229 lines
30 KiB
XML
229 lines
30 KiB
XML
<document id="8431FC46420A451DB5DCDE486994BD58" ID-DOI="10.1163/18759866-20191416" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. & Köhler, Frank" docDate="2019" docId="03F32A40FF8BB15353419DEFF7B3F9E7" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Fastosarion papillosus Stanisic 2010" docType="treatment" docVersion="5" lastPageNumber="433" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="433" updateTime="1695298548213" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1EE14686514EA8138A7CD3E1ADC3C840">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:namePart id="9C094E7818AC6D48227E42D22E00B011">Köhler, Frank</mods:namePart>
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<mods:title id="FE2971830A0BF4D61D8937FD0AD34DA5">Contributions to Zoology</mods:title>
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<mods:date id="536415A05B00C7F3A0D26944BB27FAA4">2019</mods:date>
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<treatment id="03F32A40FF8BB15353419DEFF7B3F9E7" LSID="urn:lsid:plazi:treatment:03F32A40FF8BB15353419DEFF7B3F9E7" httpUri="http://treatment.plazi.org/id/03F32A40FF8BB15353419DEFF7B3F9E7" lastPageId="83" lastPageNumber="433" pageId="82" pageNumber="433">
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<heading id="D0AD2C3AFF8BB15253419DEFF669FDE2" box="[123,542,511,538]" fontSize="11" level="2" pageId="82" pageNumber="433" reason="2">
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<taxonomicName id="4C5AE0D5FF8BB15253419DEFF669FDE2" ID-CoL="3DVWF" ID-ENA="2658020" authority="Stanisic, 2010" authorityName="Stanisic" authorityYear="2010" box="[123,542,511,538]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="papillosus">
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<emphasis id="B92E4744FF8BB15253419DEFF669FDE2" bold="true" box="[123,542,511,538]" pageId="82" pageNumber="433">
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<emphasis id="B92E4744FF8BB15253419DEFF5F7FDE1" bold="true" box="[123,384,511,538]" italics="true" pageId="82" pageNumber="433">Fastosarion papillosus</emphasis>
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<bibRefCitation id="EFCBE6A7FF8BB15252BF9DEFF669FDE2" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[389,542,511,537]" pageId="82" pageNumber="433" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic, 2010</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C340C8DDFF8BB15253BB9E33F5CEFDC6" box="[129,441,547,574]" pageId="82" pageNumber="433" type="description">
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<paragraph id="8BE59B56FF8BB15253BB9E33F5CEFDC6" blockId="82.[129,639,547,646]" box="[129,441,547,574]" pageId="82" pageNumber="433">
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<figureCitation id="136187D3FF8BB15253BB9E33F4A6FDC6" box="[129,209,547,573]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="82" pageNumber="433">Figs. 18</figureCitation>
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,
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<figureCitation id="136187D3FF8BB15253E09E33F56CFDC5" box="[218,283,547,574]" captionStart="FIGURE 15" captionStartId="60.[129,202,1418,1439]" captionTargetBox="[139,1187,188,1385]" captionTargetId="figure-123@60.[129,1187,188,1385]" captionTargetPageId="60" captionText="FIGURE 15 Genitalia of F.griseolus, MO55568, Bobby Range.A.Reproductive system. Arrow shows the junction of the penis and epiphallus.B. Penis interior. Scale bars 1 mm (B), 2 mm (A)." figureDoi="http://doi.org/10.5281/zenodo.8343091" httpUri="https://zenodo.org/record/8343091/files/figure.png" pageId="82" pageNumber="433">15G–I</figureCitation>
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,
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<figureCitation id="136187D3FF8BB152521F9E33F511FDC6" box="[293,358,547,573]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="82" pageNumber="433">19J–K</figureCitation>
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,
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<figureCitation id="136187D3FF8BB15252559E33F5CEFDC6" box="[367,441,547,573]" captionStart="FIGURE 30" captionStartId="83.[98,171,1326,1347]" captionTargetBox="[98,1156,188,1302]" captionTargetId="figure-6@83.[98,1156,188,1302]" captionTargetPageId="83" captionText="FIGURE 30 Genitalia.A-B.F.papillosus, MO78896, Mt Superbus.C-D.F.pustulosus, MO78894, Mt Fort William. A, D.Reproductive system. Arrows show the junction of the penis and epiphallus. B–C. Penis interior. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8343121" httpUri="https://zenodo.org/record/8343121/files/figure.png" pageId="82" pageNumber="433">30A–B</figureCitation>
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</paragraph>
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<subSubSection id="C340C8DDFF8BB15253A59E58F5CAFD7D" pageId="82" pageNumber="433" type="reference_group">
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<treatmentCitationGroup id="AB4ABC78FF8BB15253A59E58F5CAFD7D" pageId="82" pageNumber="433">
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<taxonomicName id="4C5AE0D5FF8BB15253A59E58F5FFFD9A" ID-CoL="6HPMW" authorityName="Stanisic" authorityYear="2010" box="[159,392,583,609]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="papillosa">
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<emphasis id="B92E4744FF8BB15253A59E58F5FFFD9A" box="[159,392,583,609]" italics="true" pageId="82" pageNumber="433">Fastosarion papillosa</emphasis>
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</taxonomicName>
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[sic!]
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<bibRefCitation id="EFCBE6A7FF8BB15252F49E57F615FD9A" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[462,610,583,609]" pageId="82" pageNumber="433" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">
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Stanisic,
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in
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<treatmentCitation id="0AFBBD47FF8BB15253BB9E7BF5CEFD7D" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[129,441,619,646]" page="304" pageId="82" pageNumber="433" year="2010">
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<bibRefCitation id="EFCBE6A7FF8BB15253BB9E7BF5CEFD7D" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[129,441,619,646]" pageId="82" pageNumber="433" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010: 304–305</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BE59B56FF8BB15253419EA3F539FD36" blockId="82.[123,639,691,968]" box="[123,334,691,717]" pageId="82" pageNumber="433">Material examined</paragraph>
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<paragraph id="8BE59B56FF8BB15253BB9EC8F615FCC2" blockId="82.[123,639,691,968]" pageId="82" pageNumber="433">
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<emphasis id="B92E4744FF8BB15253BB9EC8F4B2FD0A" box="[129,197,728,753]" italics="true" pageId="82" pageNumber="433">Types:</emphasis>
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<materialsCitation id="3B32910BFF8BB15253F19EC7F615FCC2" collectingDate="1991-05-24" collectionCode="QM" collectorName="J. Stanisic & D. Potter" latitude="-28.225" longLatPrecision="815" longitude="152.45" pageId="82" pageNumber="433" specimenCode="MO32337" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E125F4FF8BB15253F19EC7F541FD0A" box="[203,310,727,753]" pageId="82" pageNumber="433">Holotype</typeStatus>
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:
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<collectionCode id="ED4B0393FF8BB152527A9EC7F51CFD0A" box="[320,363,727,753]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF8BB152524B9EC7F5ADFD0A" box="[369,474,727,753]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">MO32337</specimenCode>
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(
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<taxonomicName id="4C5AE0D5FF8BB15252D29EC7F60DFD0A" box="[488,634,727,753]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="superbus">Mt Superbus</taxonomicName>
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, via
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<collectorName id="26AFFE80FF8BB15253979EEBF571FCEE" box="[173,262,763,789]" pageId="82" pageNumber="433">Boonah</collectorName>
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, SEQ,
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,
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, under logs, coll.
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<date id="FFE4BD96FF8BB15253D79F0FF507FCC2" box="[237,368,799,825]" pageId="82" pageNumber="433" value="1991-05-24">
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<collectingDate id="EFA0447EFF8BB15253D79F0FF507FCC2" box="[237,368,799,825]" pageId="82" pageNumber="433" value="1991-05-24">24 May 1991</collectingDate>
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</date>
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,
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<collectorName id="26AFFE80FF8BB15252409F0FF591FCC2" box="[378,486,799,825]" pageId="82" pageNumber="433">J. Stanisic</collectorName>
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,
|
||
<collectorName id="26AFFE80FF8BB15252CB9F0FF621FCC2" box="[497,598,799,825]" pageId="82" pageNumber="433">D. Potter</collectorName>
|
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE59B56FF8BB15253A59F53F553FC5F" blockId="82.[123,639,691,968]" pageId="82" pageNumber="433">
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<materialsCitation id="3B32910BFF8BB15253A59F53F5B1FCA6" box="[159,454,835,861]" collectionCode="QM" pageId="82" pageNumber="433" specimenCode="MO23314" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54E125F4FF8BB15253A59F53F579FCA6" box="[159,270,835,861]" pageId="82" pageNumber="433">Paratypes</typeStatus>
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:
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<collectionCode id="ED4B0393FF8BB15252189F53F53AFCA6" box="[290,333,835,861]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF8BB15252649F53F5B1FCA6" box="[350,454,835,861]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">MO23314</specimenCode>
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</materialsCitation>
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,
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<materialsCitation id="3B32910BFF8BB15252E19F53F60BFCA6" box="[475,636,834,861]" collectionCode="QM" pageId="82" pageNumber="433" specimenCode="MO51156" specimenCount="1" typeStatus="paratype">
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<collectionCode id="ED4B0393FF8BB15252E19F53F671FCA6" box="[475,518,835,861]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF8BB152512C9F52F60BFCA6" box="[534,636,834,861]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">MO51156</specimenCode>
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</materialsCitation>
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,
|
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<materialsCitation id="3B32910BFF8BB15253BB9F77F555FC7A" box="[129,290,870,897]" collectionCode="QM" pageId="82" pageNumber="433" specimenCode="MO55366" specimenCount="1" typeStatus="paratype">
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<collectionCode id="ED4B0393FF8BB15253BB9F77F4DBFC7A" box="[129,172,871,897]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF8BB152538F9F76F555FC7A" box="[181,290,870,897]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">MO55366</specimenCode>
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</materialsCitation>
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,
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<materialsCitation id="3B32910BFF8BB152520A9F77F5BAFC7A" box="[304,461,870,897]" collectionCode="QM" pageId="82" pageNumber="433" specimenCode="MO55410" specimenCount="1" typeStatus="paratype">
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<collectionCode id="ED4B0393FF8BB152520A9F77F52CFC7A" box="[304,347,871,897]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">QM</collectionCode>
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<specimenCode id="DBFC332DFF8BB152525E9F76F5BAFC7A" box="[356,461,870,897]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">MO55410</specimenCode>
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</materialsCitation>
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,
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<materialsCitation id="3B32910BFF8BB15252E19F77F60CFC7A" box="[475,635,870,897]" collectionCode="QM" pageId="82" pageNumber="433" specimenCode="MO55432" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED4B0393FF8BB15252E19F77F671FC7A" box="[475,518,871,897]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBFC332DFF8BB15251359F76F60CFC7A" box="[527,635,870,897]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">MO55432</specimenCode>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B32910BFF8BB15253BB9F9BF557FC5F" box="[129,288,906,933]" collectionCode="QM" pageId="82" pageNumber="433" specimenCode="MO78896" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED4B0393FF8BB15253BB9F9BF4DBFC5E" box="[129,172,907,933]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">QM</collectionCode>
|
||
<specimenCode id="DBFC332DFF8BB152538B9F9AF557FC5F" box="[177,288,906,932]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="82" pageNumber="433" type="Museum">MO78896</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FF8BB15253A59FBFF59EFC33" blockId="82.[123,639,691,968]" box="[159,489,942,968]" pageId="82" pageNumber="433">
|
||
<emphasis id="B92E4744FF8BB15253A59FBFF51CFC33" box="[159,363,942,968]" italics="true" pageId="82" pageNumber="433">Non-type material:</emphasis>
|
||
See
|
||
<tableCitation id="C6D8AEEDFF8BB15252A79FBEF592FC33" box="[413,485,942,968]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="82" pageNumber="433" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FF8BB15253419FEAF49FFBEF" blockId="82.[123,641,1018,1727]" box="[123,232,1018,1044]" pageId="82" pageNumber="433">Diagnosis</paragraph>
|
||
<paragraph id="8BE59B56FF8BB15253BB980EF641FAAC" blockId="82.[123,641,1018,1727]" pageId="82" pageNumber="433">
|
||
<emphasis id="B92E4744FF8BB15253BB980EF504FBC3" box="[129,371,1054,1080]" italics="true" pageId="82" pageNumber="433">External morphology</emphasis>
|
||
: Shell (16G–I, 19J–K) medium-sized (
|
||
<quantity id="4CA236B3FF8BB152520A9852F5BDFBA6" box="[304,458,1090,1117]" metricMagnitude="-2" metricUnit="m" metricValue="1.455" metricValueMax="1.48" metricValueMin="1.43" pageId="82" pageNumber="433" unit="mm" value="14.55" valueMax="14.8" valueMin="14.3">14.3–14.8 mm</quantity>
|
||
), golden amber, very transparent, 3.8–4.0 whorls, subglobose with a flat spire. Body beige, sole cream. Mantle lobes and shell lappets moderately small, irregularly marked with black; each lappet with row of 4–5 warts (sometimes pigmented) near lower edge. Tail slightly keeled at tip, slime network weak, caudal horn large.
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FF8BB15253A59971F00CFECA" blockId="82.[123,641,1018,1727]" lastBlockId="82.[677,1188,171,305]" pageId="82" pageNumber="433">
|
||
<emphasis id="B92E4744FF8BB15253A59971F511FA80" box="[159,358,1377,1403]" italics="true" pageId="82" pageNumber="433">Genital anatomy:</emphasis>
|
||
Genitalia (fig. 30A–B) with long bulging vagina; bursa copulatrix very short, duct distinct, bursa oval. Penis long, broad, cylindrical, proximally fused to epiphallus 2, inner penial wall smooth, with three transversely ridged longitudinal pilasters, two not reaching the distal end; 35–65% of penis contained in penial tunica. Epiphallus longer than penis; entering penis through a simple pore; epiphallus 2 much longer than epiphallus 1; half of epiphallus 2 fused to penis; epiphallic caecum of moderate length, penial retractor muscle attaching to the middle; flagellum of moderate length, slender.
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FF8BB15251A59D63F776FE76" blockId="82.[671,1189,371,1726]" box="[671,769,371,397]" pageId="82" pageNumber="433">Remarks</paragraph>
|
||
<paragraph id="8BE59B56FF8BB152519F9D87F746FC37" blockId="82.[671,1189,371,1726]" pageId="82" pageNumber="433">
|
||
The spelling of the species name is adapted herein to agree with the masculine gender of the genus name.
|
||
<taxonomicName id="4C5AE0D5FF8BB15250AA9DCFF0F0FE02" authorityName="Stanisic" authorityYear="2010" box="[912,1159,479,505]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF8BB15250AA9DCFF0F0FE02" box="[912,1159,479,505]" italics="true" pageId="82" pageNumber="433">Fastosarion papillosus</emphasis>
|
||
</taxonomicName>
|
||
is recorded from Richmond Range in northeastern NSW to Lamington NP in southeastern Qld (
|
||
<bibRefCitation id="EFCBE6A7FF8BB15250359E5AF790FD9F" author="Stanisic, J. & Shea, M. & Potter, D. & Griffiths, O." box="[783,999,586,613]" pageId="82" pageNumber="433" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. & Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010</bibRefCitation>
|
||
). In the present study we found that candidate taxa
|
||
<taxonomicName id="4C5AE0D5FF8BB15257789E7EF76EFD57" authority="BR" authorityName="Bourguignat" authorityYear="1877" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="family">Helicarionidae BR</taxonomicName>
|
||
12 from Tabletop Bush Reserve and SQ6 from Kroombit Tops, previously identified through curatorial work, also belong to this species. The Kroombit Tops population is approximately
|
||
<quantity id="4CA236B3FF8BB15250929F32F788FCC7" box="[936,1023,802,828]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="82" pageNumber="433" unit="km" value="400.0">400 km</quantity>
|
||
further north than the rest of the included populations; however, both morpho-anatomical and molecular data confirm its placement as part of
|
||
<taxonomicName id="4C5AE0D5FF8BB152519D9FA2F75CFC30" authorityName="Stanisic" authorityYear="2010" box="[679,811,945,971]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF8BB152519D9FA2F75CFC30" box="[679,811,945,971]" italics="true" pageId="82" pageNumber="433">F. papillosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE59B56FF8BB15351F99FC5F7B3F9E7" blockId="82.[671,1189,371,1726]" lastBlockId="83.[646,1157,1502,1564]" lastPageId="83" lastPageNumber="434" pageId="82" pageNumber="433">
|
||
The sister taxon to this species is the very similar
|
||
<taxonomicName id="4C5AE0D5FF8BB152503A9FEAF7E4FBE8" box="[768,915,1017,1043]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="pustulosus">
|
||
<emphasis id="B92E4744FF8BB152503A9FEAF7E4FBE8" box="[768,915,1017,1043]" italics="true" pageId="82" pageNumber="433">F. pustulosus</emphasis>
|
||
</taxonomicName>
|
||
. A comparison of the original descriptions indicates that
|
||
<taxonomicName id="4C5AE0D5FF8BB152570D980EF6BFFBA0" authorityName="Stanisic" authorityYear="2010" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF8BB152570D980EF6BFFBA0" italics="true" pageId="82" pageNumber="433">F. papillosus</emphasis>
|
||
</taxonomicName>
|
||
is slightly larger, with a row of black lappets on the right shell lappet and no pigmentation on the left lappet, and with pale pinkish grey body colouration, while
|
||
<taxonomicName id="4C5AE0D5FF8BB15250D198BEF000FB3C" box="[1003,1143,1197,1223]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="pustulosus">
|
||
<emphasis id="B92E4744FF8BB15250D198BEF000FB3C" box="[1003,1143,1197,1223]" italics="true" pageId="82" pageNumber="433">F. pustulosus</emphasis>
|
||
</taxonomicName>
|
||
has unpigmented pustules on the right shell lappet and three black stripes on the left lappet, and dark body colouration. We found some of these reported differences to be erroneous (although observations were based only on preserved data): both species were found to have a row of around five large pustules or warts (sometimes pigmented in
|
||
<taxonomicName id="4C5AE0D5FF8BB152570299DDF6BFF9F1" authorityName="Stanisic" authorityYear="2010" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF8BB152570299DDF6BFF9F1" italics="true" pageId="82" pageNumber="433">F. papillosus</emphasis>
|
||
</taxonomicName>
|
||
) on both left and right lappets, and both also had black markings on the shell lappets (particularly on the left). The most consistent external difference was the higher spire seen in
|
||
<taxonomicName id="4C5AE0D5FF8BB152503B9A90F7FAF962" box="[769,909,1663,1689]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="82" pageNumber="433" phylum="Mollusca" rank="species" species="pustulosus">
|
||
<emphasis id="B92E4744FF8BB152503B9A90F7FAF962" box="[769,909,1663,1689]" italics="true" pageId="82" pageNumber="433">F. pustulosus</emphasis>
|
||
</taxonomicName>
|
||
. The two species shared a unique genital character comprising fusion of the two epiphallus arms and the proximal portion of the penis. However, in
|
||
<taxonomicName id="4C5AE0D5FF8AB15352EE9A13F617F9E7" box="[468,608,1538,1564]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="83" pageNumber="434" phylum="Mollusca" rank="species" species="pustulosus">
|
||
<emphasis id="B92E4744FF8AB15352EE9A13F617F9E7" box="[468,608,1538,1564]" italics="true" pageId="83" pageNumber="434">F.pustulosus</emphasis>
|
||
</taxonomicName>
|
||
the penis was longer and narrower, and approximately half of the penis was fused to the epiphallus, while in
|
||
<taxonomicName id="4C5AE0D5FF8AB15352059A7FF5B2F973" authorityName="Stanisic" authorityYear="2010" box="[319,453,1646,1672]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="83" pageNumber="434" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF8AB15352059A7FF5B2F973" box="[319,453,1646,1672]" italics="true" pageId="83" pageNumber="434">F.papillosus</emphasis>
|
||
</taxonomicName>
|
||
the penis was very broad and only the proximal quarter or third was fused to the epiphallus. Moreover,
|
||
<taxonomicName id="4C5AE0D5FF8AB15351749AA6F68CFA03" box="[590,763,1502,1743]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="83" pageNumber="434" phylum="Mollusca" rank="species" species="pustulosus">
|
||
<emphasis id="B92E4744FF8AB15351749AA6F68CFA03" box="[590,763,1502,1743]" italics="true" pageId="83" pageNumber="434">F. pustulosus</emphasis>
|
||
</taxonomicName>
|
||
has one major longitudinal pilaster while
|
||
<taxonomicName id="4C5AE0D5FF8AB15351F09A13F727F9E7" authorityName="Stanisic" authorityYear="2010" box="[714,848,1538,1564]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="83" pageNumber="434" phylum="Mollusca" rank="species" species="papillosus">
|
||
<emphasis id="B92E4744FF8AB15351F09A13F727F9E7" box="[714,848,1538,1564]" italics="true" pageId="83" pageNumber="434">F. papillosus</emphasis>
|
||
</taxonomicName>
|
||
has three.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |