodo.org/record/8343065/files/figure.png" pageId="58" pageNumber="409">2</figureCitation>
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, 5b, 6;
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<treatmentCitation id="0AFBBD47FFE3B13A51C79A6EF042F963" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[765,1077,1662,1689]" page="306" pageId="58" pageNumber="409" year="2010">
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<bibRefCitation id="EFCBE6A7FFE3B13A51C79A6EF042F963" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[765,1077,1662,1689]" pageId="58" pageNumber="409" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010: 306–307</bibRefCitation>
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</treatmentCitation>
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, 329.
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</paragraph>
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<paragraph id="8BE59B56FFE3B13A519F9AB3F7C0F91B" blockId="58.[676,1187,1591,1761]" pageId="58" pageNumber="409">
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<treatmentCitationGroup id="AB4ABC78FFE3B13A519F9AB3F7C0F91B" pageId="58" pageNumber="409">
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<taxonomicName id="4C5AE0D5FFE3B13A519F9AB3F06CF946" ID-CoL="7RBRX" authority="Stanisic, 2010" authorityName="Stanisic" authorityYear="2010" box="[677,1051,1698,1725]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="58" pageNumber="409" phylum="Mollusca" rank="species" species="schelli">
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<emphasis id="B92E4744FFE3B13A519F9AB3F703F947" box="[677,884,1698,1724]" italics="true" pageId="58" pageNumber="409">Fastosarion schelli</emphasis>
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<bibRefCitation id="EFCBE6A7FFE3B13A50BB9AB2F06CF946" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[897,1051,1698,1725]" pageId="58" pageNumber="409" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">
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Stanisic,
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<quantity id="4CA236B3FFE3B13A50D19AB3F06CF946" box="[1003,1051,1699,1725]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="58" pageNumber="409" unit="in" value="2010.0">2010</quantity>
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</bibRefCitation>
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in
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<treatmentCitation id="0AFBBD47FFE3B13A57719AB2F7F7F91B" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." page="306" pageId="58" pageNumber="409" year="2010">
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<bibRefCitation id="EFCBE6A7FFE3B13A57719AB2F7F7F91B" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." pageId="58" pageNumber="409" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010: 306–307</bibRefCitation>
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, 328.
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</paragraph>
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<caption id="DF25CBDEFFE2B13B5358988BF023FADE" ID-DOI="http://doi.org/10.5281/zenodo.8343089" ID-Zenodo-Dep="8343089" httpUri="https://zenodo.org/record/8343089/files/figure.png" pageId="59" pageNumber="410" startId="59.[98,171,1179,1200]" targetBox="[98,1157,188,1147]" targetPageId="59" targetType="figure">
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<paragraph id="8BE59B56FFE2B13B5358988BF023FADE" blockId="59.[98,1142,1178,1318]" pageId="59" pageNumber="410">
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FIGURE 14 Shell microsculpture. A–C.
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<taxonomicName id="4C5AE0D5FFE2B13B52EE988BF6CBFB4B" authority=", MO" authorityName="MO" box="[468,700,1178,1200]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="410" phylum="Mollusca" rank="species" species="griseolus">
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<emphasis id="B92E4744FFE2B13B52EE988BF6E7FB54" box="[468,656,1178,1200]" italics="true" pageId="59" pageNumber="410">Fastosarion griseolus</emphasis>
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, MO
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</taxonomicName>
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55568 (A = protoconch, B = early teleoconch, C = late teleoconch). D.
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<taxonomicName id="4C5AE0D5FFE2B13B528E98A8F620FB36" baseAuthorityName="Stanisic" baseAuthorityYear="2010" box="[436,599,1208,1229]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="410" phylum="Mollusca" rank="species" species="alyssa">
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<emphasis id="B92E4744FFE2B13B528E98A8F620FB36" box="[436,599,1208,1229]" italics="true" pageId="59" pageNumber="410">Fastosarion alyssa</emphasis>
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</taxonomicName>
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, F230784 (protoconch). E–F.
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<taxonomicName id="4C5AE0D5FFE2B13B505A98A8F034FB35" authority=", MO" authorityName="MO" box="[864,1091,1208,1230]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="410" phylum="Mollusca" rank="species" species="minerva">
|
||
<emphasis id="B92E4744FFE2B13B505A98A8F060FB36" box="[864,1047,1208,1229]" italics="true" pageId="59" pageNumber="410">Fastosarion minerva</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
65418 (E = protoconch, F = teleoconch). G–I.
|
||
<taxonomicName id="4C5AE0D5FFE2B13B510698C6F747FB10" authority=", MO" authorityName="MO" box="[572,816,1237,1259]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="410" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FFE2B13B510698C6F773FB11" box="[572,772,1237,1259]" italics="true" pageId="59" pageNumber="410">Fastosarion papillosus</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
78896 (G = protoconch, H = early teleoconch, I = late teleoconch). J–L.
|
||
<taxonomicName id="4C5AE0D5FFE2B13B511698E3F77BFAF3" authority=", MO" authorityName="MO" box="[556,780,1266,1288]" class="Gastropoda" family="Helicarionidae" genus="Stanisicarion" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="410" phylum="Mollusca" rank="species" species="aquila">
|
||
<emphasis id="B92E4744FFE2B13B511698E3F697FAFC" box="[556,736,1266,1288]" italics="true" pageId="59" pageNumber="410">Stanisicarion aquila</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
49825 (J = protoconch, K = early teleoconch, L = late teleoconch). Scale bars 20 µm (A, D, E, I), 100 µm (B, F, L), 50 µm (C, G-H, J-K).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C340C8DDFFE2B13B53669993F7A6FA66" pageId="59" pageNumber="410" type="materials_examined">
|
||
<paragraph id="8BE59B56FFE2B13B53669993F547FA66" blockId="59.[92,609,1411,1761]" box="[92,304,1411,1437]" pageId="59" pageNumber="410">Material examined</paragraph>
|
||
<paragraph id="8BE59B56FFE2B13B535899B8F64DFA1D" blockId="59.[92,609,1411,1761]" pageId="59" pageNumber="410">
|
||
<emphasis id="B92E4744FFE2B13B535899B8F4D6FA3A" box="[98,161,1448,1473]" italics="true" pageId="59" pageNumber="410">Types</emphasis>
|
||
:
|
||
<materialsCitation id="3B32910BFFE2B13B538299B7F64DFA1D" ID-GBIF-Occurrence="4413347308" collectorName="Bowling Green Bay NP" latitude="-19.5" location="Bowling Green Bay" longLatPrecision="1271" longitude="146.96666" municipality="Mt Elliot" pageId="59" pageNumber="410" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54E125F4FFE2B13B538299B7F554FA3A" box="[184,291,1447,1473]" pageId="59" pageNumber="410" type="holotype">Holotype</typeStatus>
|
||
: MTQ MO40184 (
|
||
<collectorName id="26AFFE80FFE2B13B513999B7F48BFA1E" pageId="59" pageNumber="410">
|
||
<location id="8E85CD8DFFE2B13B513999B7F4A1FA1E" LSID="urn:lsid:plazi:treatment:03F32A40FFE3B13D51A59A03F053FF12:8E85CD8DFFE2B13B513999B7F4A1FA1E" latitude="-19.5" longLatPrecision="1271" longitude="146.96666" municipality="Mt Elliot" name="Bowling Green Bay" pageId="59" pageNumber="410">Bowling Green Bay</location>
|
||
NP
|
||
</collectorName>
|
||
,
|
||
<collectingMunicipality id="6B81012CFFE2B13B523C99DBF51DFA1E" box="[262,362,1483,1509]" pageId="59" pageNumber="410">Mt Elliot</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE6EFD91FFE2B13B524999DBF5B4FA1E" box="[371,451,1483,1509]" degrees="19" direction="south" minutes="30" orientation="latitude" pageId="59" pageNumber="410" precision="925" value="-19.5">19°30'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6EFD91FFE2B13B52F699DBF658FA1D" box="[460,559,1483,1510]" degrees="146" direction="east" minutes="58" orientation="longitude" pageId="59" pageNumber="410" precision="925" value="146.96666">146°58'E</geoCoordinate>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FFE2B13B535899FFF4A1F9D5" blockId="59.[92,609,1411,1761]" pageId="59" pageNumber="410">
|
||
<typeStatus id="54E125F4FFE2B13B535899FFF4A6F9F2" box="[98,209,1519,1545]" pageId="59" pageNumber="410" type="paratype">Paratypes</typeStatus>
|
||
: MTQ MO40185-40191 (same data as holotype).
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FFE2B13B53589A27F4A9F962" blockId="59.[92,609,1411,1761]" pageId="59" pageNumber="410">
|
||
<typeStatus id="54E125F4FFE2B13B53589A27F4BDF9AA" box="[98,202,1591,1617]" pageId="59" pageNumber="410" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5AE0D5FFE2B13B53D69A27F5DBF9AB" authority=": QM MO" authorityName="QM MO" box="[236,428,1590,1617]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="410" phylum="Mollusca" rank="species" species="schelli">
|
||
<emphasis id="B92E4744FFE2B13B53D69A27F53FF9AA" box="[236,328,1591,1617]" italics="true" pageId="59" pageNumber="410">F. schelli</emphasis>
|
||
: QM MO
|
||
</taxonomicName>
|
||
14106 (Magnetic I., MEQ,
|
||
<geoCoordinate id="EE6EFD91FFE2B13B53FB9A4BF565F98E" box="[193,274,1627,1653]" degrees="19" direction="south" minutes="08" orientation="latitude" pageId="59" pageNumber="410" precision="925" value="-19.133333">19°08’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6EFD91FFE2B13B52269A4AF500F98E" box="[284,375,1626,1653]" degrees="146" direction="east" minutes="51" orientation="longitude" pageId="59" pageNumber="410" precision="925" value="146.85">146°51’E</geoCoordinate>
|
||
,
|
||
<quantity id="4CA236B3FFE2B13B52BB9A4BF5B0F98E" box="[385,455,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="59" pageNumber="410" unit="m" value="200.0">200 m</quantity>
|
||
,
|
||
<date id="FFE4BD96FFE2B13B52E89A4BF62AF98E" box="[466,605,1627,1653]" pageId="59" pageNumber="410" value="1980-09-23">23 Sept 1980</date>
|
||
, J. Stanisic).
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FFE2B13B53589AB3F5ECF91A" blockId="59.[92,609,1411,1761]" pageId="59" pageNumber="410">
|
||
<materialsCitation id="3B32910BFFE2B13B53589AB3F545F91B" ID-GBIF-Occurrence="4413347329" collectionCode="QM, AM" pageId="59" pageNumber="410" specimenCode="MO16161, MO78893" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54E125F4FFE2B13B53589AB3F4B8F946" box="[98,207,1699,1725]" pageId="59" pageNumber="410" type="paratype">Paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5AE0D5FFE2B13B521A9AB3F5E7F947" box="[288,400,1698,1724]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="59" pageNumber="410" phylum="Mollusca" rank="species" species="schelli">
|
||
<emphasis id="B92E4744FFE2B13B521A9AB3F5E7F947" box="[288,400,1698,1724]" italics="true" pageId="59" pageNumber="410">F. schelli</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<collectionCode id="ED4B0393FFE2B13B52899AB2F5A9F947" box="[435,478,1698,1724]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="59" pageNumber="410" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBFC332DFFE2B13B52C19AB2F62AF947" box="[507,605,1698,1724]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="59" pageNumber="410" type="Museum">MO16161</specimenCode>
|
||
,
|
||
<collectionCode id="ED4B0393FFE2B13B53589AD6F4FAF91B" box="[98,141,1734,1760]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="59" pageNumber="410" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBFC332DFFE2B13B53A99AD6F488F91B" box="[147,255,1734,1760]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="59" pageNumber="410" type="Museum">MO78893</specimenCode>
|
||
,
|
||
<collectionCode id="ED4B0393FFE2B13B52339AD6F545F91B" box="[265,306,1734,1760]" country="Australia" name="Australian Museum" pageId="59" pageNumber="410" type="Museum">AM</collectionCode>
|
||
</materialsCitation>
|
||
C.125918.
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FFE2B13B51BC9994F7A6FA66" blockId="59.[646,977,1411,1438]" box="[646,977,1411,1438]" pageId="59" pageNumber="410">
|
||
<emphasis id="B92E4744FFE2B13B51BC9994F73AFA66" box="[646,845,1411,1437]" italics="true" pageId="59" pageNumber="410">Non-type material</emphasis>
|
||
: See
|
||
<tableCitation id="C6D8AEEDFFE2B13B50BE9993F7BBFA66" box="[900,972,1411,1437]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="59" pageNumber="410" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340C8DDFFE2B13D51BA99DBF4CAFDAB" lastPageId="61" lastPageNumber="412" pageId="59" pageNumber="410" type="diagnosis">
|
||
<paragraph id="8BE59B56FFE2B13B51BA99DBF69AFA1E" blockId="59.[640,1158,1483,1760]" box="[640,749,1483,1509]" pageId="59" pageNumber="410">Diagnosis</paragraph>
|
||
<paragraph id="8BE59B56FFE2B13C51BC99FFF5E5F962" blockId="59.[640,1158,1483,1760]" lastBlockId="60.[129,640,1555,1760]" lastPageId="60" lastPageNumber="411" pageId="59" pageNumber="410">
|
||
<emphasis id="B92E4744FFE2B13B51BC99FFF707F9F2" box="[646,880,1519,1545]" italics="true" pageId="59" pageNumber="410">External morphology</emphasis>
|
||
: Shell (fig. 7G–H) medium-sized (
|
||
<quantity id="4CA236B3FFE2B13B51C19A03F7E3F9D6" box="[763,916,1555,1582]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" metricValueMax="1.85" metricValueMin="1.43" pageId="59" pageNumber="410" unit="mm" value="16.4" valueMax="18.5" valueMin="14.3">14.3–18.5 mm</quantity>
|
||
), amber, transparent, 3.2–3.7 whorls, depressedly subglobose with a low spire. Body (fig. 8I–J)
|
||
<quantity id="4CA236B3FFE2B13B50909A4BF78FF98E" box="[938,1016,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="59" pageNumber="410" unit="mm" value="30.0">30 mm</quantity>
|
||
long, speckled brownish cream to grey, darker on head and tail; sometimes with a reddish sole. Shell lappets and mantle lobes large, each lappet with a ridged dark line close to and parallel with lower edge. Dark line along mantle collar. Tail weakly keeled, keel paler, slime network moderately strong.
|
||
</paragraph>
|
||
<caption id="DF25CBDEFFE5B13C53BB999AF7E1FA46" ID-DOI="http://doi.org/10.5281/zenodo.8343091" ID-Zenodo-Dep="8343091" httpUri="https://zenodo.org/record/8343091/files/figure.png" pageId="60" pageNumber="411" startId="60.[129,202,1418,1439]" targetBox="[139,1187,188,1385]" targetPageId="60" targetType="figure">
|
||
<paragraph id="8BE59B56FFE5B13C53BB999AF7E1FA46" blockId="60.[129,1183,1417,1469]" pageId="60" pageNumber="411">
|
||
FIGURE 15 Genitalia of
|
||
<taxonomicName id="4C5AE0D5FFE5B13C5251999AF58DFA64" authority=", MO" authorityName="MO" box="[363,506,1417,1439]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="60" pageNumber="411" phylum="Mollusca" rank="species" species="griseolus">
|
||
<emphasis id="B92E4744FFE5B13C5251999AF5BAFA65" box="[363,461,1417,1439]" italics="true" pageId="60" pageNumber="411">F.griseolus</emphasis>
|
||
, MO
|
||
</taxonomicName>
|
||
55568, Bobby Range. A. Reproductive system. Arrow shows the junction of the penis and epiphallus. B. Penis interior. Scale bars 1 mm (B), 2 mm (A).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE59B56FFE5B13D539A9AB2F4CAFDAB" blockId="60.[129,640,1555,1760]" lastBlockId="61.[98,609,171,592]" lastPageId="61" lastPageNumber="412" pageId="60" pageNumber="411">
|
||
<emphasis id="B92E4744FFE5B13C539A9AB2F52AF947" box="[160,349,1698,1724]" italics="true" pageId="60" pageNumber="411">Genital anatomy:</emphasis>
|
||
Genitalia (figs. 12C–D, 16) with medium length vagina; bursa copulatrix moderately short, duct distinct, bursa spherical to oval. Penis long, slender, slightly swollen proximally, internally with one complete longitudinal pilaster and one short longitudinal pilaster proximally, internal wall sculptured with pustules; 60–80% of penis contained in penial tunica. Penis slightly longer than epiphallus, slightly narrower in diameter; epiphallus enters penis through a swollen ring; epiphallus 2 much longer than epiphallus 1; epiphallic caecum short; flagellum short, slender, terminal portion fused into loop. Spermatophore simple, tail-pipe mostly smooth, towards terminal end sculptured with small teeth pointing towards capsule, followed by hair-like spines pointing away from capsule on terminal portion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C340C8DDFFE4B13D53669E6EF053FF12" pageId="61" pageNumber="412" type="discussion">
|
||
<paragraph id="8BE59B56FFE4B13D53669E6EF4C9FD63" blockId="61.[92,609,638,1777]" box="[92,190,638,664]" pageId="61" pageNumber="412">Remarks</paragraph>
|
||
<paragraph id="8BE59B56FFE4B13D53589EB3F64AFB2D" blockId="61.[92,609,638,1777]" pageId="61" pageNumber="412">
|
||
<taxonomicName id="4C5AE0D5FFE4B13D53589EB3F518FD47" authorityName="Scott" authorityYear="1995" box="[98,367,674,700]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="helenkingae">
|
||
<emphasis id="B92E4744FFE4B13D53589EB3F518FD47" box="[98,367,674,700]" italics="true" pageId="61" pageNumber="412">Fastosarion helenkingae</emphasis>
|
||
</taxonomicName>
|
||
is only known from the
|
||
<typeStatus id="54E125F4FFE4B13D53B69ED6F4CAFD1B" box="[140,189,710,736]" pageId="61" pageNumber="412">type</typeStatus>
|
||
locality, Mt Elliot NP near Townsville.
|
||
<taxonomicName id="4C5AE0D5FFE4B13D53589EFBF545FCFF" box="[98,306,746,772]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="schelli">
|
||
<emphasis id="B92E4744FFE4B13D53589EFBF545FCFF" box="[98,306,746,772]" italics="true" pageId="61" pageNumber="412">Fastosarion schelli</emphasis>
|
||
</taxonomicName>
|
||
has been reported from the Townsville area, including Hervey Range, Magnetic Island and Mt Stuart (
|
||
<bibRefCitation id="EFCBE6A7FFE4B13D52F29F22F4E5FC8B" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." pageId="61" pageNumber="412" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010</bibRefCitation>
|
||
). The two taxa are very similar in external appearance, differing primarily in size, shell shape and colouration:
|
||
<taxonomicName id="4C5AE0D5FFE4B13D52539F8EF5B2FC4C" box="[361,453,925,951]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="schelli">
|
||
<emphasis id="B92E4744FFE4B13D52539F8EF5B2FC4C" box="[361,453,925,951]" italics="true" pageId="61" pageNumber="412">F. schelli</emphasis>
|
||
</taxonomicName>
|
||
is larger, with a more globose shell and paler body colour (
|
||
<bibRefCitation id="EFCBE6A7FFE4B13D53569FF5F536FC04" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[108,321,997,1024]" pageId="61" pageNumber="412" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010</bibRefCitation>
|
||
). There is also a possible ecological difference:
|
||
<taxonomicName id="4C5AE0D5FFE4B13D5261981AF58FFBD8" authorityName="Scott" authorityYear="1995" box="[347,504,1033,1059]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="helenkingae">
|
||
<emphasis id="B92E4744FFE4B13D5261981AF58FFBD8" box="[347,504,1033,1059]" italics="true" pageId="61" pageNumber="412">F. helenkingae</emphasis>
|
||
</taxonomicName>
|
||
has only been found under logs in the high altitude mesophyll rainforest on the summit of Mt Elliot, while
|
||
<taxonomicName id="4C5AE0D5FFE4B13D53EC9866F547FB74" box="[214,304,1141,1167]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="schelli">
|
||
<emphasis id="B92E4744FFE4B13D53EC9866F547FB74" box="[214,304,1141,1167]" italics="true" pageId="61" pageNumber="412">F.schelli</emphasis>
|
||
</taxonomicName>
|
||
is found semi-arboreally in dry vine thicket on granite, in the foothills of the Hervey Range and on Magnetic Island.
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FFE4B13D53BB98F1F501F97D" blockId="61.[92,609,638,1777]" pageId="61" pageNumber="412">
|
||
We have found that the two species agree very closely in reproductive anatomy, the only difference being slightly coarser pustules in the penis interior of
|
||
<taxonomicName id="4C5AE0D5FFE4B13D526D995DF5C1FA9D" box="[343,438,1356,1382]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="schelli">
|
||
<emphasis id="B92E4744FFE4B13D526D995DF5C1FA9D" box="[343,438,1356,1382]" italics="true" pageId="61" pageNumber="412">F. schelli</emphasis>
|
||
</taxonomicName>
|
||
. They are also genetically very similar. Also, additional material examined herein includes
|
||
<taxonomicName id="4C5AE0D5FFE4B13D52FF9985F628FA55" authorityName="Scott" authorityYear="1995" box="[453,607,1428,1454]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="helenkingae">
|
||
<emphasis id="B92E4744FFE4B13D52FF9985F628FA55" box="[453,607,1428,1454]" italics="true" pageId="61" pageNumber="412">F.helenkingae</emphasis>
|
||
</taxonomicName>
|
||
from the foothills of Mt Elliot and specimens of
|
||
<taxonomicName id="4C5AE0D5FFE4B13D534599CDF4ADFA0D" box="[127,218,1500,1526]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="schelli">
|
||
<emphasis id="B92E4744FFE4B13D534599CDF4ADFA0D" box="[127,218,1500,1526]" italics="true" pageId="61" pageNumber="412">F. schelli</emphasis>
|
||
</taxonomicName>
|
||
from under logs. We do not believe that the minor genetic or morpho-anatomical differences warrant maintaining these taxa as separate species and we therefore treat the two names as synonyms.
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FFE4B13D53BB9A80F053FF12" blockId="61.[92,609,638,1777]" lastBlockId="61.[645,1156,171,234]" pageId="61" pageNumber="412">
|
||
<taxonomicName id="4C5AE0D5FFE4B13D53BB9A80F5F9F952" authorityName="Scott" authorityYear="1995" box="[129,398,1679,1705]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="61" pageNumber="412" phylum="Mollusca" rank="species" species="helenkingae">
|
||
<emphasis id="B92E4744FFE4B13D53BB9A80F5F9F952" box="[129,398,1679,1705]" italics="true" pageId="61" pageNumber="412">Fastosarion helenkingae</emphasis>
|
||
</taxonomicName>
|
||
is allopatric with respect to its congeners. It can be distinguished from other members of Clade A by its generally smaller size and by the presence of a pigmented ridge on the shell lappets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |